Ichnofossils from the Miocene Middle Bhuban Formation, Mizoram,Northeast India and their Paleoenvironmental Significance

An ichnofossil assemblage is preserved in sediments of the Middle Bhuban Formation,Surma Group,of Miocene age from the Durtlang and Chanmari sections,Aizawl district,Mizoram,northeastern India.The studied sections comprise a thick,bioturbated and thinly bedded sandstone–silty shale facies.The preserved ichnofossils show high density and low diversity with the assemblage represented by feeding and resting traces,i.e.,Psilonichnus upsilon Frey et al.,1984,Ophiomorpha isp.,Teichichnus spiralis Mikulá.1990,Skolithos isp.,Palaeophycus isp.,and other horizontal burrows.Sedimentary structures associated with these traces are mainly cross bedding,flaser bedding and slump structures.Considering the distribution of the Psilonichnus ichnogenus across an integrated ichnological–sedimentological framework,the stratigraphic interval of the Middle Bhuban Formation,in which they occur,are interpreted to have been deposited under a shallow,marginal-marine channel complex dominated by tidal channels developed in quiet,brackish-water portions of a delta plain.     

New Early Cretaceous Pterosaur-Bird Track Assemblage from Xinjiang, China: Palaeoethology and Palaeoenvironment

A pterosaur-bird track assemblage from a sandstone-siltstone-mudstone sequence of the Lower Cretaceous Tugulu Group of Xinjiang comprises the first pterosaur track record from this province and the largest specimen thus far known from China.The pterosaur tracks are assigned to the ichnogenus Pteraichnus based on the triangular overall-shape,the four elongate digit traces and the robust manual digit trace Ⅲ.Supposed trackmakers were dsungaripterid pterodactyloids whose skeletal remains are well known from the Tugulu Group.The bird tracks that occur on the same surface,are those of typical shorebirds,known from different other localities in southeast Asia.The congruence with Koreanaornis dodsoni described from the same stratigraphic level justifies an assignment to this ichnospecies.This is a further evidence of the co-occurrence of pterosaurs and birds in a typical iakeshore environment with possible seasonal alteration of water supply and aerial exposure indicated by wave ripples,mudcracks and repeated cycles of coarse to fine sediment.Pterosaurs and birds frequented the shoreline and may have fed also on the numerous invertebrates such as the Scoyenia tracemaker that left abundant burrows.     

Research on Dinoasur Footprints in Zizhou, Shaanxi Province, China

In recent years, the discoveries of dinosaur footprints have been successively reported from Dianshi Town, Zizhou City, Shaanxi Province. The footprints include the tracks of theropod, ornithopod and sauropod as well. Actually, the dinosaur footprints were found by local inhabitants much earlier in history, but not for science. The slabs bearing the dinosaur footprints were collected for domestic use, such as building stones, millstones, cellar covers, sheepfold fences, windlass holders, etc. This paper is to describe the dinosaur footprints on both sides of three slabs used for cellar covers, sheepfold fences and windlass holders by the local people. 24 dinosaur footprints and 4 trackways have been recognized and all of them belong to theropod. Four kinds of dinosaur footprints are identified, including 1 new ichnogenus and 2 new ichnospecies: (1) Shanbeipus caudatus ichnogen. et ichnosp. nov.; (2) Pengxianpus yulinensis ichnosp. nov.; (3) Eubrontes ichnosp; (4) Kayentapus ichnosp. Dinosaur-footprint-bearing beds were initially identified as the Fuxian Formation of the Lower Jurassic. Multipal dinosaur footprints associated with fragment plant fossils suggest a humid fluvio-lacustrine environment setting in the Ordos Basin during the early period of the Jurassic.     

A Possible Long-tailed Bird with a Pygostyle from the Late Mesozoic Yixian Formation, Western Liaoning, China

AbstractAn incomplete caudal vertebral series (IVPP Vl1309) from the Yixian Formation of late MesozoicJianshangou area of Beipiao, western Liaoning Province, may represent a new bird. The tail is composed of at least 12 free caudal vertebrae and the most distal 5 caudal vertebrae co-ossified into a pygostyle. The pygostyle is plate-like and slightly curved dorsally. The anterior free caudals are amphiplatyan. The anterior caudal surfaces of the last three free caudals are concave, but their posterior articular surfaces are convex. The pygostyle is regarded as the first appeared flight apparatus during the evolutionary process from Archaeopteryx to neornithes. The pygostyle appeared in most early fossil birds and almost all the modem birds. Although their morphologies are different, they are basically formed by at least four last caudal vertebrae. The specimen V11309 is regarded as a bird rather than a non-avian theropod dinosaur based on the following characters: short caudal vertebrae, numerous pits present on the surfaces of the centra and, a foramen present between the basal part of the fused neural spines, which is similar to that of Struthio camelus. The discoveries of pygostyles from the therizinosauroids and oviraptorosaurs may provide strong evidence for supporting the origin of birds from small theropod dinosaurs. The structure of the pygostyle in specimen Vl1309 is different from those of Beipiaosaurus (Therizinosauroid) and Nomingia (oviraptorosaur). The most parsimonious interpretation is that these pygostyle-like structures are independently acquired by Beipiaosaurus and Nomingia during their evolutionary process.     

Early Jurassic basal sauropodomorpha dominated tracks from Guizhou,China: Morphology,ethology, and paleoenvironment

The newly discovered large(350 m^2) Yantan dinosaur tracksite, in the Lower Jurassic Ziliujing Formation of Guizhou Province, China, reveals at least 250 footprints of which ~97 can be resolved into trackways of sauropodomorphs. All the trackways are sub parallel likely indicating gregarious behavior. One theropod track(cf. Grallator) was recorded. The sauropodomorph tracks predominantly represent quadrupedal progression(Morphotype A), and footprint morphology is similar to the ichnospecies Liujianpusshunan, characterized by outward pes rotation. Three trackways indicate bipedal progression, and two of these(Morphotype B) indicate inward pes rotation, accompanied by elongate pes digit scratch marks. For the latter phenomenon three possible scenarios are discussed:(1) significant rotation changes accompanying changes in gait,(2) swimming behavior,(3) formation of undertracks.Sedimentological evidence indicates the tracks were made on a linguloid rippled, muddy, immature sandstone substrate characterized by significant differences in substrate consistency across the trackbearing surface. Microbially induced sedimentary structures(MISS) characterized by distinctive wrinkle marks indicate a stressed, probably semi-arid, paleoenvironment that was not conducive to habitation by invertebrate organisms. This is consistent with other evidence that Lower Jurassic sauropodomorph tracks are often associated with semi-arid paleoenvironments.     

Trace Fossils from Lower Cambrian Wusongger Formation in Xiaoerbulake Outcrop,Kalpin Area,Xinjiang

Based on the analysis of trace fossils collected from the typical outcrop of the Lower Cambrian Wusongger Formation in the Kalpin area, ten ichnospecies of six ichnogenus were identified in the upper member of the Wusongger Formation, with most of them being found for the first time. The trace fossils are described seriatim. The ichnospecies are mainly represented by such common trace fossils as Ophiomorpha nodosa, Helminthopsis hieroglyphica, Helminthopsis ichnosp., Planolites beverleyensis, Planolites vulgaris, Planolites montanus, Palaeophycus striatus, Palaeophycus curvatus?, Cochlichnus anguineus and Rituichnus elongatum. The trace fossils are of high diversity and low abundance, and can be diagnosed as the Cruziana ichnofacies. They can be interpreted as having formed in a shallow water environment.     

Geochemistry of the trace elements and rare-earth elements at the boundary between Cambrian Series 2 and Series 3 at Jianshan, South China: Paleoenvironmental and stratigraphic implications

The Jianshan section in Jianhe County,Guizhou Province,is a very important auxiliary section to the candidate stratotype at the boundary between Cambrian Series 2 and Series 3.Twenty four samples collected from the Jianshan section of the Kaili Formation were analyzed for the concentrations of trace element and rare-earth elements(REEs) by inductively coupled plasma mass spectrometry.The trace element and REE geochemical characteristics of the Kaili Formation at the Jianshan section show that trace element composition and REE distribution patterns across the section are covariant.Notably,at 44 m above the base of the Kaili Formation,most trace element contents and REEs concentrations are lower than those values observed below 44 m.Above 44 m,an increasing trend in the concentration of both trace elements and REEs is observed.These geochemical data indicate important,but subtle,changes at the 44 m horizon.If this is,in fact,verified by work in progress,geochemical criteria can provide additional supports for defining the top surface of Cambrian Series 2,especially in sections where Oryctocephalus indicus is absent.Trace element ratios,specifically Ni/Co,V/Cr,V/(Ni+V) and Th/U as well as Ce and Eu anomalies indicate that the sedimentary environment of the Kaili Formation at the Jianshan section was oxidizing,that benthic redox conditions were less oxic,and may have been influenced by freshwater as compared to the Balang area.Moreover,the Eu anomalies observed in the Kaili Formation are similar to the negative Eu anomalies observed in post-Archean sedimentary rocks,and reveal fluctuations in oxygen content from the bottom to the top of the Kaili Formation.     

Neogene Paleoseismic Events and the Shanwang Biota’s Burial in the Linqu Area, Shandong Province, China

Several paleoseismic events are recorded in the Neogene Linqu Group, exposed in the Linqu area, Shandong Province, China. The events were interpreted on the basis of fieldwork and laboratory analysis, which showed the presence of seismites with plastically deformed soft-sediment deformation structures in the Shanwang Formation, and of seismic volcanic rocks in the Yaoshan Formation which show brittle deformation. The earthquake-triggered soft-sediment deformations in the seismites include load structures, ball-and-pillow structures, flame structures, pillow-like beds, boudinage structures, slump folds, syn-depositional faults, veins of liquefied sand, and dikes of liquefied sandy lime-mud. The seismic activity is also reflected in what might be called ‘brittle seismites'; these originated when, under the influence of seismic vibrations, semi-consolidated conglomerate was shattered. Moreover, volcanic activity is related to intense earthquakes that affected basalts intercalated with sand layers; these successions are known as ‘seismic volcanic rocks', which are characterized by veins of liquefied sand intruding the basalts. All above traces of paleoseismic activity were left from one single time span of 4 Ma with active seismicity that took place 14–10 Ma. This time span is known as ‘the Linqu Neogene Paleoseismic Active Period', which is divided into four paleoseismic episodes, which were responses to tectonic extension and basin rifting in this area. It even includes the activity of the Yishu Fault Zone during the Miocene and the Neogene. The ratios of trace elements in the seismites, w(La)/w(Sc) and w(La)/w(Th) are higher than the average value of the upper crust, but w(Th)/w(Sc) is lower; this is geochemical evidence for the basin rifting that resulted in a high sedimentation rate. The intense and frequent paleo-earthquakes are held responsible for the rapid burial of the Shanwang Biota. Secondary earthquake-induced processes(e.g. slumping of a lake shore and the strongly increased lacustrine sedimentation rate) contributed to the rapid burial of the biota.     

Largest dinosaur tracksite in China(Cretaceous,Zhaojue area,Sichuan Province): On the verge of destruction

The Zhaojue(ZJ) tracksites represent multiple steeply-inclined, track-bearing exposures in the clastic Feitianshan Formation of Sichuan Province that have been sequentially, excavated, mapped, expanded and destroyed, by quarrying,erosion or collapse, for almost three decades. The quarried area extends for more almost 1.0 km from north to south and ～0.5 km from east to west. Four important track-bearing surfaces have been identified and identified as ZJ-I, ZJ-II,ZJ-IIN and ZJ-III, some of which are newly exposed surfaces or expanded surfaces that were previously partially described. Others represent surfaces lost since they were first reported in papers published in 2014–2016. The ZJ-II site represents a hugely expanded continuation of a much smaller track-bearing surface described in 2014. It was mapped using drone technology, and now represents the largest tracksite in China, with a minimum count of 933 recognizable tracks, and the longest recorded sauropod and ornithopod trackways from China(～80 m and ～52 m respectively), which also reveal a pronounced change in the direction of trackmaker movement. The ZJ-II surface represents the best Zhaojue paleo-census sample giving a count of 68 individuals from 61 trackways(37 ornithopod, 10 theropod, 14 sauropod) and isolated tracks(representing 7 individuals). Thus, ornithopods(both large and small trackmakers) represent ～54% of the total number of trackways. The smaller ZJ-III surface was mapped using traditional methods, and reveals at least 6 ornithopod-and 4 theropod trackways. Both the ZJ-II and ZJ-III surfaces reveal parallel ornithopod trackways suggestive of social or gregarious behavior. The combined data from all four Zhaojue surfaces reveal a total of 1928 tracks, and include a few previously reported pterosaurs and theropod swim tracks. The Zhaojue quarry complex provide a good example of multiple track-bearing sites(surfaces) that require long term study and monitoring to extract as much trackway data as possible before in situ physical evidence is lost.     

A New Troodontid (Theropoda: Troodontidae) from the Lower Cretaceous Yixian Formation of Western Liaoning, China

A specimen collected from the Lower Cretaceous Yixian Formation of western Liaoning, China, represents a new genus and species of troodontid theropod. The new taxon is named and described on the basis of the holotype and the only known specimen, which comprises an articulated skeleton with the presacral vertebral, shoulder girdle and forelimbs missing as preserved. Diagnostic features of the new species include nasals that are sinusoid in lateral view, absence of a passage connecting the antorbital and maxillary fenestrae, relatively large teeth, plate-like chevrons forming a band along most of the length of the tail, and a long neck between the femoral head and shaft. The temporal constraints of the three paravian groups (Troodontidae, Dromaeosauridae and Aves) combined with the character distributions among the earliest known troodontids indicate a rapid evolution at the base of the Troodontidae.     

First record of dinosaur trackway from Tibet,China

Three sauropod trackways comprise eight tracks that resemble Brontopodus tracks have been found at the Morong track site in Changdu Prefecture, Tibet, China. These wide- (or sub-wide)-gauge tracks suggest that there was a large sauropod, possibly a member of Titanosauriformes, in Changdu Prefecture during the Early-Middle Jurassic. The sauropod fauna from Changdu Prefecture, Tibet not only has elements in common with the sauropod fauna from the Sichuan Basin, but may include more diverse faunal components.     

Intact theropod dinosaur eggs with embryonic remains from the Late Cretaceous of southern China

In spite of tremendous discoveries of dinosaur eggs and nests over the world, dinosaur embryonic remains are rare. Until today, dinosaur embryos associated with eggshells are only known in about 20 cases, most of which were recovered from broken or crushed eggs. Here we report intact dinosaur eggs in a partial clutch with preserved embryonic remains in eggs from the Upper Cretaceous Nanxiong Formation of Ganzhou in the southern Jiangxi Province of China. Based on macro- and microscopic features of eggs and eggshells, these eggs were identified as Elongatoolithus sp., and possibly belong to oviraptorosaurs. A CT scanning topography revealed that all the eggs contain accumulations of embryonic bones, including possible femur and tibia. Our discovery suggests that dinosaur embryos might be preserved more than we have recognized, because intact eggs potentially confine embryonic remains.     

New perleidid fishes from the Middle Triassic strata of Yunnan Province

Three new genera and species of perleidid fishes from the Middle Triassic strata of Yunnan Province, i.e., Fuyuanperleidus dengi gen. et sp. nov., Luopingperleidus sui gen. et sp. nov., and Diandongperleidus denticulatus gen. et sp. nov., are described in this paper. Fuyuanperleidus dengi is distinguishable from other perleidids by the shape of the maxilla, the big teeth, the ornamentation of the skull bones, maxilla fused with the first infraorbital and the deepened flank scales. Luopingperleidus sui is distinguishable from other perleidids by the following features: the triangular main branchiostegal rays, four horizontal rows of deepened flank scales anterior to the pelvic fin, and three anal scales. Diandongperleidus denticulatus is distinguishable from other perleidids by the following features: the anterior border of some anterior fin rays of both the paired fins and the dorsal and anal fins bearing denticles, and posterior strongly serrated scales. The new discoveries not only add to the diversity of perleidid fishes in South China, but also shed new light on the radiation of Triassic perleidid fishes in the world.