New U/Th ages for Pleistocene megafauna deposits of southeastern Queensland,Australia

Arguments over the extinction of Pleistocene megafauna have become particularly polarised in recent years. Causes for the extinctions are widely debated with climate change, human hunting and/or habitat modification, or a combination of those factors, being the dominant hypotheses. However, a lack of a spatially constrained chronology for many megafauna renders most hypotheses difficult to test. Here, we present several new U/Th dates for a series of previously undated, megafauna-bearing localities from southeastern Queensland, Australia. The sites were previously used to argue for or against various megafauna extinction hypotheses, and are the type localities for two now-extinct Pleistocene marsupials (including the giant koala, Phascolarctos stirtoni). The new dating allows the deposits to be placed in a spatially- and temporally constrained context relevant to the understanding of Australian megafaunal extinctions. The results indicate that The Joint (Texas Caves) megafaunal assemblage is middle Pleistocene or older (>292 ky); the Cement Mills (Gore) megafaunal assemblage is late Pleistocene or older (>53 ky); and the Russenden Cave Bone Chamber (Texas Caves) megafaunal assemblage is late Pleistocene (～55 ky). Importantly, the new results broadly show that the sites date prior to the hypothesised megafaunal extinction ‘window’ (i.e., ～30–50 ky), and therefore, cannot be used to argue exclusively for or against human/climate change extinction models, without first exploring their palaeoecological significance on wider temporal and spatial scales.     

The Olduvai buffalo Pelorovis and the origin of Bos

The origin of the genus Bos is a debated issue. From ∼ 0.5 Ma until historic times, the genus is well known in the Eurasian large mammal assemblages, where it is represented by Bos primigenius. This species has a highly derived cranial anatomy that shows important morphological differences from other Plio-Pleistocene Eurasian genera of the tribe Bovini such as Leptobos, Bison, Proamphibos-Hemibos, and Bubalus. The oldest clear evidence of Bos is the skull fragment ASB-198-1 from the middle Pleistocene (∼ 0.6-0.8 Ma) site of Asbole (Lower Awash Valley, Ethiopia). The first appearance of Bos in Europe is at the site of Venosa-Notarchirico, Italy (∼ 0.5-0.6 Ma). Although the origin of Bos has traditionally been connected with Leptobos and Bison, after a detailed anatomical and morphometric study we propose here a different origin, connecting the middle Pleistocene Eurasian forms of B. primigenius with the African Late Pliocene and early Pleistocene large size member of the tribe Bovini Pelorovis sensu stricto. The dispersal of the Bos lineage in Western Europe during middle Pleistocene times seems to coincide with the arrival of the Acheulean tool technology in this continent.     

Southern dispersal and Palaeoecological implications of woolly rhinoceros (Coelodonta antiquitatis): review of the Iberian occurrences

Cold-adapted large mammal populations spread southward during the coldest and driest phases of the Late Pleistocene reaching the Iberian Peninsula. Presence of woolly rhinoceros (Coelodonta antiquitatis) can be identified from 23 Iberian sites, which is compiled and analyzed herein, and the fossil specimens from seven of these sites are described here for first time.Morphological and biometrical analyses demonstrate that the Iberian woolly rhinoceros did not significantly differ from individuals of other European populations, but represent the westernmost part of a continuous Eurasian belt of distribution.The first presence of woolly rhino in the Iberian Peninsula has been identified during the late Middle Pleistocene and early Late Pleistocene. However, the highest abundance of this species is recorded during MIS 3 and 2. The latest Iberian occurrences can be dated around 20 ka BP. The presence of woolly rhinoceros in the Iberian Peninsula correlates with periods of extreme dry and cold climatic conditions documented in Iberian terrestrial and marine sediment sequences.From a palaeobiogeographic point of view, the maximum southern spread of C. antiquitatis on the Iberian Peninsula was registered during the late Middle Pleistocene or early Late Pleistocene, reaching the latitude of Madrid (about 40°N). Subsequently, during MIS 3 and 2, all Iberian finds were restricted to the Northern regions of Iberia (Cantabrian area and Catalonia). The southern expansion of C. antiquitatis during the Late Pleistocene in the Iberian Peninsula reached similar latitudes to other Eurasian regions.The ecological composition of fossil assemblages with presence of woolly rhinoceros was statistically analyzed. Results show that temperate ungulate species are predominant at Iberian assemblages, resulting in a particular mixture of temperate and cold elements different of the typical Eurasian cold-adapted faunal associations. This particular situation suggests two possible explanations: a) Eventual migrations during the coldest time spans, resulting in a mixing of cold and temperate faunas, instead a faunal replacing; b) Persistence of woolly rhinoceros populations in the Iberian Peninsula during interglacial episodes confined at cryptic southern refugia.     

Quaternary coral reefs of the Red Sea coast, Egypt: diagenetic sequence, isotopes and trace metals contamination

This study focuses on the diagenetic sequence under marine and meteoric conditions as well as isotopes and trace metals contamination in Quseir and Gebel Zeit areas along the Egyptian Red Sea coast through a series of modern and fossil corals, Porites lutea and Favites pentagona. The diagenetic sequence begins with deposition of thin fringes of syntaxial aragonite and micritic high-magnesian calcite in the modern corals to completely altered Porites and partially altered Favites to low-magnesium calcite in the oldest Pleistocene unit. Average δ¹⁸O and δ¹³C values of Pleistocene corals in the two studied areas were lower than those of modern corals. Values of modern corals and lower fossil unit indicated coralline limestone, while those of middle and upper fossil units indicated fresh water influences. Average values of trace metals in modern corals were higher than those of Pleistocene counterpart except for Mn. Modern coral samples recorded enrichment in the average values of Pb, Zn, and Mn at Quseir area and enrichment in Co, Cu, and Ni at Gebel Zeit area. This may be attributed mostly to different tourist activities, landfill due to increase urbanization and nearby of Quseir area from the old phosphate harbor at El Hamrawin area, as well as oil exploration and production activities in the Gulf of Suez area. Also, results indicated that most samples of Porites have high concentration of trace metals than in Favites, especially in Cu, Zn, Mn, and Pb. This may due to high amounts of intergranular porosity and high total surface area of Porites in contrast to Favites.     

松嫩平原东部中、晚更新世地层界限：光释光年龄及元素变化

松辽平原东部受第四纪差异性构造活动影响,形成特殊的更新世沉积地层。由于第四纪划分方案和地域差异,松辽平原东部中、晚更新世界限仍处于不确定状态。通过黑龙江省方正剖面的研究、中更新统顶部(149.6±10.4) ka光释光年龄和主要氧化物、微量元素、稀土元素分析以及区域剖面对比,认为将松辽盆地东缘中、晚更新世的界限年龄置于140 ka更适合地层实际情况。中更新世地层以河湖相、河流相和洪积相沉积物为主,完整的晚更新世地层应该包括下部的沼泽漫滩相黏土、亚黏土和上部亚砂土、黄土沉积。黑龙江省荒山剖面哈尔滨组下部可能由于相变而缺少沼泽漫滩相的沉积物。方正剖面与榆树东岗组剖面地层可以对比。中、晚更新世沉积环境发生了明显变化,主要氧化物、微量元素和稀土元素含量在中、晚更新世的地层界限处都发生了明显的突变。     

Middle Pleistocene bivalves of the İznik lake basin (Eastern Marmara, NW Turkey) and a new paleobiogeographical approach

?znik Lake is a tectonically originated basin mainly controlled by the E–W trending middle strand of the North Anatolian Fault (NAF) system. Pleistocene sediments occurring in front of the faults are well exposed both in the northern and in the southern shorelines of the basin. In this study, two endemic brackish water bivalve species, Didacna subpyramidata Pravoslavkev 1939 and Didacna nov. sp. were found in the oldest terrace of the northern Pleistocene sequence. Having characterized morphology, these species serve as stratigraphic indicators in the regional Pleistocene stratigraphy of the Ponto-Caspian region, and thus are well correlated to the assemblages of the early Khazarian subhorizon (Middle Pleistocene). Hence, these data demonstrate that the early Khazarian brackish water sea covered the study area. Additionally, a model for the formation of the basin is proposed: the ?znik lake basin was a gulf of the former Marmara Sea in the early Khazarian, connecting the Marmara to the Black Sea and the Caspian Sea. The subsequent regional prograding uplifts, main dextral strike-slip fault and many normal faults of the NAF Zone cut off the marine connections to the basin, leading to its present location and topographic level.     

Proneofiber, a new genus of vole (Cricetidae: Rodentia) from the Pleistocene Seymour Formation of Texas,and its evolutionary and stratigraphic significance

A new genus and species of a microtine rodent, Proneofiber guildayi, is named on the basis of rooted teeth with cement from the warm Gilliland local fauna, Seymour Formation of Pleistocene age, Knox County, Texas. Proneofiber guildayi is considered to be ancestral to Neofiber, the round-tailed water rat, now living in the extreme southeastern United States. The presence of Pleistocene faunas prior to the first continental glaciation and the existence of three Pearlette-like volcanic ashes in the Plains Region suggest a revision of the age assignment of Pleistocene deposits from the nonglaciated region.     

L'escarpement de faille de la Têt est-il le résultat de la tectonique active Plio-Pléistocène ou d'une exhumation Pléistocène ?

The Têt Fault looks like an active fault. Its activity, attested for Pliocene, remains discussed for Pleistocene. We propose a new scenario of evolution for the Têt Fault, highlighting the importance of Pleistocene exhumation processes. It is based on a relationship between morphology and basin filling. During Pleistocene, the fault activity is weak and vertical displacements seem to have no considerable impact on the drainage features. This study concludes that Pliocene offset may also be reduced: the fault activity generating relief is anterior to Pliocene. To cite this article: J.-M. Carozza, S. Baize, C. R. Geoscience 336 (2004).     

First record of Procyon cancrivorus (G. Cuvier, 1798) (Carnivora,Procyonidae) in stratigraphic context in the Late Pleistocene of Brazil

Although five genera of procyonids are currently present in South America, only two of the extant genera, Procyon and Nasua are represented in the South American fossil record. A recent discovery of a procyonid lower second molar in Late Pleistocene deposits of Aurora do Tocantins, northern Brazil, offers potential to further our understanding of the stratigraphic and temporal range of South American fossil procyonids. We use geometric morphometric analysis of two-dimensional landmarks and semilandmarks to explore morphological variation in the lower second molars of extant Procyon lotor and Procyon cancrivorus and multivariate methods to support the identification of the Pleistocene specimen as P. cancrivorus. This material represents the second fossil record of P. cancrivorus in South America Procyonids entered South America in two phases: the first comprising by Cyonasua and Chapadmalania during the Late Miocene, and the other recent genera, beginning in the Late Pleistocene. These Late Miocene procyonids were more carnivorous than Late Pleistocene-Recent omnivorous taxa and possible went extinct due to competition with other placental carnivorans that entered South America and diversified during the latest Pliocene-Early Pleistocene.     

中国大陆第四纪地壳运动时程

本文从中国大陆有代表性的青藏高原、黄土高原、三峡地区着手,进行了新构造运动剖析。并结合全国火山活动期次,总结出中国大陆第四纪地壳运动的时程变化。从而为进一步探讨新构造运动特点和规律,为国民经济建设及人类生存环境提供了一定的地质背景资料。     

Magnetostratigraphy of the Orce region (Baza Basin), SE Spain: New chronologies for Early Pleistocene faunas and hominid occupation sites

The Orce fossil quarries, in the Baza Basin of southeastern Spain, are a rich source of Early Pleistocene Palaeolithic tools and vertebrate remains. Geologic fieldwork during the last decade has placed these fossiliferous strata within the context of a thick Neogene continental sequence. Detailed lithostratigraphic and magnetostratigraphic results indicate that at least the upper 60 m of this sequence are of Early Pleistocene age. The quarried strata (Venta Micena, Barranco León and Fuentenueva-3) are from a narrow time span (<100 ka) starting before 1.3 Ma. A new, lower excavation at ∼1.5 Ma (Fuentenueva-1 quarry) has a distinctly older fauna. These Orce strata provide a high resolution, Early Pleistocene record of grassland fauna that shows the end of Mammal Neogene fauna (MN17) in the Fuentenueva-1 site (with Gazella borbonica, Equus stenonis) and the beginning of more characteristic Pleistocene fauna in the Venta Micena site (with Hippopotamus antiquus, Equus granatensis, Homo sp.). Thus far, no evidence for human occupation has been found within the earlier Fuentenueva-1 quarry, although many of the same terminal MN17 species have been found with hominids on the other side of Europe at the Dmanisi site (earliest Pleistocene) in the Republic of Georgia.     

A new record of Pleistocene hippopotamus from River Severn terrace deposits, Gloucester, UK—palaeoenvironmental setting and stratigraphical significance

A new Pleistocene vertebrate assemblage from fluvial deposits of the River Severn in Gloucester, England, has yielded the remains of hippopotamus (Hippopotamus amphibius), a new record for this terrace system, with additional material from probable bison (cf. Bison priscus) and elephant (Elephantidae sp.). The presence of these taxa indicates fully temperate climatic conditions and the occurrence of hippopotamus, a significant biostratigraphical indicator for the British Late Pleistocene, suggests an age for the assemblage within MIS 5e (the Last Interglacial). This would contradict the older MIS 7-6 age for the gravel body that is currently accepted on the basis of deposit mapping and imply a more complex mode of deposition than presently envisaged in the valley.     

First megafossil evidence of Cyatheaceous tree fern from the Indian Cenozoic

A part of the compressed tree fern axis with leaf and adventitious root scars in unusual arrangement from Plio–Pleistocene sediments of Arunachal Pradesh, India is described as Cyathea siwalika sp. nov. This record suggests that Cyathea was an important component of tropical evergreen forest in the area during the Plio–Pleistocene and this group subsequently declined in importance within the local vegetation.     

The northernmost record of Catagonus stenocephalus (Lund in Reinhardt, 1880) (Mammalia,Cetartiodactyla) and its palaeoenvironmental and palaeobiogeographical significance

During fieldwork carried out in January 2009 at Aurora do Tocantins (Tocantins State, northern Brazil), we recovered a fragmentary right maxilla (UNIRIO-PM 1006) of Catagonus stenocephalus from a sedimentary deposit of presumed late Pleistocene age in a karstic cave. This paper aims to: (1) provide the first record of C. stenocephalus in the northern region of Brazil (and consequently, also the northernmost one); (2) update the geographic distribution of C. stenocephalus; (3) present a date for the specimen; and (4) discuss the palaeoenvironmental and palaeobiogeographical implications of the finding. The species C. stenocephalus (Lund) is known from the Bonaerian (middle Pleistocene) and Lujanian (late Pleistocene to earliest Holocene) ages in Argentina, Uruguay, Brazil and Bolivia. The new record presented here extends the geographical distribution of C. stenocephalus more than 1000 km north from the former northernmost record (caves of Lagoa Santa region). Peccaries of the genus Catagonus have several morphological features associated with cursorial habits in relatively open and dry environments. The new distributional range of C. stenocephalus is coincident with the Chacoan subregion, characterized by dry climates and open areas. As the studied material comes from the top of the carbonate layer, this may suggest that the deposition of the C. stenocephalus remains described here is synchronous with the onset of a wetter climate phase. This argument is also in accordance with the datation results, around 20 ky BP, just after the last glacial maximum. This increasingly wet climate, which may also be related to the climatic changes that occurred during the late Pleistocene/early Holocene, could be a factor in the extinction of C. stenocephalus in South America.     

Late Pleistocene vegetation of Kings Canyon,Sierra Nevada,California

Seven packrat midden samples make possible a comparison between the modern and late Pleistocene vegetation in Kings Canyon on the western side of the southern Sierra Nevada. One modern sample contains macrofossils and pollen derived from the present-day oak-chaparral vegetation. Macrofossils from the six late Pleistocene samples record a mixed coniferous forest dominated by the xerophytic conifers Juniperus occidentalis, Pinus cf. ponderosa, and P. monophylla. The pollen spectra of these Pleistocene middens are dominated by Pinus sp., Taxodiaceae-Cupressaceae-Taxaceae (TCT), and Artemisia sp. Mesophytic conifers are represented by low macrofossil concentrations. Sequoiadendron giganteum is represented by a few pollen grains in the full glacial. Edaphic control and snow dispersal are the most likely causes of these mixed assemblages.The dominant macrofossils record a more xeric plant community than those that now occur on similar substrates at higher elevations or latitudes in the Sierra Nevada. These assemblages suggest that late Wisconsin climates were cold with mean annual precipitation not necessarily greater than modern values. This conclusion supports a model of low summer ablation allowing for the persistence of the glaciers at higher elevations during the late Wisconsin. The records in these middens also suggest that S. giganteum grew at lower elevations along the western side of the range and that P. monophylla was more widely distributed in cismontane California during the Pleistocene.     

Geological and ecological perspectives on the Middle Pleistocene

This paper evaluates the geological and paleoecological implications of a Wenner-Gren symposium Stratigraphy and Patterns of Cultural Change in the Middle Pleistocene. The deep-sea, glacial-eustatic, loess, alluvial, and palynological records suggest between six and eight cold-warm cycles since the Brunhes-Matuyama magnetic reversal of 700,000 BP. Till and outwash stratigraphies are inadequate to provide a valid nomenclature for the numerous glacials preceding the Würm. Since at least five of the glacials since 700,000 BP were sufficiently severe to produce permafrost in midlatitude Europe, the “glacial Pleistocene” begins with the Brunhes-Matuyama. Although earlier cold-warm cycles extend well back into the early Pleistocene, with extensive glaciation and repeated floral decimations in higher latitudes, the first record of permafrost in the Rhine Basin 700,000 BP argues that major climatic oscillations of the Brunhes epoch were of glacial-interglacial amplitude. It is therefore recommended that a Lower-Middle Pleistocene boundary be linked to the practicable and universally applicable chronometric horizon provided by the Brunhes-Matuyama reversal, while the Middle-Upper Pleistocene limit can continue to be drawn at the base of the last, Eemian Interglacial (130,000 BP). The tropical African record presently contributes little to general understanding of the Middle Pleistocene, while the climatic cycles of higher latitudes are of limited value in analyzing mid-Pleistocene records of the tropical continents. Problems of stratigraphic control and environmental contexts for archeological sites are discussed.     

Mauer – the type site of Homo heidelbergensis: palaeoenvironment and age

The mandible of Homo heidelbergensis was found 1907 in the sand pit Grafenrain at Mauer in coarse fluvial sands 24 m below the surface, deposited in a former course of the Neckar River. These ‘Mauer sands’ are overlain by a series of glacial-climate loess deposits with intercalated interglacial palaeosols, which can be correlated with Quaternary climate history, thus indicating an early Middle Pleistocene age for H. heidelbergensis. The ‘Mauer sands’ are famous for their rather rich mammal fauna, which clearly indicates interglacial climate conditions. The faunal evidence – in particular the micromammals – place the ‘Mauer sands’ into MIS 15 or MIS 13 although most stratigraphic arguments favour correlation to MIS 15 and therefore to an age of ca 600 ka.     

The first record of “spelaeoid” bears in Arctic Siberia

For a long time, “spelaeoid” (cave-bear-like) bears, Ursus (Spelearctos) spp., were believed to be almost purely European animals. Their geographic range has recently been extended to the east, in southern Siberia, Transbaikalia, Kirghizia, Mongolia and Korea. Two unexpected new findings, presented here in detail, significantly change existing views on the distribution of cave bears; both were found in North-Eastern Siberia, far beyond the Arctic Circle, more than 1500 km to the north-east of the previously accepted range.One of the fossils is a mandible, found near the town of Cherskiy at 68.73°N, 161.38°E. The analysis of local geology and accompanying mammal fossils suggests that it comes from the Olyorian Fauna (Early to early Middle Pleistocene). Morphologically, the Cherskiy mandible is closest to Ursus savini, a small middle Pleistocene cave bear from the British Cromer Forest-bed Formation, but differs in having a slightly more advanced dentition, and thus it is described as a new subspecies Ursus savini nordostensis. Another newly recognized fossil of the “spelaeoid” bear is an astragalus found at the Oskhordokh site at 67.54°N, 135.67°E, on a large gravel bar on the right bank of the Adycha River. This specimen is attributed to Ursus cf. deningeri.The paper also presents an interesting example of the interaction between classical and “molecular” palaeontology.The new finds significantly change existing ideas on the ecology and evolution of cave bears, some of the most remarkable members of the extinct Pleistocene megafauna.