宁城中生代道虎沟生物群中两类具原始羽毛翼龙的发现及其地层学和生物学意义

本文实事求是地介绍了道虎沟生物群的组成及相关地层的岩性,并根据长尾型嘴口龙类(Rhamphorhynchoidea)翼龙与短尾型翼手龙类(Pterodactyloidea)翼龙同期共生的特点,确信道虎沟生物群的地质时代为晚侏罗世。此外,本文还研究了道虎沟生物群中翼龙身上发育的皮肤衍生物,初步认为它们是原始羽毛(protofeathers),至少是与羽毛同源的(feather-homologous),这对于了解羽毛(广义)的演化历史和早期的形态结构将具有十分重要的意义。     

The Jurassic orthopteran Allaboilus gigantus Ren and Meng, 2006 (Prophalangopsidae) from Beipiao,Northeast China and its biostratigraphical significance

The orthopteran Allaboilus gigantus Ren and Meng, 2006 (Prophalangopsidae) is reported based on a male forewing from the Middle-Upper Jurassic Haifanggou Formation of Beipiao, western Liaoning, China. This discovery greatly extends the distribution of the species from Daohugou in Inner Mongolia eastwards to Beipiao in western Liaoning. It further provides new evidence for correlation of the Daohugou Beds with the Haifanggou Formation indicating a Callovian-early Oxfordian age for the Daohugou Beds.     

A New Elaterid from the Middle Jurassic Daohugou Biota (Coleoptera: Elateridae: Protagrypninae)

A new genus and species of fossil elaterid namely Clavelater ningchengensis gen. et sp. nov. assigned to the subfamily Protagrypninae based on one specimen from the Middle Jurassic Jiulongshan Formation near Daohugou Village, Ningcheng County, Inner Mongolia, northeastern China is described and illustrated. This specimen displays a pair of large antennae that easily differs from those of most Mesozoic elaterids. There are around 36 species within 20 genera referred to elateroids described from the Early Jurassic to Miocene of China, with another three species within three genera reviewed, but the taxonomic assignment of at least seven species within three genera is doubted. The systematic position of fossil elateriform beetles reported from China is reviewed and the evolution of Mesozoic elateroids briefly discussed.     

中国中侏罗世始水虻科莎水虻属二新种记述

我国中生代始水虻科昆虫多样性非常丰富,本文新记录了来自内蒙古宁城道虎沟中侏罗世地层莎水虻属 Sharasargus 下的两个新种：长脉莎水虻 Sharasargus longivenus Wang, Feng et Wang, sp. nov. 和道虎沟莎水虻 Sharasargus daohugouensis Wang, Feng et Wang, sp. nov. 。其中,长脉莎水虻具有较长的Rs和bM1+2与其他种明显分开,而道虎沟莎水虻的R5明显延长与其他种明显区分;此外,本文还编制了莎水虻属分种检索表。     

对《自然》、《科学》杂志报道的热河生物群、道虎沟生物群重大发现与研究进展的述评

20世纪90年代以来,中国辽宁西部及其邻区晚中生代热河生物群和道虎沟生物群研究取得了令世人瞩目的成果,尤其是在鸟类起源及其早期分异、羽毛起源及其早期演化模式、真兽类起源与早期兽类的生态辐射、被子植物起源等研究方面获得了许多重大进展。截至2006年底,共有46篇关于热河生物群的研究论文以及3篇道虎沟生物群的研究论文发表在世界著名的《自然》和《科学》杂志上,其中第一作者单位为中国科研机构的论文数量分别达39篇和2篇。《自然》和《科学》杂志是世界上自然科学领域两个国际性学术周刊,其覆盖范围广、传播速度快、读者群广泛,其连续刊登热河生物群和道虎沟生物群的研究论文,使这些重要研究成果在短时期内为全世界的学者所认可和接受,为推动该两个重要生物群的研究发挥了重要的作用。     

粗壮假碾磨齿兽的发现及其科学意义

本文简要回顾了蜀兽类哺乳动物的研究历史,论述了在哺乳动物牙齿演化和中生代陆相地层对比研究中的科学意义。粗壮假碾磨齿兽的发现有力地支持了哺乳动物中“三磨楔齿形臼齿趋同演化”的假设,并再一次以直接的化石证据表明内蒙古宁城道虎沟地区含蜀兽类化石地层的时代为中侏罗世。     

A new species of Eomeropidae (Insecta: Mecoptera) from the Middle Jurassic of China

The Eomeropidae is a species-poor family with only 11 known fossil species from the Early Jurassic to Palaeogene. A new species of fossil eomeropid, Tsuchingothauma gongi sp. nov., is described based on a well-preserved wing from the Middle Jurassic Daohugou deposits of Inner Mongolia, China. Our new species is distinguished from the type species T. shihi mainly in having very numerous crossveins and cells; more longitudinal veins: 13 and 11 terminal branches in radial sector and medial field, CuA with 3 terminal branches and A with 7 terminal branches.     

内蒙古宁城道虎沟地区首次发现中侏罗世蝌蚪化石

中国中生代的两栖类化石十分稀少,已报道的有尾两栖类仅见产于河北丰宁县凤山炮樟沟晚侏罗世的东方塘螈(Laccotriton subsolanus)、凤山中华螈(Sinerpeton fengshanensis)、辽宁葫芦岛市水口子早白垩世的钟健辽西螈(Liaoxitriton zhongjiani)、内蒙古宁城道虎沟地区中侏罗世的天义初螈(Chunerpeton tianyiensis)和奇异热河螈(Jeholotriton paradoxus),以及辽西早白垩世热河生物群中的无尾两栖类葛氏辽蟾(Liaobatrachus grabaui)、三燕丽蟾(Callobatrachus sanyanensis)北票中蟾(Mesophryne beipiaoensis)。本文记述了一件采自内蒙古宁城道虎沟地区中侏罗世的蝌蚪化石,这在中国乃至亚洲尚属首次报道,不仅填补了中国中侏罗世无尾两栖类化石分布的空白,而且进一步丰富了燕辽生物群的内容,对早期蛙类的地理分布、形态发生、生存环境等研究具有重要的科学价值。     

Current Knowledge of Mesozoic Coleoptera from Daohugou and Liaoning (Northeast China)

<正>The present paper is devoted to an overview on fossil Coleoptera studied from Inner Mongolia, Daohugou(Middle Jurassic,Jiulongshan Formation)and Liaoning(Upper Jurassic-Lower Cretaceous,Yixian Formation)deposited in Chinese collections.As a result,species of the tribe Sperchopsini and Hydrophilini from Hydrophilidae,families and subfamilies Silphidae,Syndesinae from Lucanidae,Pleocomidae,Trogidae, Trogissitidae,Pyrochroidae,Diaperinae from Tenebrionidae,and Cerambycidae were first registered in the Mesozoic and some families were defined as new.It was shown that many superfamilies represented in the Recent Fauna were formed within the Middle Jurassic and Lower Cretaceous.The materials examined confirm the hypothesis that Cucujiformian beetles are a younger group than other infraordera of Polyphaga(Staphyliniformia and Elateriformia)and,therefore,they appeared in the fossil record only in the late Mesozoic.It was shown and confirmed that most superfamilies appeared in the fossil records before Cucujoidea.The synonymy of Notocupes Ponomarenko,1964;Sinocupes Lin,1976,syn.nov.;Amblomma Tan,Ren et Liu 2005,syn.nov.;Euryomma Tan, Ren et Shih,2006,syn.nov.,non Stein,1899 and Ovatocupes Tan et Ren,2006,syn.nov.;synonymy of Tetraphalerus Waterhouse,1901 and Odontomma Tan,Ren et Ge 2006,syn.nov.;and synonymy of Priacmopsis Ponomarenko,1966 and Latocupes Tan et Ren,2006,syn.nov.are proposed.Sinorhombocoleus papposus Tan et Ren,2009 is transferred from the family Rhombocoleidae to Schizophoridae.Cervicatinius complanus Tan,Ren et Shih,2007 and Forticatinius elegans Tan,Ren et Shih,2007 are transferred from the family Catiniidae(suborder Archostemata)to superfamily Cleroidea(suborder Polyphaga:first among the family Peltidae and second as a closely related group to the latter family).The family Parandrexidae is transferred from the superfamily Cucujoidea to Cleroidea.The ecological circumstances of the past ecosystems and hypotheses of historical development of the order Coleoptera are discussed.The age of faunas examined is considered.The list of the taxa described from Daohugou and Liaoning is compiled.     

辽西及其毗邻地区中生代珍稀动、植物化石的分布及层位

近十几年来,辽宁西部及毗邻的内蒙古道虎沟地区下自海房沟组上至孙家湾组中发现了大量的龙鸟等珍稀动、植物化石.这些珍稀化石被分散描述和报道于国内外的各种期刊、文集及专著中,查找起来多有不便,而且有很多珍稀化石的具体产出层位模糊不清,不便于对比研究.为使有关从事科研、教学、科普工作者及广大动、植物化石爱好者对该地区的珍稀化石属种有全面细致的了解,作者在对部分化石产出层位进行了野外考察的基础上,整理出己报道的珍稀化石属种名单,并按自然分类系统排列,每个属种后面附有拉丁学名、产地、时代及层位.     

A New Species of Liadotaulius (Insecta: Trichoptera) from the Middle Jurassic of Daohugou,Inner Mongolia

Diverse caddis flies are recently collected from the Middle Jurassic Daohugou fauna in Ningcheng County, Inner Mongolia, northern China, and different types of caddis cases have been reported in the same locality. A new species of caddisfly, Liadotaulius daohugouensis sp. nov. is described and illustrated; the establishment is based on a well-preserved isolated forewing collected from the Middle Jurassic Jiulongshan Formation of Daohugou. The family status of Liadotaulius is discussed on the basis of new material and is referred to the family Philopotamidae. The Daohugou fauna continues to provide new data for studying the early evolution of Trichoptera.     

A new family of aphids (Hemiptera: Aphidoidea) in mid-Cretaceous Myanmar amber

A new family of aphids, the Isolitaphidae fam. nov. (Hemiptera: Aphidoidea), is described for the new genus and species, Isolitaphis prolatantennus gen. et sp nov., in mid-Cretaceous amber from Myanmar (Burma). The new species is compared with previously described Mesozoic Aphidoidea, including members of the families Parvaverrucosidae and Burmitaphididae that were described earlier from Myanmar amber. The new taxon can be separated from all previously described extant and extinct Aphidoidea by its 10-segmented antenna, the presence of an ovipositor, siphons, a three-branched M vein and forked C vein. The new species adds to the morphological diversity of Mesozoic Aphidoidea.     

Taxonomy and Stratigraphy of Late Mesozoic Anurans and Urodeles from China

Three anurans (Callobatrachus sanyanensis, Liaobatrachus grabaui, Mesophryne beipiaoensis) and six urodeles (Laccotriton subsolanus, Liaoxitriton zhongjiani, Jeholotriton paradoxus, Sinerpeton fengshanensis, Chunerpeton tianyiensis, Liaoxitriton daohugouensis) are reported from the late Mesozoic tuff-interbedded lacustrine deposits (mostly of the Jehol Group) in northeastern China. They document the first discovery of Chinese Mesozoic lissamphibians, and their old geological age, superb preservation condition, and large taxoncmic diversity are unique compared with other findings worldwide. The anurans occupy a higher evolutionary position than typical Jurassic taxa, supporting a post-Late Jurassic age of the fossil horizons. The urodeles all have unicapitate ribs, suggesting an evolutionary grade at the cryptobranchoid level, and are advanced in osteological features over non-urodeles from the Middle and Upper Jurassic in England and Central Asia. Some urodeles (Jeholotriton and Chunerpeton) exhibit neot     

The first Mesozoic palmetto beetle (Coleoptera: Smicripidae) in Upper Cretaceous Burmese amber

Fossil smicripid beetles are very rare. Here we describe and figure the first Mesozoic representative of Smicripidae. Smicrips cretacea sp. nov., a new species belonging to the extant Smicrips LeConte is preserved in the Upper Cretaceous amber from northern Myanmar. Our discovery suggests that Smicrips is an ancient group, originating at least in the earliest Late Cretaceous. Together with other two fossil Smicrips species from the Eocene of Europe, it reinforces the hypothesis that the Recent distribution of Smicripidae is relict, and the family was more widespread in the Mesozoic and early Cenozoic than it is in the Recent.     

A new afrograptid (Diplostraca: Estheriellina) from the Lower Cretaceous of southern England

The Family Afrograptidae is a ‘conchostracan’ group with multiple radial costae reaching to the umbo on their carapaces. It comprises four described genera: Afrograpta, Camerunograpta, Congestheriella and Graptoestheriella with a total of thirteen described species which are occasionally reported from the Jurassic and the Cretaceous in Africa, Europe and South America (i.e. Afrograpta from the Upper Cretaceous of Cameroon; Camerunograpta from the Jurassic to Cretaceous of Cameroon; Congestheriella from the Jurassic to Upper Cretaceous of the Congo Basin, Brazil, Bulgaria, Venezuela and Argentina; and Graptoestheriella from the Upper Jurassic to Lower Cretaceous of Brazil). A new genus and a new species, Surreyestheria ockleyensis gen. et sp. nov., belonging to the Family Afrograptidae from the Lower Cretaceous (lower Barremian) Upper Weald Clay Formation of Ockley Village, Surrey County, southern England is described in this paper. The new genus mainly differs from the other four genera by the special reticulate ornamentation on its carapace. It indicates that the Family Afrograptidae was more diverse and more widely distributed in the late Mesozoic than previously supposed. Afrograptidae is a special branch of Estheriellina the latter originating in the late Palaeozoic and the former in the early Mesozoic. Afrograptids, as a whole had been widespread across Pangea in the Early Jurassic.     

内蒙古宁城县道虎沟叶肢介化石群的时代

内蒙古赤峰宁城县道虎沟含丰富的叶肢介壳瓣及许多珍罕的软体化石 ,计有 1属 4种 :Euestheria ziliu-jingensis Chen,Euestheria haifanggouensis Chen,Euestheria jingyuanensis Chen,Euestheria luanpingensis Shen etNiu。这些分子都是我国中侏罗世早期自流井真叶肢介 (Euestheria ziliujingensis)动物群的重要成员 ,以个体较小、壳瓣圆或椭圆形、生长带具小网状装饰、种类单调、密集群生为特点 ,主要出现于中侏罗世早中期 (Bajocian- MiddleBothonia)。特别是 Euestheria luanpingensis Shen et Niu及其软体化石在道虎沟的发现 ,证明其层位应与冀北滦平九龙山组相当。道虎沟叶肢介群与中侏罗世晚期的假线叶肢介 (Pseudograpta)动物群的面貌相差甚远 ,与比其层位更高的大北沟组的尼斯脱叶肢介 (N estoria)动物群及义县组的东方叶肢介 (Eosestheria)动物群迥然不同 .     

New mantises (Insecta: Mantodea) in Cretaceous ambers from Lebanon,Spain, and Myanmar

Diverse new material of mantises found in the Cretaceous amber-bearing deposits from Lebanon (Barremian), Spain (Albian), and Myanmar (Albian–Cenomanian) are described and figured. The Lebanese and Spanish forms are nymphs; while the one from Myanmar is an adult specimen. The Lebanese nymph corresponds to a new specimen of Burmantis lebanensis Grimaldi, 2003 while the adult Burmese (Myanmar) specimen belongs to the new species Burmantis zherikhini. The Spanish specimen represents a new genus and species and is established as Aragonimantis aenigma, but is considered family incertae sedis. The Spanish specimen is the first record of Mesozoic mantises from western-European amber deposits. A revised phylogenetic hypothesis for Cretaceous mantises is proposed.     

A New Axymyiid Genus with Two New Species from the Middle Jurassic of China (Diptera: Nematocera: Axymyiidae)

The Axymyiidae is one of the small families of the suborder Nematocera within the Diptera.Up to date,three genera and seven species of extant axymyiids have been described from the Holarctic Region,including Canada,China,Hungary,Japan,Russia,and USA,with three genera and three species of fossil taxa described from the Middle Jurassic of Daohugou,northeastern China.A new genus Raraxymyia gen.nov.and two new species,R.parallela and R.proxima gen.et sp.nov.from the Middle Jurassic Jiulongshan Formation of Daohugou Village,Inner Mongolia,China,are herein described based on their well-preserved wings and body characters.A revised key to the genera of axymyiids is provided.     

A New Fossil Scorpionfly (Insecta: Mecoptera: Holcorpidae) with Extremely Elongate Male Genitalia from Northeastern China

Since Holcorpa maculosa was first described in 1878, it has attracted lots of attention due to its extremely elongate abdominal segments from the sixth to the eighth. Fossil records of family Holcorpidae are very rare, comprising two species in one genus represented by three fossil specimens, resulting in limited knowledge on the structure and function of holcorpid’s unique male organ, as well as its evolutionary relationship with other scorpionflies. Herein we describe a new genus with a new species, Conicholcorpa stigmosa gen. et sp. nov., of the Holcorpidae from the Middle Jurassic Jiulongshan Formation near Daohugou Village, Inner Mongolia, China. This is the first holcorpid documented from the Daohugou locality, albeit two previously reported species with extremely elongate male organs but deemed as Family Incertae sedis. C. stigmosa gen. et sp. nov. is the earliest holcorpid hitherto, extending its existence from the Early Eocene to the Middle Jurassic by 114 million years and providing one more early case of presenting exaggerated male body parts for sexual display and/or selection. Our analysis supports the sister relationship between Holcorpidae and Orthophlebiidae. Our results not only enhance our knowledge of the morphology of Holcorpidae but also elucidate the relationship of known species of this family.