Changes in late Quaternary vegetation and insect communities in southwestern Ontario

The Gage Street site in Kitchener, Ontario, is a peat/marl sequence representing continuous lacustrine sedimentation from the time of deglaciation (ca. 13,000 yr B.P.) through 6900 yr B.P. Insect, pollen, and plant macrofossil remains isolated from the sediments indicate that from ca. 13,000 to 12,500 yr B.P. the region was characterized by parkland-tundra vegetation existing within thermal conditions more analogous to those today of the midboreal forest. The transition from parkland to coniferous forest at ca. 12,500 yr B.P. occurred within a climate that was only gradually warming. By the time of the spruce/pine transition at 10,500 yr B.P., an insect fauna had become established that is typical of southwestern Ontario today. The replacement of this fauna at ca. 8400 yr B.P. by one characteristic of the lowlands of the east-central United States represents the beginning of Hypsithermal conditions in southern Ontario. Vegetation and insects indicate that the climate continued to gradually warm through the mid-Holocene.     

Gould Pond, Maine: late-glacial transitions from marine to upland environments

Gould Pond sediments are unusual in North America in that they include a continuous record of change from marine to freshwater conditions during the late-glacial period, with a wide array of micro- and macrofossils deposited during a period of high sedimentation rate. Marine waters, much colder than those in the present Gulf of Maine, covered the site at the time of deglaciation (c. 13,200 BP). Plants characteristic of modem tundra grew on nearby uplands. Marine recession, due to isostatic rebound of the land, occurred from c . 12,800–12,200 BP. The lake water was completely freshened by 12,000 BP. A sparse shrub-herb tundra became established around Gould Pond as marine waters receded. Subsequent to 11,300 BP, sedges and other herbs became more abundant, and willow and Dryas less abundant, signifying increased warmth and decreased frost action. At least six tree species, all now common in the area, arrived around Gould Pond between c . 10,800 and 10,500 BP. This rapid transition was coincident with the most rapid major non-anthropogenic change of vegetation at sites across eastern North America during the postglacial period.     

Boreal peatland margins as repository sites of long-term natural disturbances of balsam fir/spruce forests

A multidisciplinary, high-resolution paleoecological study (Lepidoptera and plant remains, macroscopic charcoal, pollen) was conducted on a 4000-yr peat monolith extracted from the margin of an ombrotrophic peatland on Anticosti Island (Gulf of St. Lawrence, eastern Canada) to reconstruct the long-term natural disturbances (insect outbreaks, forest fires) of a balsam fir/spruce forest. We hypothesized that an activity of insect defoliators (spruce budworm, hemlock looper) was the main disturbance factor of conifer forests during the Late Holocene. The earliest remains of spruce budworm and hemlock looper were found ca. 3220 and 2350 cal yr BP, respectively. Peaks of insect head capsules occurred from ca. 1640 to ca. 625 cal yr BP. Low balsam fir pollen concentrations during this period suggest a lengthy episode ( 1000 yr) of high insect activity, resulting in extensive fir dieback and mortality. The long-term dynamics of the pristine balsam fir/spruce forests were mainly governed by the activity of insect defoliators. The limited extent and possibly the low occurrence of forest fires in the maritime environment of Anticosti Island allowed the development of mature coniferous stands propitious for insect infestations. Insect head capsules appeared to be a useful and effective tool for establishing insect presence and activity during the Holocene.     

Holocene insect fossils from two sites at Ennadai Lake, Keewatin, Northwest Territories, Canada

Ennadai Lake, in the forest-tundra ecotonal region of Keewatin, Northwest Territories, Canada, has been the subject of several paleoecological investigations (palynology, plant macrofossils, fossil soils). This study concerns Holocene insect fossils at Ennadai, a new approach in a region shown to be sensitive to climatic change. The Ennadai I site yielded 53 taxa, representing 13 families of Coleoptera and 7 families of other insects and arachnids, including abundant ants. These fossils range in age from about 6300 to 630 yr B.P. The Ennadai II site produced fossils of 58 taxa, including 13 beetle families and 15 families of other arthropods, ranging in age from 4700 to 870 yr B.P. The insect evidence suggests the presence of trees in the Ennadai region from 6000 to 2200 yr B.P. A conifer pollen decline from 4800 to 4500 yr B.P. at Ennadai has previously been interpreted as an opening up or retreat of forest in response to climatic cooling, but the insect fossils reveal the continued presence of trees during this interval. Both insect assemblages suggest trends of forest retreat and tundra expansion between about 2200 and 1500 yr B.P., presumably due to climatic cooling, with a return of woodland by about 1000 yr B.P.     

Postglacial diatom stratigraphy of Kirchner Marsh,Minnesota

Fossil diatom assemblages from a 12-m core from Kirchner Marsh were compared with modern surface assemblages from 159 Minnesota and labrador lakes using cluster analysis. The deepest levels of the core (spruce pollen zone 13,000 to 10,200 yr B.P.) resemble modern diatom assemblages from deep oligotrophic lakes of northeastern Minnesota. Diatom assemblages of the pine pollen zone (about 10,200 to 9500 yr B.P.) have few modern analogs. In the oak zone (9500 yr B.P. to present) after a brief pulse of diatom species indicative of eutrophication, the assemblages are dominated by species characteristic of shallow lakes, suggesting a drop in the lake water level during the prairie period (5500 to 7500 yr B.P.). Macrofossil data of W. A. Watts and T. C. Winter (1966, Geological Society of America Bulletin77, 1339–1360) show that this shift to shallow-water diatoms occurred when aquatic macrophytes appeared at the site in abundance.     

Plant and insect fossils at Norwood in south-central Minnesota: A record of late-glacial succession

The Norwood site in Sibley Co., Minnesota, contains 1.6 m of silt resting on till and overlain by peat. The base of the peat has been radiocarbon dated at 12,400 ± 60 and the top at 11,200 ± 250 yr B.P. The pollen, plant macrofossils, and insect remains in the basal silt consist of boreal species inhabiting open environments, but not tundra. No modern analogue exists for the insect assemblage, which includes elements of boreal forest, tundra-forest, and western affinities. The transition from an unstable open environment to a stable coniferous forest is reflected by both plant and insect fossils and is interpreted as a successional rather than a climatic event. During this time of significant biologic change, the climate is inferred to have been relatively uniform, with temperatures similar to those presently existing in the boreal forest south of the tundra-forest transition zone. The geologic and ecologic succession at Norwood is generally similar to that presently associated with ice stagnation of the Klutlan Glacier in the Yukon Territory. Localized successional sequences similar to those at Norwood are conceived to have occurred repeatedly during the melting of the Laurentide ice, and thus the proposed model has potentially broad application to the interpretation of late-glacial sequences.     

长白山自然保护区历史森林景观的初步重建

本文在对环境因子(包括年均温、年降水量、海拔高度、坡度和坡向)进行空间表达的基础上,建立了长白山自然保护区当前森林景观垂直带(包括阔叶红松林、云冷杉林、岳桦林和苔原)与这些环境因子间的逻辑斯蒂回归模型,然后利用1997年各环境因子的空间数字面来反推1975年森林景观带的空间分布,并根据1975年MSS遥感影像计算机监督分类的结果,用Kappa指数对模型预测结果进行验证。结果表明,苔原、岳桦林、云冷杉林和阔叶红松林1975年的预测分布面积分别为7243.83hm2、6517.08hm2、125570.16hm2和34264.80hm2,与遥感分类结果相比,苔原和云冷杉林的面积分别减少了29.04%和3.05%;岳桦林和阔叶红松林的面积分别增加了246.45%和8.6%。用逻辑斯蒂回归模型可以较好地预测苔原在保护区内的分布范围,同时在一定程度上也能预测阔叶红松林的分布范围,但是对岳桦林和云冷杉林分布范围的预测结果较差。尽管如此,作为一种方法性的探讨,本文可以为森林景观空间分布和环境因子变化的关系以及森林景观对气候变化的反应研究提供一种新思路。     

The Holocene thermal maximum and late-Holocene cooling in the tundra of NE European Russia

To investigate the Holocene climate and treeline dynamics in the European Russian Arctic, we analysed sediment pollen, conifer stomata, and plant macrofossils from Lake Kharinei, a tundra lake near the treeline in the Pechora area. We present quantitative summer temperature reconstructions from Lake Kharinei and Lake Tumbulovaty, a previously studied lake in the same region, using a pollen–climate transfer function based on a new calibration set from northern European Russia. Our records suggest that the early-Holocene summer temperatures from 11,500 cal yr BP onwards were already slightly higher than at present, followed by a stable Holocene Thermal Maximum (HTM) at 8000–3500 cal yr BP when summer temperatures in the tundra were ca. 3°C above present-day values. A Picea forest surrounded Lake Kharinei during the HTM, reaching 150 km north of the present taiga limit. The HTM ended with a temperature drop at 3500–2500 cal yr BP associated with permafrost initiation in the region. Mixed spruce forest began to disappear around Lake Kharinei at ca. 3500 cal yr BP, with the last tree macrofossils recorded at ca. 2500 cal yr BP, suggesting that the present wide tundra zone in the Pechora region formed during the last ca. 3500 yr.     

A full-glacial biota from southeastern Iowa,USA

Wisconsinan full-glacial silts filling a swale exposed in Conklin Quarry, Johnson Co., Iowa, contain a large and diverse biota that includes pollen, bryophytes, vascular-plant macrofossils, small mammals, molluscs, and insects. Radiocarbon dates on wood from the top, middle and bottom of the swale fill respectively were 16710 ± 270, 17 170 ± 205, and 18090 ± 190 yr BP. The pollen diagram is dominated by Picea (spruce), Pinus (pine), and Cyperaceae (sedge), and it records low pollen accumulation rates. Plant macrofossils include a number of tundra species along with Picea and Larix (larch) needles and small pieces of wood. The insect fauna contains many species now confined to the forest-tundra transition zone of northwestern Yukon and Alaska. Small mammals include the tundra indicators Dicrostonyx (collared lemming), and probably Microtus miurus (singing vole) together with boreal forest taxa. The molluscs include extinct and relict species and show the widest range in present geographic distribution, but Rocky Mountain and especially northern elements predominate in the swale fill. All these lines of evidence lead to consistent palaeoclimato-logical interpretation and palaeoecological reconstruction. The dominant habitats represented by the biota and sedimentary environment collectively included open calcareous silty to sandy or gravelly upland sites, minerotrophic fens (wetlands), pond- or stream-side clayey to sandy shores, and shallow (possibly ephemeral), cold, clear-water ponds. Mean July temperatures were probably 11° to 13°C cooler than at present. The biota indicates that a Picea-Larix krummholz with extensive tundra openings was present in southeastern Iowa between 18090 and 16710 yr BP.     

Seismic risk assessment at the proposed site of Gemsa wind power station,southwestern coast of Gulf of Suez,Egypt

Gemsa has been chosen as the site for one of a new generation of power stations along the south-western margin of the Gulf of Suez. This site has been affected by a number of destructive earthquakes (M_w> 5), in addition to large number of earthquakes with magnitudes of less than 5. In this study seismic activities in the region were collected and re-evaluated, and the main earthquake prone zones were identified. It is indicated that this site is affected by the southern Gulf of Suez, northern Red Sea and Gulf of Aqaba source zones. The southern Gulf of Suez source zone is the nearest to the proposed site. The stochastic simulation method has been applied to estimate the Peak GroundAcceleration at the site of the proposed Gemsa power plant. It was noticed that the pseudo-spectral acceleration (PSA) reaches 175 cm/sec2 resulting from the southern Gulf of Suez seismic source. In addition, the response spectrum was conducted with a damping value of 5% of the critical damping, and the predominant period reached 0.1sec at the site. These results should be taken into consideration by civil engineers and decision-makers for designing earthquake resistant structures.     

Advance of Norway spruce (Picea abies) onto mafic Lommoltunturi fell in Finnish Lapland during the last 200 years

As a result of global changes, shifts of alpine tree lines towards higher elevations have been recorded, but the role of the spatial variability of the snowpack and zonal‐pattern soil‐nutrient regimes is poorly understood. Norway spruce (Picea abies (L.) Karst) is best suited to fertile soils, and hence we applied soil physical‐chemical and snow measurements and the age chronology of Norway spruce along an elevational gradient (380–557 m a.s.l.) to address a vertical soil zonality hypothesis on mafic Lommoltunturi fell in Finnish Lapland. With regard to increasing elevation, we found an increase in soil N_TOT, C_TOT and Al, but a decrease in soil Ca, Mg and Ca:Al ratio as well as in electrical conductivity (EC). In addition, the snowpack was significantly thicker in low‐elevation forest than in the tree line and open tundra. In the 1840s, spruce established on low‐elevation soils with a Ca:Al ratio of 2.2. Starting from the 1920s a significant shift of spruce occurred such that it took 60 years to expand the tree line by 55 m in elevation. The spruce tree line has advanced, and the age distribution indicates new colonization of spruce in closed forest up to tundra. The poor soil Ca:Al ratio of 0.02 on tundra apparently is a constraint for spruce. Spruce forest is young (<165 years), and hence we argue that spruce has expanded onto formerly tree‐free sites of this mafic fell. This paper demonstrates that vertical soil zonality is a potential driver for the diffuse tree line of Picea abies on mafic Fennoscandian fells.     

Postglacial vegetation community change over an elevational gradient on the western Kenai Peninsula,Alaska: pollen records from Sunken Island and Choquette Lakes

The Kenai Peninsula of south‐central Alaska is a region of high topographic diversity with a complex glacial history. The sedimentary record of two small lakes [Sunken Island (SIL; 76 m a.s.l.) in the Kenai Lowlands; Choquette (CL; 527 m a.s.l.) in the Caribou Hills upland] exemplifies the postglacial development of the conifer–hardwood forest over an elevational range there. A herb–shrub tundra was established at both sites after deglaciation. By ~10.7 ka, poplar (Populus sp.) and alder (Alnus) dominated the lowland forest, while alder with minor poplar occurred at the upland site. Lake levels lower than today occurred during the early Holocene until ~8 ka. Subsequently at SIL, the near‐modern Kenai birch (Betula kenaica) – white spruce (Picea glauca) forest maintained prominence throughout the Holocene. However, at CL, alder dominated with dwarf birch and other subshrubs; small amounts of white spruce arrived ~5.2 ka. Black spruce (Picea mariana) grew around SIL by ~4 ka, but never gained prominence at CL. Fire, a prominent agent of disturbance in the Kenai Lowlands since ~8 ka, was essentially absent at the hardwood‐dominated upland site before ~6 ka, and rare thereafter. This suggests an important link between fire and spruce in Kenai forests.     

Isla San Pedro Nolasco as a Late Miocene intrusive record at the eastern margin of the Gulf of California: Insights from geological,geochemical and geochronological studies

Isla San Pedro Nolasco (ISPN) is a structural high bounded by inactive dextral oblique-slip faults in the east-central part of the Gulf of California rift zone and is composed of intrusive rocks not exposed on other Gulf of California islands. Here we present the reconnaissance results from geological mapping, as well as first geochemical and geochronological data for the ISPN intrusive complex. The intrusive rocks compose a sheet-like body of intermediate and felsic composition intruded by an intermediate and acidic dike swarm. All intrusive rocks (host and dikes) range in age from ca. 9 Ma to 10 Ma (⁴⁰Ar/³⁹Ar) and show a hydrous ferromagnesian mineral association (amphibole and biotite) with a calc-alkalic and transitional affinity. This hydrated mineralogical association has not been recognized in the coeval rocks along the onshore western margin of the North American plate (coastal Sonora). However, such hydrous mineralogical association is found in the coeval rift transitional volcanic rocks from the Baja California Microplate at Santa Rosalía and Bahía de Los Ángeles – Bahía de Las Ánimas. The ISPN continental block, at least 40 km long, has been pulled apart by transtensional faulting of the late Miocene Gulf of California shear zone before the westward migration of the North America-Pacific plate boundary at ca. 3–2 Ma. Eventually, ISPN became isolated as an island during the late Miocene flooding of the Gulf of California seaway.     

Quaternary vegetation history of the Mississippi embayment

Nonconnah Creek, located in the loess-mantled Blufflands along the eastern wall of the Lower Mississippi Alluvial Valley in Tennessee displays a sedimentary sequence representing the Altonian Substage through the Woodfordian Substage of the Wisconsinan Stage. The site has a biostratigraphic record for the Altonian and Farmdalian Substages that documents warm-temperate upland oak-pine forest, prairie, and bottomland forest. At 23,000 yr B.P., white spruce and larch migrated into the Nonconnah Creek watershed and along braided-stream surfaces in the Mississippi Valley as far as southeastern Louisiana. The pollen and plant-macrofossil record from Nonconnah Creek provides the first documentation of a full-glacial locality in eastern North America for beech, yellow poplar, oak, history, black walnut, and other mesic deciduous forest taxa. During the full and late glacial, the Mississippi Valley was a barrier to the migration of pine species, while the adjacent Blufflands provided a refuge for mesic deciduous forest taxa. Regional climatic amelioration, beginning about 16,500 yr B.P., is reflected by increases in pollen percentages of cooltemperate deciduous trees at Nonconnah Creek. The demise of spruce and jack pine occurred 12,500 yr B.P. between 34° and 37° N in eastern North America in response to postglacial warming.     

Holocene vegetation history of the Prince William Sound region, south-central Alaska

Vegetation history during the Holocene is interpreted from the pollen and sedimentary records of nine sections of peat deposits located in sedge tundra at sites in the northern and northwestern parts of the Prince William Sound region. Basal radiocarbon ages of the deposits are between 10,015 and 580 yr B.P. Modern surface pollen data from these and 25 additional sites, ranging from lowlands to an altitude of 675 m in the alpine tundra, were used to aid in the interpretation of the fossil records. Both frequency and influx pollen diagrams of the oldest section disclose a sequence of communities beginning with sedge tundra, containing thickets of willow and alder, followed by alder, which became predominant at about 8300 yr B.P. Later, alder declined, and an inferred growth of sedge tundra and the establishment of colonies of mountain hemlock and Sitka spruce with some western hemlock occurred about 2680 yr B.P. Finally, regrowth of sedge tundra accompanied by the development of forest communites took place over the past 2000 yr. The influence of glacier advances on the vegetation in the fjords occurred during Neoglacial episodes dated at 3200–2500 yr B.P. and during recent centuries. Regional Holocene tectonic activity was also an influential factor, especially at the time of the 1964 earthquake.     

Correlation of gravity with the Maquereau Group, southern Gaspe Peninsula, Quebec

Gravity observations at 150 locations in the Port-Daniel and Chandler area, southeastern Gaspé, Quebec, delineate an E-W to ENE-WSW trending elliptical positive Bouguer gravity anomaly with an amplitude of 19 mGal and a residual gravity anomaly of circa 22 mGal. This anomaly correlates closely with the Lower Cambrian and/or Hadrynian Maquereau Group, the steep gradients defining it coinciding with the contact between the group and the surrounding Ordovician and Siluro-Devonian sedimentary rocks. On the basis of its similar geological setting, this gravity anomaly may be compared with the positive Bouguer anomalies observed over the Fleur de, Lys zone to the east of the Baie Verte lineament in Newfoundland. Tridimensional modelling of the gravity data is inconsistent with a serpentinite dome as originally proposed and it suggests that the Maquereau Group represents an upthrusted slice of deep-seated volcanosedimentary sequences. These were originally located on the lower segment of the continental slope; they constituted a topographic high during the final process of the mountain building at the Taconic phase (Middle to Upper Ordovician time). The most probable way of emplacement of the Maquereau block is through a tear-fault running E-W to ESE-WNW in the southern Gulf of St. Lawrence west of Cape Breton and resulting from Early Paleozoic continental collision between a Eurafrican continent and a North American continent which had a salient located beneath the site of the modern Gulf. The Maquereau may then have been reactivated with a left-lateral motion accompanied of a rotation about a sub-horizontal axis. The presumed Baie Verte-Brompton line ought to be displaced to the north and the Humber zone is narrowed.     

The contemporary distribution of pollen in eastern North America: A comparison with the vegetation

By mapping and summarizing 478 pollen counts from surface samples at 406 locations in eastern North America, this study documents the relationships between the distributions of pollen and vegetation on a continental scale. The most common pollen types in this region are pine, birch, oak, and spruce. Maps showing isopercentage contours or isopolls for 13 important pollen types reflect the general N-S zonation of the vegetation. The maps and tabulations of average pollen spectra for the six major vegetational regions indicate high values for the following pollen types in each region: (1) tundra-nonarboreal birch, sedge, and alder; (2) forest/tundra-spruce, nonarboreal birch and alder; (3) boreal forest-spruce, jack pine (type), and arboreal birch with fir in the southeastern part; (4) conifer/hardwood forest-white pine, arboreal birch, and hemlock with beech, maple, and oak in the southern part; (5) deciduous forest-oak, pine, hickory, and elm, with beech and maple in the northern part, and highest values of oak and hickory west of the Appalachian crest; and (6) southeastern forest-pine, oak, hickory, tupelo, and Myricaceae. In some cases, less abundant pollen types are diagnostic for the region, e.g., bald cypress in the southeast. In the conifer-hardwood region and southward, pollen of weeds associated with deforestation and agriculture is abundant. The maps also show that much of southeastern U.S. and the area just to the east of Hudson Bay are in need of additional sampling. At 51 of the sites, absolute pollen frequencies (APF; grains/ml lake sediment) were obtained. These confirm the major conclusions from the percentage data, but differences are evident, e.g., the percentages of alder pollen peak in the tundra whereas alder APFs peak in the boreal forest, and spruce percentages peak in the forest-tundra whereas spruce APFs peak in the boreal forest. Because the APF data reflect the patterns of absolute abundance of individual taxa in the vegetation as well as the overall forest densities, future counts of modern pollen should include APF determinations. The effects of sedimentation processes on APF quantities indicate that APF samples should be obtained from moderate size lakes of similar morphology and hydrology and that, in each lake, several samples from the profundal zone should be pooled to create a sample representative of that lake.     

青藏高原苔原自然带谱及复式三维地带模式

首次依空间结构将青藏高原划分为外围高山、外缘高山、高原面高山、高原面及高原盆地等五种地貌体;厘清了高原不存在纬度地带性的误判,并结合作者前期高原苔原的论证,创建了高原苔原带谱体系及其三维仿真地带模型,分别为顶锥型(外围离散高山及高原面高山)、带型(外缘带状高山)和平面型(高原面).既具有一般高山苔原(alpine tundra)垂直带谱,又呈现高原山原苔原所特有的"复式三维地带性模式",在全球具有唯一性.首次编制了表征北极苔原与高原苔原关系的北半球苔原带模式图,指出长白山为东北亚高山苔原南界的论点不能成立,青藏高原苔原才是中国及东北亚苔原的南界.目前高原是第三极称谓皆无顶极生物带苔原带景观为前提,仅是高度上的寓意,从地理概念上明确了其与南北极并列的世界第三极定位.     

Oxygen isotope and deuterium composition of snow cover on the profile of Western Siberia from Tomsk to the Gulf of Ob

The purpose of this work is to study the variability of the isotope composition (δ¹⁸О, δD, d _exc) of the snow cover on a long transect of Western Siberia from the southern taiga to the tundra. The study of the snow cover is of paleogeographic, paleogeocryological, and paleohydrological value. The snow cover of western Siberia was sampled on a broadly NS transzonal profile from the environs of Tomsk (southern taiga zone) to the eastern coast of the Gulf of Ob (tundra zone) from February 19 to March 4, 2014. Snow samples were collected at 31 sites. Most of the samples represented by fresh snow, i.e., snow that had fallen a day before the moment of sampling were collected in two areas. In the area of Yamburg, the snow specimens collected from the surface are most probably settled snow of different ages. The values of δ¹⁸О in the snow from Tomsk to Yamburg varied from–21.89 to–32.82‰, and the values of δD, from–163.3 to–261.2‰. The value of deuterium excess was in the range of 4.06–19.53‰.     

Early postglacial environmental conditions in the northeastern Laptev Sea region

A comparison of the first results of a comprehensive micropaleontological analysis (pollen, spores, organic-walled microfossils, diatoms, ostracods) and radiocarbon ages (AMS¹⁴C) from sediment core recovered in the northeastern outer shelf of the Laptev Sea (51 m water depth) revealed a temporal coincidence between terrestrial and marine environmental changes that occurred between 11.2–10.3 cal ka. This interval provided evidence for a landscape transition from grass tundra to shrub tundra and the development of a freshwater estuarine basin with the strong influence of riverine discharge and a minor advection of North Atlantic waters. The establishment of a warmer and wetter climate promoted the expansion of shrub tundra habitats. The interval of 9.5–7.5 cal ka recorded a transition from a shrub tundra environment to forest-tundra vegetation. This interval also revealed a series of short-term temperature fluctuations, when summer temperatures were 3–4°C higher than today. The active advection of North Atlantic waters and the increase in salinity were also recorded by this interval.