Ecological evolution across the Permian/Triassic boundary at the Kangjiaping Section in Cili County,Hunan Province,China

The stratigraphic sequence of calcimicrobialite facies at the Permian-Triassic (P/Tr) boundary has well recorded the biotic and environmental transition across the end-Permian catastrophic events. The biostratigraphy, microfacies, carbon isotopes, and fossil records across the P/Tr boundary have been studied at the Kangjiaping Section in Cili County, Hunan Province. Three biostratigraphic zones, Palaeofusulina-Colaniella Zone, Hindeodus parvus Zone, and Isarcicella staeschei Zone, are identified. The excursion of δ¹³C_carb exhibits a sharp negative shift in the calcimicrobialite at the P/Tr boundary, which is roughly accordant with the abrupt bioclastic decline. In addition, five types of microfacies are recognized, including algal-foraminifer bioclastic limestone, algal-laminated calcimicrobial limestone, oolitic grainstone, vermiculate limestone, and intraclastic wackstone. The results indicate that the changeover of ecosystem from metazoan reef to calcimicrobialite in Cili is a classic case of marine ecological evolution during the Paleozoic-Mesozoic transition. Supported by National Natural Science Foundation of China (Grant Nos. 40830212, 40621002, 40730209) and National Basic Research Program of China (Grant No. 2006CB80640)     

Geological event sequences of the Permian-Triassic transition recorded in the microfacies in Meishan section

The microfacies sequence in the key interval of Beds 24―29 of the Meishan section comprehensively recorded the geological events during the Permian-Triassic transition, including the anoxia, storm disturbance, hard-ground/firm-ground form, volcanic eruption, weathering input, and microbialite development. This investigation of event sequences on the microfacies provided synthetically some clues to clarifying the previously proposed mechanisms of the end-Permian extinction. The deposit succession in Bed 24 o...     

Depositional model of Early Permian reef-island ocean in Eastern Kunlun

Many fusulinid fossils have been found in thin- to middle-bedded limestones which are distributed between the Early Permian limestone hills and formerly considered as Early Triassic. The fusulinid fossils, identified asNeoshwagerina sp.,Verbeekina sp. andSchwagerina sp., can also be found in massive limestone hills. At the same time, Early Permian radiolarian chert of deep basin facies was discovered in Animaqing. All the above show that the massive limestone hills, thin- to middle-bedded limestones and radiolarian chert belong to syndeposits in Early Permian ocean. The sediments in the study area can roughly be divided into three types: shallow facies, basin facies and transitional facies. The carbonate buildup can be subdivided into massive bioclastic limestone and reef framestone. Basin facies contains thin- or middle-bedded limestone, abyssal red mudstone or ooze, blue-green mudstone and radiolarian chert. Transitional facies includes reef talus and platformal skirt facies. The Early Permian ocean in Eastern Kunlun is recognized as a kind of reef-island ocean environment according to distribution and composition of different facies. The reef-island ocean in Eastern Kunlun is characterized by reef islands (or carbonate buildups) alternating with basins, complicated sea-floor topography, sharp facial change and well-developed reefs.     

Sedimentary facies of Carboniferous–Permian mid-oceanic carbonates in the Changning–Menglian Belt, West Yunnan, Southwest China: Origin and depositional process

Huge carbonate rock bodies ranging in age from the Visean (Middle Mississippian/Early Carboniferous) to the Changhsingian (Lopingian/Late Permian) overlie a basaltic basement in the Changning–Menglian Belt, West Yunnan, Southwest China. These carbonates lack intercalations of terrigenous siliciclastic material throughout. These lines of evidence indicate that they formed upon an isolated and continuously subsiding mid-oceanic island (or plateau), probably of hotspot origin. The carbonates are grouped into a shallow-water carbonate platform facies regime observed in the Yutangzhai section and a relatively deep-water carbonate slope facies regime typically represented in the Longdong section. These two facies regimes developed contemporaneously as parts of a carbonate depositional system on and around a mid-oceanic volcanic edifice. The carbonate platform is subdivided into four facies, including platform-margin, shoal, lagoon, and peritidal facies. Along the measured Yutangzhai section of the platform facies regime, the vertical facies succession from the platform-margin facies into inner-platform facies such as the shoal and lagoon facies is recognized. This facies succession is explained as resulting from the progradation of the carbonate platform. Worm tubes occur as a main reef builder in platform-margin facies of the Mississippian. Their occurrence as major constituents in a high-wave-energy reef is peculiar to Carboniferous reef distributions of the world. The occurrences of other reef- and/or mound-building organisms and peritidal dolo-mudstone are almost consistent in timing with those of Panthalassan counterparts such as the Akiyoshi and Omi limestones of Japan, and probably exhibit the worldwide trend.     

Two episodes of C-depletion in organic carbon in the latest Permian: Evidence from the terrestrial sequences in northern Xinjiang, China

New analyses reveal two intervals of distinctly lower δ¹³C values in the terrestrial organic matter of Permian–Triassic sequences in northern Xinjiang, China. The younger negative δ¹³C_org spike can be correlated to the conspicuous and sharp δ¹³C drops both in carbonate carbon and organic carbon near the Permian–Triassic event boundary (PTEB) in the marine section at Meishan. The geochemical correlation criteria are accompanied by a magnetic susceptibility pulse and higher abundances of distinctive, chain-like organic fossil remains of Reduviasporonites.The older negative δ¹³C_org spike originates within a latest Permian regression. Significant changes in organic geochemical proxies are recorded in the equivalent interval of the marine section at Meishan. These include relatively higher concentrations of total organic carbon, isorenieratane, C₁₄–C₃₀ aryl isoprenoids and lower ratios of pristane/phytane that, together, indicate the onset of anoxic, euxinic and restricted environments within the photic zone. The massive and widespread oxidation of buried organic matter that induced these euxinic conditions in the ocean would also result in increased concentrations of ¹³C-depleted atmospheric CO₂. The latest Permian environmental stress marked by the older negative δ¹³C_org episode can be correlated with the distinct changeover of ostracod assemblages and the occurrences of morphological abnormalities of pollen grains. These observations imply that biogeochemical disturbance was manifested on the land at the end of the Permian and that terrestrial organisms responded to it before the main extinction of the marine fauna.     

徐州大北望地区馒头组化石碳酸盐岩微相分析

江苏省徐州市大北望剖面早寒武世馒头组的碳酸盐岩地层发育较好,沉积类型多样,成岩特征明显,是研究碳酸盐岩沉积及成岩作用的理想剖面之一。文章采用微相分析的方法 ,对该剖面碳酸盐岩的生物组合、颗粒成分、基质类型及其结构等微相特征做了研究。根据颗粒类型、沉积结构和化石,本区的碳酸盐岩可以被分为3个大类和10种化石碳酸盐岩微相类型:泥晶颗粒灰岩(微相1至微相5),颗粒泥晶灰岩(微相6至微相8)和泥晶灰岩(微相9和微相10)。每一种微相对应特定的沉积环境及演化特征。综合分析表明,研究区下寒武统形成于泻湖、局限台地、开阔台地和台地边缘的浅滩环境,据此建立了沉积相模式。在徐州大北望地区进行的关于碳酸盐岩微相的研究表明早寒武世晚期的馒头组继承了早寒武世中期的特点,且海侵持续扩大的持续沉降活动。     

物探方法在临清坳陷某区块山西组沉积相分析中的应用

临清坳陷某区块缺少岩心、薄片、录井及常规测井资料。为了研究其目的层山西组沉积相对储层的控制作用,指导非常规天然气的高效开发。通过利用较少的录井岩性统计资料,结合地震物探方法,分析地震剖面特征、优选地震均方根振幅及相对波阻抗属性,确定该区目的层沉积相。通过上述地震属性研究的物探方法,确定山西组储层为河控浅水三角洲相和碳酸盐台地相,主要发育分流河道、分流间湾及局限台地沉积微相。其中三角洲相是主要的沉积相类型,分流河道是有效储层发育的主要微相类型。      

Anachronistic facies in the Lower Triassic of South China and their implications to the ecosystems during the recovery time

The end-Permian mass extinction not only severely distressed the Paleozoic ecosystems but also dramatically changed the sedimentary systems, resulting in a peculiar Early Triassic ecosystem and submarine environment during the recovery time following the mass extinction. The Lower Triassic is characteristic of the wide occurrence of various distinctive sediments and related sedimentary structures, such as flatpebble conglomerates, vermicular limestone, subtidal wrinkle structures, microbialite, carbonate seafloor fans, thin-bedded limestone and zebra limestone-mudstone. These sediments were common in the Precambrian to Early Ordovician marine settings, and then they occurred only in some extreme and unusual environments with the expansion of metazoan faunas. However, the Early Triassic witnessed an "anachronistic" reappearance of some distinctive sedimentary records in normal shallow marine settings. The study of these anachronistic facies should be of great importance for the understanding of the unique ecosystem and marine environment through the great Paleozoic-Mesozoic transition. The anachronistic facies characterized by vermicular limestone have been documented in many localities in South China and occur at various horizons of the Lower Triassic. Most types of re- ported distinctive sediments over the world have been observed in the Lower Triassic of South China. This provides an excellent opportunity for understanding the Early Triassic environment and its co- evolution with the biotic recovery. Among the anachronistic facies the vermicular limestone is the most characteristic and common distinctive sediments in the Lower Triassic of South China but has received relatively few investigations. Taking it as a case study, we will detail the variation of vermicular limestone and its stratigraphic distribution in the Three Gorges area, Hubei Province. The investigation on the vermicular limestone and other distinctive sediments from the Lower Triassic of South China further indicates that the appearance of anachronistic facies immediately following the mass extinction and the elimination from normal shallow marine facies with the radiation of Mesozoic marine faunas imply the natural response of the sedimentary systems and ecosystems to the great Paleozoic-Mesozoic transitional events and their induced harsh environments. Therefore, the ups and downs of the anachronistic facies may act as a proxy for the evolution of ecosystems independent of fossil analyses.     

Depositional model of Permian Luodianian volcanic island and its impact on the distribution of fusulinid assemblage in southern Qinghai, Northwest China

Pan-riftizational tectonic activity reached climax at Luodianian (Permian) in the East Tethyan Domain, Qinghai-Tibet Plateau. Because of eruptive volcanics and influence of terrigenous materials, a complex volcanic-sedimentary landform formed on the sea floor in southern Qinghai. Four sedimentary facies types were recognized based on detailed field mapping. Spatially, platform facies volcanic-limestone type was located at the center belt approximately trending NWW, surrounded by shallow water slope facies tuff/tuffite type at the two flanks and deep water slope facies breccia/calcirudite at the most outside. The depression facies sandstone-mudstone type, which comprised mainly mudstone, de-posited between volcanic islands (platform facies volcanic-limestone type). Based on the field map-ping and stratigraphic section data, seven rift-related sedimentary facies were recognized and a depo-sitional model for volcanic island was proposed. It is revealed that some volcanic island chain formed quickly and intermittently in the Qamdo Block during violent eruption, and small carbonate reef, shoal, platform occurred above or on edge of volcanic island, and some slope sedimentary facies surrounded volcano island chain during dormant period of volcanic activities. Three types of fusulinid assemblages were distinguished in the carbonate rocks, which deposited in varied positions of a palaeo-volcanic island: (1) Misellina-Schwagerina assemblage occurred above or on edge of volcanic island, (2) Para-fusulina assemblage was located at restricted depression facies among volcanic islands or carbonate platform, and (3) the reworked Pseudofusulina-Schwagerina assemblage occurred at slope facies near margin of volcanic island, which originally deposited in the shallow-water carbonate platform, then collapsed along the volcanic island margin with fusulinid-bearing grain-supported carbonate con-glomerate or calcirudite, and finally re-deposited on the deeper slope. The sedimentary sequence re-sulting from calm shallow water was deposited at the interior of the Qamdo Block from the Devonian to early Early Permian. At the beginning of the peak period of activity of pan-riftzation (Luodianian), al-ternate volcanic island and shallow marine environment within continent crust came into being. Uni-form and stable shallow-water carbonate platform was formed during the Xiangboan. This suggested that the activity of rift basin was evidently weakened. Subsequently the instability of the basin appre-ciably increased with the occurrence of basalt in late Kuhfengian. At last the whole Qamdo Block turned into the closure period of rift during the Late Permian.     

Lithostratigraphy of Permian marine sequences, Khao Pun Area, central Thailand: Paleoenvironments and tectonic history

Geologic mapping and subsurface lithostratigraphic investigations were carried out in the Khao Pun area (4 km²), central Thailand. More than 250 hand specimens, 70 rock slabs, and 70 thin sections were studied in conjunction with geochemical data in order to elucidate paleoenvironments and tectonic setting of the Permian marine sedimentary sequences. This sedimentary succession (2485 m thick) was re‐accessed and re‐grouped into three lithostratigraphic units, namely, in ascending order, the Phu Phe, Khao Sung and Khao Pun Formations. The Lower to lower Upper Permian sedimentary facies indicated the transgressive/regressive succession of shelf sea/platform environment to pelagic or abyssal environment below the carbonate compensation depth. The sedimentological and paleontological aspects, together with petrochemical and lithological points of view, reveal that the oldest unit might indicate an Early Permian sheltered shallow or lagoonal environment. Then the depositional basin became deeper, as suggested by the prolonged occurrence of bedded chert‐limestone intercalation with the local exposure of shallower carbonate build‐up. Following this, the depositional environment changed to pelagic deposition, as indicated by laminated radiolarian (e.g. Follicucullus sp.) cherts. This cryptic evidence might indicate the abyssal environment during middle Middle to early Late Permian; whereas, previous studies advocated shelf‐facies environments. Following this, the depositional condition might be a major regression on the microcontinent close to Indochina, from the minor transgressive/regressive cycles that developed within a skeletal barrier, and through the lagoon with limited circulational and anaerobic conditions, on to the tidal flat to the sheltered lagoon without effective land‐derived sediments.     

End-Permian conodont fauna from Dongpan section: Correlation between the deep-and shallow-water facies

This paper systematically investigated the conodonts from the uppermost Permian to the Lower Triassic at the Dongpan Section, Southern Guangxi, South China, and obtained abundant Late Permian conodonts from the syndepositional limestone lenses of beds 3 and 5-2 at this section. One genus and eight species of conodont P1 element including one new species, Neogondolella dongpanensis sp. nov., have been identified. The feature of conodont fauna indicates that conodonts collected from beds 3 and 5 at the Dongpan Section belong to the Neogondolella yini conodont zone, and correspond to bed 24 at the Meishan Section. Based on these conodont data, we suggest that the Neoalbaillella optima radiolarian zone at the Dongpan Section at least extended to the upper part of the N. yini conodont zone.     

Microbialite of anoxic condition from Permian-Triassic transition in Guizhou,China

Spherical microfossils are present in the Hindeodus parvus zone of the Lower Triassic in Ziyun,Guizhou Province. They generally range from 0.15 to 0.30 mm across,with micritic wall and filled by micro-sparry calcites,and are evenly scattered in micritic matrix. Their abundance makes the rock as-signed to microbialites. The accompanied organisms include ostracods and algal mat,but no gastro-pods or bivalves. Presence of small (<7 μm) pyrite framboids indicates that this bed formed in anoxic conditions. In some sections,this bed is overlain and underlain by tidal-flat micritic limestone with microgastropods and small burrows. Occurrence only in deposits on reef top indicates that the micro-bial organism was benthic,and needed sunlight in life. The size of the microbial fossil exceeds that of any bacteria or cyanobacteria. Thus,it does not belong to these two phyla. It may belong to lower green algae,and is assigned to a new species in a new genus,Ziyunosphaeridium sinensis gen. et sp. nov. Up to now,the rocks described as microbialites from the Permian-Triassic transition include six types: (1) porous micritic limestone such as that descried from Japan by Sano and Nakashima (1997),(2) limestone with rich globular microfossils such as that described from the Nanpanjiang Basin in China by Lehrmann (1999),(3) dendroidal limestone such as that described from the vicinity of Chongqing by Kershaw et al. (1999),(4) spherical microbial limestone adapted to anoxic environments described from Ziyun,Guizhou reported in this paper,(5) limestone with cyanobacterial fossils such as that described by Wang et al. (2005),and (6) stromatolites. All these microbialites are not reefs in the proper sense,and the argument that Permian reef ecosystems extended into the Mesozoic is incorrect.     

Fusulinoidean faunal succession of a Paleo–Tethyan oceanic seamount in the Changning–Menglian Belt, West Yunnan, Southwest China: An overview

Abstract   Fusulinoidean faunal succession from Paleo–Tethyan seamount-type carbonates of the Yutangzhai section in the Central zone of the Changning–Menglian Belt of West Yunnan, Southwest China, is presented for the first time. The Changning–Menglian Belt is one of the orogenic belts that represent the closed main Paleo–Tethys in East Asia. The Yutangzhai section is represented by basalts and overlying carbonates, about 1100 m thick. It exhibits a continuous faunal succession composed of 17 fusulinoidean assemblages ranging from the Serpukhovian (late Mississippian/late Early Carboniferous) to Midian/Capitanian (late Middle Permian/late Guadalupian). No significant faunal break can be recognized in this section. The generic and some specific composition of the Yutangzhai assemblages indicates that the faunal succession is similar to those observed in Tethyan and Panthalassan areas and is of tropical Tethyan type although their generic diversity is definitely lower than those of Paleo–Tethyan shelves, such as South China, Indochina, and Central Asia. Throughout the Yutangzhai section, the carbonate rocks are essentially massive, very pure in composition, and devoid of terrigenous siliciclastic inputs. These lithologic characters are identical to those observed in accreted shallow-marine carbonate successions of seamount origin in Permian and Jurassic accretionary complexes of Japan, for example the Akiyoshi Limestone. This evidence further demonstrates the seamount origin of the basalt–limestone succession in the Central zone of the Changning–Menglian Belt from the viewpoint of lithofacies. In middle Mississippian (middle Early Carboniferous) time, oceanic submarine volcanism that was probably related to hot spot activities formed a number of seamounts and oceanic plateaus. It was active not only in the Panthalassa, but also in the Paleo–Tethys.     

Size variation of the end Permian conodont <Emphasis Type="Italic">Neogondolella</Emphasis> at Meishan Section,Changxing, Zhejiang and its significance

This study is based on both a generic and species level investigation of the individual size of the latest Permian conodont Neogondolella Pa elements collected from Meishan Section A, Changxing, Zhejiang Province. In this study, an obvious size reduction of Neogondolella Pa elements within bed 24e of the upper Changxing Limestone is recognized. The size variation of the Neogondolella occurs simultaneously with some important events including the negative shift of δ ¹³C, change in the ratio of the abundance of cyanobacterial biomarkers versus that of other general bacterial biomarkers and the shallowing of the sea water. Through the investigation of the paleoenvironmental changes and the analysis of the paleoecology of the conodont genus Neogondolella, the authors propose that the major factors for the size reduction of species of the conodont genus Neogondolella are food shortages caused by the mass extinction, the shallowing of the sea water as well as the increase in opacity of the sea water during the end Permian. The same phenomenon of Neogondolella size reduction is also observed in preliminary research from the same horizon at Shangsi Section, Sichuan Province. All the evidence suggests that there was a mass extinction that occurred at the horizon of bed 24e, and the evidence supports the viewpoint of a multi-phase mass extinction during the Permian and Triassic transition in South China.     

Mega-shoaling in carbonate platform of the Middle Triassic Leikoupo Formation,Sichuan Basin,southwest China

Shoaling is a common type of sedimentation in the evolution of carbonate platform,and commonly has poor continuity.This paper presents a newly discovered and rare type of shoaling,i.e.,mega-shoaling in nearly basin scale,which is developed in the Middle Triassic Leikoupo Formation of the Sichuan Basin,southwest China.During the Leikoupo time,the studied Sichuan Basin experienced hot and dry climate conditions and developed a carbonate platform within a restricted epicontinental sea.In B sub-layer of the Lei-1-1 sub-member of the Leikoupo Formation a series of grainstones of shoal facies accumulated throughout almost the entire basin,thereby generating features associated with basin-scale mega-shoaling.By detailed core examination and microscopic observation of thin sections,it is shown that the lithology of this set of grainstones is dominated by doloarenite(calcarenite)followed by oolitic dolomite(limestone).In addition,it contains three types of sedimentary sequences characterized by upward-coarsening and upward-shallowing as the followings:restricted lagoon to platform interior beach;restricted lagoon to platform interior beach and to platform flat;and tidal flat to peritidal beach.Subsequently,a multicyclic stratigraphic division and correlation revealed that this set of grainstones can be well traced and compared horizontally,and is generally isochronous.In addition,a template for logging facies,established based on core calibrations and logging data,was employed to analyze the 235 wells in the basin.The results demonstrate the shoal grainstones to be 10–40 m thick with a15×104km2continuous distribution area.These findings indicate that the carbonate platform developed mega-shoals within a short period of time.The genesis of such a mega-shoaling was investigated by focusing on various shoaling conditions,such as paleo-tectonics,paleo-geomorphology,paleo-climate,sea-level changes,and palaeo-hydrodynamics.A specific combination of independent geological factors creates beneficial geomorphologic conditions for the mega-shoaling including a quiescent paleo-tectonic environment,relatively flat paleo-geomorphology and evaporites filling up and leveling off.In addition,a stably settling carbonate platform underwent sea-level fluctuations through swift transgressions and protracted regressions,which is not only conducive to continuous,multicyclic and superimposed vertical development of grain beaches but also beneficial for the horizontal migration,coalescence and superimposition of individual grain beaches.As a consequence,large-scale and continuously-distributed grain beach sedimentation emerges and mega-shoals develop.     

泰国Khorat盆地二叠纪石灰岩的重磁化研究

泰国Ｋｈｏｒａｔ盆地西部的晚二叠世石灰岩的古地磁研究表明磁铁矿为稳定剩磁的主要载体多组分磁分量分离技术揭示了高温磁组分（或高矫顽力）具有呈对分布的正、反极性．但是，应用逐渐展平岩层法可以发现各采样点的平均特征磁化方向在岩层展平至３０％时．磁化方向最为集中．这一发现表明二叠纪石灰岩中所揭示出的磁化方向很可能形成于褶皱（期）过程中．野外观察表明，二叠纪石灰岩在印支期发生强烈褶皱并被晚三叠世湖相石灰岩角度不整合覆盖．所以二叠纪石灰岩的重磁化很可能发生在中、晚三叠世的印支期．这些石灰岩样品切片后经显微镜、扫描电镜和电子探针分析，次生磁铁矿多数与方解石微晶和铁质碳酸钙粒共生，且多分布在方解石脉附近．重磁化很可能是由于印支期造山运动时，铁质碳酸钙受碳水化合物流体的蚀变作用所引起的。     

Size variation of the end Permian conodont Neogondolella at Meishan Section,Changxing,Zhejiang and its significance

Recent works reveal that the size reduction of or- ganisms during the Permian-Triassic transition is of great importance for research on biotic mass extinction and recovery in this period[1,2]. Initially, Urbanek (1993) observed a decrease in the size of Silurian graptolites during the biotic crisis, and used the term “Lilliput Effect” to define the size reduction in sur- viving animal taxa[3]. Subsequently, researchers re- ported a decrease in size of late Devonian conodonts[4], heart urc…     

Timing of the terrestrial Permian-Triassic boundary biotic crisis: Implications from U-Pb dating of authigenic zircons

The Late Permian to Early Triassic transition represents one of the most important Phanerozoic mass extinction episodes. The cause of this event is still in debate between catastrophic and gradual mechanisms. This study uses the U-Pb method on zircons from the uppermost Permian/lowermost Triassic clay deposits at Chahe (Guizhou Province, SW China) to examine time constraints for this event. The results of both this and previous studies show that the ages of Bed 68a and 68c (the upper clay bed of the terrestrial Permian-Triassic boundary (PTB)) respectively are 252.6±2.8 and 247.5±2.8 Ma. This age (within the margin of error) almost accords with the upper clay bed (Bed 28) age of Meishan and the eruption age of Tunguss Basalt, and is so far the most accurate age obtained from terrestrial PTB. The claystone of Bed 68 was formed in the earliest Triassic. The biotic crisis occurred at nearly the same time in terrestrial and marine environments during Permian-Triassic interval; however the extinction patterns and processes are different. The extinction pattern of the terrestrial plants shows a major decline at the PTB after long-term evolution, followed by a retarded extinction of the relicts in the earliest Triassic.     

Community replacement sequences and paleoenvironmental changes in reef areas of South China from Late Permian to Early Triassic exemplified by Panlongdong section in northeastern Sichuan Basin

The well-known Permian Changhsingian calcisponge reef located at Panlongdong section,Xuanhan county,northeastern Sichuan Basin has attracted wide attention.Due to severe dolomitization and poor quality of the fossils,the P-T boundary in this section is difficult to determine.This study,for the first time,recognized six communities in the Upper Permian Changhsingian Changxing Formation through the Lower Triassic Induan Feixianguan Formation of the Panlongdong section.They are Bryozoan-Archaeolithoporella-calcisponge Community,Calcareous green algae-foraminifer-crinoid Community,Microgastropod-foraminifer Community,Ostracod-microgastropod-cystic microbe Community,Ostracod-small brachiopod Community,and Non-calcified cyanobacteria Community.By using community replacement and palaeoenvironmental analysis,for the first time,we set the P-T boundary of the Panlongdong section at the middle of the calcimicrobialite containing cystic microbes.The community replacement sequence in the Panlongdong section is similar to that in other contemporaneous sections in reef areas of South China,indicating universal palaeoenvironmental changes during the Permian-Triassic transition.The results show that:(1)Changhsingian calcisponge reefs in South China generally vanished before the mass extinction and may be related to the large regression in the Late Permian.(2)The Calcareous green algae-foraminifer-crinoid Community replaced reef community and continued till the mass extinction.The mass extinction was probably related to the global sea-level drop.(3)The first aftermath community was dominated by specialized microgastropods,followed by the microbes,and then by the specialized microgastropods and small brachiopods.The succession of the three communities reflected the change in environmental conditions from dysoxic to anoxic and again to dysoxic.(4)In the Early Triassic,the relict community in shallow sea had very low diversity and low abundance,and was dominated by crinoids,gastropods,and bivalves.     

An accurately delineated Permian-Triassic Boundary in continental successions

The Permian-Triassic Boundary Stratigraphic Set (PTBST), characteristic of the GSSP section of Meishan and widespread in marine Permian-Triassic Boundary (PTB) sequences of South China, is used to trace and recognize the PTB in a continental sequence at Chahe (Beds 66f―68c). Diversified Permian plant fossils extended to the PTBST, and a few relicts survived above that level. Sporomorphs are dominated by fern spores of Permian nature below the PTBST, above which they are replaced by gymnosperm pollen of Triassic aspect. In the nearby Zhejue Section, the continental PTBST is charac- terized by the fungal 'spike' recorded in many places throughout the world. The boundary claybeds (66f and 68a,c) of the PTBST are composed of mixed illite-montmorillonite layers analogous with those at Meishan. They contain volcanogenic minerals such as β quartz and zircon. U/Pb dating of the upper claybed gives ages of 247.5 and 252.6 Ma for Beds 68a and 68c respectively, averaging 250 Ma. In con- trast to the situation in Xinjiang and South Africa, the sediment sequence of the Permian-Triassic tran- sition in the Chahe section (Beds 56―80) become finer upward. Shallowing and coarsening upward is not, therefore, characteristic of the Permian-Triassic transition everywhere. The occurrence of relicts of the Gigantopteris Flora in the Kayitou Fm. indicates that, unlike most marine biota, relicts of this pa- leophytic flora survived into the earliest Triassic. It is concluded that Bed 67 at Chahe corresponds to Bed 27 at Meishan, and that the PTB should be put within the 60-cm-thick Bed 67b④, now put at its base tentatively. This is the most accurate correlation of the PTB in continental facies with that in the marine GSSP.