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1.
The traction capacity of the mining machine is greatly in?uenced by the traction rheological properties of the deep-sea sediments. The best simulative soil was prepared for substituting the deep-sea sediment based on the deep-sea sediment collected from the Paci?c C-C mining area. Traction rheological properties of the simulative soil were studied by a home-made test apparatus. In order to accurately describe the traction rheological properties and determine traction rheological parameters, the Newtonian dashpot in Maxwell body of Burgers model was replaced by a self-similarity spring-dashpot fractance and a new rheological constitutive model was deduced by fractional derivative theory. The results show the simulative soil has obvious non-attenuate rheological properties. The transient creep and stable creep rate increase with the traction, but they decrease with ground pressure. The fractional derivative Burgers model are better in describing non-attenuate rheological properties of the simulative soil than the classical Burgers model. For the new traction rheological constitutive equation of the simulative soil, the traction rheological parameters can be obtained by ?tting the tested traction creep data with the traction creep constitutive equation. The ground contact length of track and walking velocity of the mining machine predicted by the traction rheological constitutive equation can be used to take full advantages of the maximum traction provided by the soil and safely improve mining effciency.  相似文献   

2.
In order to identify environmental factors driving the distribution and functioning of deep-sea fauna and the spatial scales of interactions, we carried out a multiple-scale investigation in the Mediterranean basin in which we compared two bathyal plains, located at the same depth (ca. 3000 m), but characterised by contrasting trophic conditions. We investigated meiofaunal abundance, biomass, community structure and biodiversity (expressed as richness of taxa) in relation to sediment characteristics, downward fluxes and food availability in the sediment. Samples were collected at all spatial scales (from small to macroscale) in two seasons. Our results indicated that deep-sea systems with different trophic conditions displayed different responses to the distribution of available energy and its spatio-temporal variability in the sediment. The analysis at a macroscale (>1000 km) indicated that meiofauna were controlled primarily by the trophic inputs to the deep-sea system. Spatial variability of meiofaunal parameters at a mesoscale (>50 km) was highest in the eastern Mediterranean and lowest in the western Mediterranean. Such differences are the consequence of the unpredictable inputs of organic matter in the oligotrophic eastern Mediterranean versus a more homogeneous distribution of food inputs in the mesotrophic western Mediterranean. At a smaller scale (local scale 7 km), in the western Mediterranean, the distribution of meiofaunal parameters was highly homogeneous, reflecting the homogeneous distribution of the food availability in the sediment. Our results indicated that the highly variable input and distribution of food sources in the deep eastern Mediterranean did not provide any “insurance” for the sustainability of the deep-sea faunal assemblages in the long term, thus leading to an uncoupling between resource availability and distribution of organisms. We conclude that the influence of energy availability on the deep-sea faunal distributions change at different spatial scales and that the analysis of spatial variability at mesoscales is crucial for understanding the relationships between deep-sea benthic fauna and environmental drivers.  相似文献   

3.
The impact load (equivalent impact height) applied to deep-sea sediment by a walking mining machine was first deduced by the energy conservation principle, and the simulative soil was prepared based on the deep-sea sediment collected from the Pacific C-C mining area. The self-designed impact compressive creep tests of the simulative soil were conducted under different ground stresses and impact heights, in order to determine impact compressive creep parameters using a K-H rheological model. Test results show that the impact compressive creep curves have three similar creep stages (transient creep, unstable creep, and stable creep) to static compressive creep curves, where the transient creep deformation and total deformation at the unstable creep stage decrease with the impact load. Among the three impact compressive creep parameters (K1, K2, β) of the simulative soil, K1 is first increased with impact height and finally fluctuated to a certain stable value, while K2 and β are approximately linearly increased with impact height. The maximum subsidence of the mining machine under a specific designed ground stress and walking velocity predicted by the impact compressive creep constitutive equation can be used for safety assessment of the mining machine.  相似文献   

4.
I am deeply honored to have been awarded the 1988 Okada Prize of the Oceanographical Society of Japan. The present paper reviews my previous works regarding the ecology of deep-sea meiobenthos in the western Pacific area. The outline cen be summarized as follows:
1)  On the basis of multivariate analyses, it was found that the rate of the organic-matter flux to the sea bed and the amount of the interstitial space within the sediment are the main factors regulating the abundance of meiofauna in the deep sea.
2)  Two indices were proposed to characterize the vertical distribution of meiofauna in the sediment profile,i.e. their maximum depth in the sediment and the degree of their concentration in the surface layer of the sediment. The index of maximum depth was closely related to the oxygen concentration in the interstitial water. On the other hand, the index of degree of concentration in the surface of the sediment tended to be higher where the supply of food seemed lower.
3)  Some taxonomically important species were found from the axis of the Izu-Ogasawara Trench. They arePliciloricus hadalis, the first species of the newest phylum Loricifera from the Pacific area, from the hadal deep sea and the fine clay sediment, andOccultammina profunda, the first infaunal species of Xenophyophorea, a group of rhizopod Protozoa.
4)  The vertical distribution ofOccultammina profunda coincided well with the unusual distribution of210Pb in the sediment profile. The organisms concentrated as high as 500 dpm g–1 of210Pb in their stercomare and granellare. The steady state model confirmed that the species made the peculiar subsurface peak of210Pb in the sediment.
5)  The depth in the sediment profile where the distinct peak of MnO2 can be seen showed strong correlation with the vertical distribution of meiofauna. This relationship suggests that oxygen concentration in the interstitial water regulates the structure of the characteristic three layers of the calcareous ooze in the deep sea. This idea was proven by the vertical distribution of free oxygen in the sediment calculated on the basis of respiration rate of deep-sea meiofauna measured using the cartesian diver technique.
6)  Radio-isotope techniques used to measure the rates at which particulated organic matter was ingested and dissolved organic matter was absorbed, suggested that deep-sea meiofauna obtained a significant fraction of their energy by absorption.
7)  The work carried out so far has revealed the important role of meiofauna in the benthic ecosystem, and emphasized the necessity for intensive research from various points of views on these microscopic organisms.
  相似文献   

5.
The abundance of baroduric bacteria from nine sediment cores (0–10 cm below sea floor) was examined for their response to simulated benthic disturbance in the Central Indian basin (CIB). While the total counts of bacteria decreased from 109 to 106 g?1 dry wt sediment, the average retrievable counts (CFU-colony forming units) improved by two orders of magnitude, i.e., from 102 104 g?1 dry wt sediment. The baroduric retrievable forms were dominated by Acinetobacter and Moraxella sp before the disturbance. After the disturbance the generic diversity was represented by Staphylococcus sp, Enterobacter sp, Micrococcus sp, Coryneforms sp, and Pseudomonas sp in addition to Acinetobacter. These observations were corroborated by changes in enzymatic activities of the retrievable bacteria, which could lead to changes in the biochemical characteristics of the sediment. Thus the simulated disturbance brought about an increase in culturable abundance, taxonomic and functional diversity of deep-sea sediment of the CIB.  相似文献   

6.
通过网格定点法对我国东南沿海区域性台风危险性进行了分析。利用对各网格点有影响的历史台风数据,建立了各网格点的台风关键参数的最优概率模型。利用Monte-Carlo方法产生每个网格点1000年间的虚拟台风事件。采用YanMeng(YM)风场模型模拟了100个历史台风的最大风速,通过使这些最大风速与观测的最大风速误差和最小,建立了一组新的计算最大风半径Rmax和Holland气压参数B的公式,结果表明新的台风参数计算方案效果良好。利用新的参数计算方案、YM风场模型、特定地点的台风衰减模型以及合适的极值分布模型,预测了各个网格点不同重现期的极值风速,进而绘制了台风多发区域的设计风速图。最后研究了不同B模型、Rmax模型和极值分布模型对预测的极值风速的影响。可以为结构抗风设计和台风防灾减灾提供新的参考。  相似文献   

7.
海底底质的物理力学参数不同于陆地土壤,其极低的抗剪强度和承压强度对深海采矿车的行走性能提出高要求。分析基于车辆地面力学理论,开展了底质土力学特性试验研究,建立了深海底质力学模型。根据深海重载作业采矿车样机结构参数,在大型动力学仿真软件Recurdyn中建立了仿真模型。通过直线行走多体动力学仿真与直线行走海试试验的对比,验证了仿真模型的准确性。在此基础上开展了采矿车样机在深海软底质上的多种行走工况动力学仿真,分析与评价其行走性能。结果表明,采矿车样机可以顺利完成转弯、爬坡、越障等基本功能。该研究成果可为深海采矿车海底行走性能评估提供理论参考,为深海采矿车和软底质的相互作用力学研究提供借鉴。  相似文献   

8.
The Zenisu deep-sea channel originates on the Izu-Ogasawara island arc, and disappears in the Shikoku Basin of the Philippine Sea. The geomorphology, sedimentary processes, and the development of the Zenisu deep-sea channel were investigated on the basis of swath bathymetry, side-scan sonar imagery, submersible observations, and seismic data. The deep-sea channel can be divided into three segments according to the downslope gradient and channel orientation. They are the Zenisu Canyon, the E–W fan channel, and the trough-axis channel. The sediment fill is characterized by turbidite and debrite deposition and blocky–hummocky avalanche deposits on the flanks of the Zenisu Ridge. In the Zenisu Canyon and the Zenisu deep-sea channel, sediment transport by turbidity currents generates sediment waves (dunes) observed during the Shinkai 6500 dive 371. The development of the Zenisu Canyon is controlled by a N–S shear fault, whereas the trough-axis channel is controlled by basin subsidence associated with the Zenisu Ridge. The E–W fan channel was probably affected by the E–W fault and the basement morphology.  相似文献   

9.
Resuspension, transport, and deposition of sediments over the continental shelf and slope are complex processes and there is still a need to understand the underlying spatial and temporal dynamical scales. As a step towards this goal, a two-dimensional slice model (zero gradients in the alongshore direction) based on the primitive flow equations and a range of sediment classes has been developed. The circulation is forced from rest by upwelling or downwelling winds, which are spatially uniform. Results are presented for a range of wind speeds and sediment settling speeds. Upwelling flows carry fine sediments (low settling speeds) far offshore within the surface Ekman layer, and significant deposition eventually occurs beyond the shelf break. However, coarser sediments quickly settle out of the deeper onshore component of the circulation, which can lead to accumulation of bottom sediments within the coastal zone. Downwelling flows are more effective at transporting coarse sediments off the shelf. However, strong vertical mixing at the shelf break ensures that some material is also carried into the surface Ekman layer and returned onshore. The concentrations and settling fluxes of coarse sediments decrease offshore and increase with depth under both upwelling and downwelling conditions, consistent with trends observed in sediment trap data. However, finer sediments decrease with depth (upwelling) or reach a maximum around the depth of the shelf break (downwelling). It is shown that under uniform wind conditions, suspended sediment concentrations and settling fluxes decay offshore over a length scale of order τs/ρf|ws|, where τs is the wind stress, ρ the water density, f the Coriolis parameter, and ws is the sediment settling velocity. This scaling applies to both upwelling and downwelling conditions, provided offshore transport is dominated by wind-driven advection, rather than horizontal diffusion.  相似文献   

10.
Rates of transformation, recycling and burial of nitrogen and their temporal and spatial variability were investigated in deep-sea sediments of the Porcupine Abyssal Plain (PAP), NE Atlantic during eight cruises from 1996 to 2000. Benthic fluxes of ammonium (NH4) and nitrate (NO3) were measured in situ using a benthic lander. Fluxes of dissolved organic nitrogen (DON) and denitrification rates were calculated from pore water profiles of DON and NO3, respectively. Burial of nitrogen was calculated from down core profiles of nitrogen in the solid phase together with 14C-based sediment accumulation rates and dry bulk density. Average NH4 and NO3-effluxes were 7.4 ± 19 μmol m−2 d−1 (n = 7) and 52 ± 30 μmol m−2 d−1 (n = 14), respectively, during the period 1996–2000. During the same period, the DON-flux was 11 ± 5.6 μmol m−2 d−1 (n = 5) and the denitrification rate was 5.1 ± 3.0 μmol m−2 d−1 (n = 22). Temporal and spatial variations were only found in the benthic NO3 fluxes. The average burial rate was 4.6 ± 0.9 μmol m−2 d−1. On average over the sampling period, the recycling efficiency of the PON input to the sediment was 94% and the burial efficiency hence 6%. The DON flux constituted 14% of the nitrogen recycled, and it was of similar magnitude as the sum of burial and denitrification. By assuming the PAP is representative of all deep-sea areas, rates of denitrification, burial and DON efflux were extrapolated to the total area of the deep-sea floor (>2000 m) and integrated values of denitrification and burial of 8 ± 5 and 7 ± 1 Tg N year−1, respectively, were obtained. This value of total deep-sea sediment denitrification corresponds to 3–12% of the global ocean benthic denitrification. Burial in deep-sea sediments makes up at least 25% of the global ocean nitrogen burial. The integrated DON flux from the deep-sea floor is comparable in magnitude to a reported global riverine input of DON suggesting that deep-sea sediments constitute an important source of DON to the world ocean.  相似文献   

11.
Meiofauna in a cold-seep community off Hatsushima,central Japan   总被引:2,自引:0,他引:2  
The community structure of the bathyal meiofauna of a cold-seep community found off Hatsushima in Sagami Bay, central Japan, was compared with the community composition outside the influence of the seep, using sediments collected during dives 226 and 227 of the deep-sea submersibleShinkai 2000. The sediment from the Hatsushima seep site (HSS) was very coarse, black in color, and with an odor of hydrogen sulfide, suggesting reduced thiobiotic conditions. The sediment from the control area was well-oxygenated, fine silt. Despite the differences in the characteristics of the sediments, the abundance of meiofauna in the HSS was not very different from that in the control area. However, its composition even at the major taxonomic group level was distinct; for example, a high nematode/copepod ratio occurred in one of the samples collected at the HSS. At the species level, nematodes were less diverse at the HSS than at the control area. The composition of the nematode fauna at the HSS showed stronger affinity with that collected at the adjacent control area than with a community sampled from other deep-sea environments or another seep community in shallow water. This emphasizes that the adaptation of nematodes to the thiobiotic condition is controlled by local conditions.  相似文献   

12.
To accurately characterize sound speed dispersion of shallow sediments in the Southern Yellow Sea, three types of sediments, i.e., silt, clayey silt, and silty clay, were selected to measure the sound speeds at 25–250?kHz. Over the frequency range, the sound speeds vary approximately from 1,536 to 1,565?m?s?1 in silt sediment, from 1,511 to 1,527?m?s?1 in clayey silt sediment, and from 1,456 to 1,466?m?s?1 in silty clay sediment. The sound speed exhibits a slow increase with frequency in a nearly linear gradient, but these three types of sediments have different sound speed dispersion characteristics. The silt sediment with relatively coarse grains has the most significant sound speed dispersion, while the sound speed dispersions of the two others are relatively weak. Comparison between the measured dispersions and the model predictions shows that the grain-shearing model can match the measured data at most of frequencies. Nevertheless, when the grain bulk modulus was assigned 3.2?×?1010?Pa according to relevant references, the Biot–Stoll model predictions were higher than the measured values at high frequencies; when it was assigned a relatively small value of 2.8?×?1010?Pa, the model predictions achieved optimal matching with the measured values.  相似文献   

13.
曾志刚 《海洋与湖沼》2021,52(6):1333-1349
海底热液活动调查研究是深海进入、深海探测和深海开发的切入点之一。近十年来,中国在西太平洋弧后盆地、东太平洋海隆、大西洋洋中脊和印度洋脊,发现了一批新的海底热液活动区,围绕着热液活动区的硫化物、流体、热液柱、生物等热液产物开展了调查研究,构建了海底热液地质学,提出了热液活动、冷泉及天然气水合物的同源异汇假说,出版了《海底热液地质学》、《现代海底热液硫化物成矿地质学》、《现代海底热液活动》、《东太平洋海隆热液地质》专著,获得了一批调查研究成果。未来,聚焦海底热液活动的深部过程及其资源环境效应关键问题,发展海底热液活动探测技术,拓展极地海底热液活动调查研究新领域,围绕烟囱体、热液柱、含金属沉积物、流体以及热液区生物等热液产物,开展深入、系统的调查研究工作,无疑将推动海底热液地质学取得新的进展。  相似文献   

14.
Key aspects of deep-ocean fluid dynamics such as basin-scale (residual) and tidal flow are believed to have changed over glacial/interglacial cycles, with potential relevance for climatic change. To constrain the mechanistic links, magnitudes and temporal succession of events analyses of sedimentary paleo-records are of great importance. Efforts have been underway for some time to reconstruct residual-flow patterns from sedimentary records. Attempts to reconstruct tidal flow characteristics from deep-sea sediment deposits, however, are at a very early stage and first require a better understanding of the reflection of modern tides in sediment dynamics. In this context internal (baroclinic) tides, which are formed by the surface (barotropic) tide interacting with seafloor obstacles, are believed to play a particularly important role. Here we compare two modern deep-sea environments with respect to the effect of tides on sediment dynamics. Both environments are influenced by kilometre-scale topographic features but with vastly different tidal forcing: (1) two sites in the Northeast Atlantic (NEA) being surrounded by, or located downstream of, fields of short seamounts (maximum barotropic tidal current velocities ~5 cm s?1); and (2) a site next to the Anaximenes seamount in the Eastern Mediterranean (EMed) (maximum barotropic tidal current velocities ~0.5 cm s?1). With respect to other key fluid-dynamical parameters both environments are very similar. Signals of sedimentary particle dynamics, as influenced by processes taking place in the bottom boundary layer, were traced by the vertical water-column distribution of radioactive disequilibria (daughter/parent activity ratios≠1) between the naturally occurring, short-lived (half-life: 24.1 d) particulate-matter tracer 234Th relative to its very long-lived and non-particle-reactive parent nuclide 238U. Activity ratios of 234Th/238U<1 in water samples collected near the seafloor indicate fast 234Th scavenging onto particles followed by fast settling of these particles from the sampled parcel of water and, therefore, imply active sediment resuspension and dynamics on time scales of up to several weeks. In the Northeast Atlantic study region tides (in particular internal tides) are very likely to locally push total current velocities near the seafloor across the critical current velocity threshold for sediment erosion or resuspension whereas in the Eastern Mediterranean the tides are much too weak for this to happen. This difference in tidal forcing is reflected in a difference of the frequency of the occurrence of radioactive disequilibria <1 between total 234Th and 238U: In the near-bottom water column of the Northeast Atlantic region 59% of samples had detectable 234Th/238U disequilibria whereas at the Eastern Mediterranean site this fraction was only 7% (including a few disequilibria >1). The results of this study, therefore, add to the evidence suggesting that tides in the deep sea of the open oceans are more important for sediment dynamics than previously thought. It is hypothesised that (a) tide/seamount interactions in the deep open ocean control the local distribution of erosivity proxies (e.g., distributions of sediment grain sizes, heavy minerals and particle-reactive radionuclides) in sedimentary deposits and (b) the aforementioned topographically controlled sedimentary imprints of (internal) tides are useful in the reconstruction of past changes of tidal forcing in the deep sea.  相似文献   

15.
Abstract

Particle fluxes were measured 7 m above the sea bottom during the predisturbance, disturbance, and postdisturbance periods by using time series sediment traps attached to seven deep-sea moorings deployed in the INDEX experiment site in the Central Indian Basin. The predisturbance particle fluxes varied between 22.3 to 55.1 mg m?2 day?1. Increased and variable particle fluxes were recorded by the sediment traps during the disturbance period. The increase observed was 0.5 to 4 times more than the background predisturbance fluxes. The increases in particle fluxes (~4 times) recorded by the sediment trap located in the southwestern direction (DMS-1) were the greatest, which could be the result of preferential movement of resuspended particles generated during the deep-sea benthic disturbance along the general current direction prevailing in this area during the experimental period. Also, the traps located closer to the disturbance area recorded greater fluxes than did the traps far away, across the Deep Sea Sediment Resuspension System path. This variability in recorded particle fluxes by the traps around the disturbance area clearly indicates that physical characteristics such as grain size and density of the resuspended particles produced during the disturbance had an important effect on particle movement. The postdisturbance measurements during ~5 days showed a reduction in particle fluxes of ~50%, indicating rapid particle settlement.  相似文献   

16.
A sediment disk reactor was tested in once flow-through mode to retrieve kinetic parameters for the Monod rate law that describes sulfate reduction. The experimental method was compared with a previously described procedure by the authors where a sediment plug-flow reactor was operated in a recirculation mode. In recirculation mode, accumulation of metabolic byproducts in certain cases may result in negative feedback, thus preventing accurate determination of kinetic information. The method described in this article provides an alternative to the recirculation sediment plug-flow-through reactor technique for retrieving kinetic parameters of microbially mediated reactions in aquatic sediments.For sulfate reduction in a saltmarsh site, a maximum estimate of the half-saturation concentration, Ks, of 204±26 μM and a maximum reaction rate, Rm, of 2846±129 nmol cm(wetsediment)3 d−1 was determined. The Ks value obtained was consistent with the one estimated previously (Ks=240±20 μM) from a different site within the same saltmarsh mud flat using a recirculating reactor. From the Rm value and reduction rates determined using 35SO42− incubation experiments, we infer that sulfate reduction is limited in the field. Substrate availability is not the main contributor for the limitation, however. Competition from other microbes, such as iron reducers affects the activity of sulfate reducers in the suboxic to anoxic zones, whereas aerobes compete in the oxic zone. High sulfide concentration in the pore water may also have acted as a toxin to the sulfate reducers in the field.  相似文献   

17.
Zai-Jin You   《Ocean Engineering》2004,31(16):1955-1965
A laboratory study was carried to qualitatively investigate the effect of suspended sediment concentration C on the settling velocity ws of cohesive sediment in quiescent water. A bay mud sample was mixed with water in a cylindrical container, and three optical back scatterance sensors were then used to measure suspended sediment concentrations of the mud–water mixture at three levels every 15 s for 5 h while sediments were settling in the quiescent water. Based on the measured sediment concentrations, the settling velocities at different concentrations were derived from the depth-integrated mass balance equation. This study has found that the settling velocity ws is independent of C in the free settling regime of C<0.3 g/l, and then increases nonlinearly with C in the enhanced settling regime of 0.3<C<4.3 g/l, and finally decreases sharply with C in the hindered settling regime of C>4.3 g/l. The maximum settling velocity occurs at C≈4.3 g/l and is about nine times faster than the settling velocity in the free settling regime. A single empirical formula is also proposed to calculate the settling velocities at different sediment concentrations.  相似文献   

18.
贻贝营半固着生活,依靠足丝将自己牢固地黏附在底物上.足丝的结构和性能对贻贝的正常生命活动至关重要.为研究热液区极端环境下深海贻贝足丝的结构和性能,采集了印度洋龙旂深海热液区的贻贝(Bathymodiolus marisindicus)足丝,对其力学特性、结构特征和组分进行了分析.力学性能测试发现与近海贻贝(Mytilu...  相似文献   

19.
Deposit-feeding holothurians often dominate the megafauna in bathyal deep-sea settings, in terms of both abundance and biomass. Molpadia musculus is particularly abundant at about 3400 m depth in the Nazaré Canyon on the NE Atlantic Continental Margin. However, these high abundances are unusual for burrowing species at this depth. The objective of this research was to understand the reasons of the massive occurrence of these molpadiid holothurians in the Nazaré Canyon. To address this question we investigated possible trophic interactions with bacteria at sites where the organic content of the sediment was different (Setúbal and Cascais Canyons, NE Atlantic Continental Margin). The molecular fingerprinting technique of Denaturing Gradient Gel Electrophoresis (DGGE) with band sequencing, combined with non-metric multi-dimensional scaling and statistical analyses, was used to compare the bacterial community diversity in canyon sediments and holothurian gut contents. Our results suggest that M. musculus does not need to develop a specialised gut bacterial community to aid digestion where the sediment is rich in organic matter (Nazaré Canyon); in contrast, such a community may be developed where the sediment is poorer in organic matter (Cascais Canyon).  相似文献   

20.
Bathymetric distribution, abundance, substrate choice and gametogenesis have been investigated in two species of deep-sea actiniarians found in the northeast Atlantic Ocean. Phelliactis hertwigi occurs at 719–1448m in the Porcupine Seabight, usually encloses a bolus of sediment within its highly concave pedal disc (79.7%), and has abundances of up to 14.5 individuals 1000m−2. Phelliactis robusta occurs at 1600–2173m in the Porcupine Seabight, but extends deeper in the Bay of Biscay. In areas of soft sediment it is associated strongly with clinker (92.7%) and attains densities of 2.9 individuals 1000m−2.Both species are dioecious. Their sperm appear similar to those of the related intertidal Calliactis spp. Previtellogenesis and vitellogenesis of the oocytes have been defined by ultrastructural and histochemical studies. In P. hertwigi mature oocytes measure up to 180μm, and to 210μm in P. robusta. In P. hertwigi oogenesis takes 8–9 months, with spawning in October/November, whereas in the deeper-living P. robusta oogenesis occupies 15 to 19 months and spawning occurs in April/ May. Evidence is produced to suggest that these two contrasting cycles are related to the rate and seasonality of deposition of organic matter to the deep-sea floor.  相似文献   

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