New lacewings (Insecta: Neuroptera) from the Lower Cretaceous Wealden supergroup of Southern England

Eight new genera and thirteen new species of lacewings (Neuroptera) are described from the Lower Cretaceous Wealden Supergroup, Weald Sub-basin: Principiala rudgwickensis sp. nov. (Ithonidae), Stenomylina medialis gen. et sp. nov., Protosmylina bifasciata gen. et sp. nov. (Osmylidae), Cretapsychops corami gen. et sp. nov., Micropsychops parallelus gen. et sp. nov., Valdipsychops logunovi gen. et sp. nov., V. brigidae gen. et sp. nov., V. maculosus gen. et sp. nov., V. proudlovei gen. et sp. nov., V. minimus gen. et sp. nov., Psychopsites rolandi gen. et sp. nov. (Psychopsidae), Actinophlebioides valdensis gen. et sp. nov. and Protohemerobius perexiguus gen. et sp. nov. (Neuroptera familia incertae sedis). Psychopsidae are shown to dominate the neuropteran fauna, while the presence of Brongniartiellidae has not been confirmed in the Wealden, due to previously thought representatives of this family being in fact psychopsids. This is the first fossil record of Ithonidae and Osmylidae in Britain. The range of Kalligrammatidae has been extended in the Cretaceous of Britain from the mid-Valanginian to early Barremian.     

A new neoselachian shark from the marine Early Cretaceous of Southern England

Rare, isolated teeth of Corysodon multicristatus sp. nov. are described from two levels in the Atherfield Clay Formation (Early Aptian, Early Cretaceous) of Atherfield Point on the Isle of Wight, UK. Ten teeth of the new species were recovered from 1095 kg of washed and graded sediment residues. The teeth themselves are very small (around 0.5 mm high) and possess a distinctive crown bearing a tiered series of transverse crests adapted for rasping. Details of the dental architecture of the Atherfield Clay Formation specimens clearly indicate that the Cretaceous material differs significantly from the teeth of the type species for the genus, Corysodon cirinensis, recorded from the Kimmeridgian of northern France and Switzerland. C. multicristatus is the first substantiated record of the genus from the Early Cretaceous, thereby extending the stratigraphic range of the genus from the latest Jurassic, and the geographical range from continental Europe to the UK.     

准噶尔盆地南缘下白垩统层序界面的识别

准噶尔盆地南缘下白垩统内部的层序及界面单纯在地震剖面或电测曲线上都较难识别,但它们在岩石地球化学等特征上有较明显的反映.研究发现,层序边界处具有Ro值的突变,有机质含量(TOC)的低谷,ΔlogR曲线处于基线,粘土矿物组合突变,坡缕石、伊/蒙混层的含量明显增加;Cu、Li、Zn、Cr、V、Ba等微量元素的含量处于低谷;锆石等稳定重矿物含量以及黄铁矿含量较低,褐铁矿含量高等特征.结合地震剖面、测井曲线等特征,将准噶尔盆地南缘下白垩统划分出2个三级层序,每个层序又分别由低位体系域、湖侵体系域和高位体系域组成.     

The non-marine Lower Cretaceous Wealden strata of southern England

The non-marine Lower Cretaceous Wealden strata of the Wessex-Weald Basin (southern England) are introduced, with reference to the depositional model developed by Professor Percival Allen FRS (Allen, 1975). To demonstrate this model and the development of Wealden palaeoenvironments through time, Wealden sites have been selected for the Geological Conservation Review programme. Site selection rationale is briefly outlined.     

A New Eusauropod Dinosaur from the Lower Cretaceous of Guangxi Province, Southern China

<正>A new sauropod dinosaur,Liubangosaurus hei gen.et sp.nov.,is erected based on a specimen represented by five articulated middle-caudal dorsal vertebrae,which was discovered in the Lower Cretaceous Napai Formation of Guangxi Province,southern China.This new taxon is diagnosed by a unique combination of derived features:prezygapophysis closely contacts with parapophysis,with the prdl and prpl absent;presence of cavity on the dorsal surface of the diapophysis;neural spine very low,with its distal end level with that of diapophysis;distal end of the neural spine strongly expanded laterally to form a platform;marked fossa formed between the infradiapophyseal lamina and the parapophysis;broad,flat area of featureless bone on lateral surface of neural arch;vertically directed infradiapophyseal lamina expands or bifurcates ventrally to form a inverted "Y";highly positioned parapophyses large and tear-drop in shape.The discovery of this new taxon increases the diversity of sauropods in China during the Early Cretaceous.     

A new afrograptid (Diplostraca: Estheriellina) from the Lower Cretaceous of southern England

The Family Afrograptidae is a ‘conchostracan’ group with multiple radial costae reaching to the umbo on their carapaces. It comprises four described genera: Afrograpta, Camerunograpta, Congestheriella and Graptoestheriella with a total of thirteen described species which are occasionally reported from the Jurassic and the Cretaceous in Africa, Europe and South America (i.e. Afrograpta from the Upper Cretaceous of Cameroon; Camerunograpta from the Jurassic to Cretaceous of Cameroon; Congestheriella from the Jurassic to Upper Cretaceous of the Congo Basin, Brazil, Bulgaria, Venezuela and Argentina; and Graptoestheriella from the Upper Jurassic to Lower Cretaceous of Brazil). A new genus and a new species, Surreyestheria ockleyensis gen. et sp. nov., belonging to the Family Afrograptidae from the Lower Cretaceous (lower Barremian) Upper Weald Clay Formation of Ockley Village, Surrey County, southern England is described in this paper. The new genus mainly differs from the other four genera by the special reticulate ornamentation on its carapace. It indicates that the Family Afrograptidae was more diverse and more widely distributed in the late Mesozoic than previously supposed. Afrograptidae is a special branch of Estheriellina the latter originating in the late Palaeozoic and the former in the early Mesozoic. Afrograptids, as a whole had been widespread across Pangea in the Early Jurassic.     

A new family of scorpionflies (Insecta; Mecoptera) from the Lower Cretaceous of England

Seven specimens of fossil scorpionflies (Mecoptera) not assignable to any known family were discovered in the Wealden Supergroup (Lower Cretaceous) of southern England. They were found at Rudgwick Brickworks, West Sussex and Smokejacks Brickworks, Surrey and came from the Upper Weald Clay Formation, dated as Barremian (∼129.4–125 Ma). A new family – Englathaumatidae fam. nov., new genus – Englathauma gen. nov. and two new species E. crabbi sp. nov. and E. mellishae sp. nov. are described. A discussion of systematic position of these new taxa within the order Mecoptera is given. Englathaumatidae fam. nov. has been a nomen nudum since the year 2002, due to the first author's untimely death.     

Leptolepid otoliths from the Hauterivian (Lower Cretaceous) Lower Weald Clay (southern England)

Teleostean saccular otoliths from the upper part of the late Hauterivian Lower Weald Clay Formation of the Wealden Supergroup exposed at Langhurstwood Quarry, West Sussex, UK, and Clockhouse Brickworks, Surrey, UK are described for the first time. Two new species of the genus Leptolepis, Leptolepis wealdensis and Leptolepis toyei are described. Many of the specimens are densely packed on individual bedding planes and they are interpreted as coprocoenotic accumulations. Additional mechanisms of deposition and concentration are discussed, in particular wave action. Ontogenetic series show isometric growth of the otoliths, and some specimens show growth rings on two orders of magnitude.     

The Wealden (non-marine Lower Cretaceous) of the Wessex Sub-basin,southern England

The Wealden Beds (non-marine Lower Cretaceous) of the Wessex Sub-basin, southern England, are exposed principally in coastal sections on the Isle of Wight and in Dorset. Geological Conservation Review sites within these strata have been extensively documented since the earliest days of geological enquiry in Great Britain. The succession is dominated by the alluvial Wessex Formation which demonstrates a broad east–west transition from meanderplain lithofacies to coarser-grained alluvial sediments, in relative proximity to the Cornubian source massif. The meanderplain sediments on the Isle of Wight are of international importance for their plant and animal fossils, the latter including many dinosaurs and their trackways. Upper Barremian transgression resulted in the spread of muddy lakes and coastal lagoons from the Weald Sub-basin into the eastern part of the Wessex Sub-basin, around or through the Purbeck–Isle of Wight structure. The resulting richly fossiliferous mudrock-dominated strata are now represented by the Vectis Formation on the Isle of Wight and in Swanage Bay, Dorset. The Geological Conservation Review sites in the Wessex Sub-basin are documented and interpreted, with particular reference to research history, chronostratigraphy, structural context, palaeoenvironments, palaeobiology and palaeoclimatology. New directions for research are proposed, as applicable.     

The Wealden (non-marine Lower Cretaceous) of the Weald Sub-basin,southern England

The Wealden strata (non-marine Lower Cretaceous) of the Weald Sub-basin outcrop in the Weald district of south-east England; the Wealden type-area. The succession is made up of the mixed alluvial–lacustrine–lagoonal Hastings Beds Group below and the predominantly lacustrine–lagoonal Weald Clay Group above. Deposition was strongly influenced by tectonism amongst surrounding massifs, and the warm to hot, periodically wet Wealden climate. Geological Conservation Review sites within the Weald district are dominated by inland sites, but also include extensive coastal cliff and foreshore exposures near Hastings, East Sussex. The Wealden strata have been documented and interpreted since the earliest days of geological enquiry in Great Britain. Collectively, the selected sites demonstrate the key elements of a depositional model for the Wealden of the Weald, developed and published by Professor Percival Allen FRS (1917–2008) in these Proceedings (Allen, 1975). The sites are documented and interpreted, with special reference to research history, chronostratigraphy, structural context, palaeoenvironments, palaeobiology and palaeoclimatology. New directions for research are proposed, as applicable.     

塔南凹陷早白垩世构造演化及其对沉积充填和砂体分布的控制

塔南凹陷早白垩世经历了断陷初期冲积-滨浅湖盆层序、强烈断陷期半深湖-深湖湖盆层序及断坳期三角洲-湖泊盆型层序的沉积演化序列。受盆地整体格架的控制,凹陷内可划分出5个次级构造单元,各次级构造单元古构造格架决定了沉积相带展布、砂体的发育和分布。     

A re-evaluation of Ophiomorpha burrows in the Wealden Group (Lower Cretaceous) of southern England

Attribution of burrows in the Wealden Group of southern England to Ophiomorpha is rejected. The burrows are essentially cylindrical, unlined and with a meniscate fill. Any outer knobbly appearance is due to diagenetic poikilotopic cementation or to differential weathering of a mudchip-sand fill. The variable nature of meniscate fill reflects passage of the producer through the thin-bedded, alternating sand-mud sediments or along sand-mud interfaces. The burrows are assigned to Beaconites, though, since the identity of this ichnotaxon has been questioned, reference is also made to Taenidium. Two ichnoassociations are recognized: (1) a Beaconites antarcticus-Scoyenia (or Taenidium-Scoyenia) association (Weald Clay) of marginal lacustrine situation with fluvial input, and (2) a Beaconites barretti-Planolites (or Taenidium-Planolites ) association of the fluvial (lacustrine delta) of the Lee Ness Sandstone (Ashdown Formation). The Wealden burrows offer no inherent indications of palaeosalinity, and inferences made on supposed occurrences of Ophiomorpha in the Wealden Group must be reassessed. Other occurrences of Ophiomorpha in non-marine facies are questioned.     

A new titanosaur from the Lower Cretaceous of Brazil

Here we describe a new record of a sauropod dinosaur from the Lower Cretaceous (Hauterivian–Barremian) Rio Piranhas Formation, Sousa Basin, NE Brazil. Dinosaur fossil bones from this deposit were unknown until now. Thus, the discovery of a sauropod fibula from this locality is highly significant. Our discovery represents an indeterminate titanosaur and the earliest stratigraphic occurrence of this group in central Gondwana. When compared to chronocorrelate titanosaur trackmakers of this geological unit, this fossil specimen appears substantially smaller. Histological analysis of the fibula suggests that this is a relatively young individual (approximately 40–50% adult body size) that had passed its most rapid phase of early juvenile growth, but had not yet attained somatic maturity. Thus, the fibula recovered is from a young individual rather than from a small-bodied adult titanosaur.     

Lacewings (Insecta: Neuroptera) from the Lower Cretaceous Purbeck Limestone Group of southern England

Eight new genera and 12 new species are described from the Lower Cretaceous Purbeck Limestone Group, southern England. Sophogramma wimbledoni sp. nov. (Kalligrammatidae) is described from the Purbeck of Wiltshire. Pterinoblattina peverilensis sp. nov., Pterinoblattina fasciata sp. nov., Purbepsychopsis parallela gen. et sp. nov. (Psychopsidae), Ovalorobius edmondsi gen. et sp. nov. (Prohemerobiidae), Mesosmylidus vulgaris gen. et sp. nov., Osmylochrysa anomala gen. et sp. nov., Osmylochrysa fragilis gen. et sp. nov., Stenochrysa gradata gen. et sp. nov. (Osmylidae), Mesypochrysa minuta sp. nov. (Chrysopidae), Purbemerobius medialis gen. et sp. nov. (?Hemerobiidae), Epimesoberotha parva gen. et sp. nov. (Berothidae) and Pseudocorydasialis alleni (Neuroptera familia incertae sedis) are described from Durlston Bay, Dorset. The species Pterinoblattina penna Scudder, Pterinoblattina pluma (Giebel) (Psychopsidae) Sialium sipylus (Nymphidae) and Osmylopsis duplicata (Osmylidae) are re-examined, described and figured. The genus Valdipsychops Jepson et al. has been synonymized with Pterinoblattina.     

Possible persistence of the morganucodontans in the Lower Cretaceous Purbeck Limestone Group (Dorset, England)

‘Triconodonts’ are relatively rare in the new Purbeck Limestone Group mammal fauna, especially in comparison with the finds made in the nineteenth century in the same beds. However, besides some remains of the already known Triconodontidae, a new possible morganucodontan of uncertain familial affinities, Purbeckodon batei gen. et sp. nov., has been discovered, extending considerably the chronological range of the order while constituting another testimony to the vitality and diversity of Mesozoic mammals.     

The marginal Wealden (non-marine Lower Cretaceous) between Wiltshire and Buckinghamshire,England

Non-marine Lower Cretaceous beds of Wealden aspect have long been known from the northern margin of the Wessex–Weald Basin, between Wiltshire and the south Midlands. Termed the Whitchurch Sands Formation, these badly exposed and generally poorly fossiliferous strata appear to represent interdigitating alluvial and brackish-marine units separated by significant sedimentary breaks. Geological Conservation Review sites within the Whitchurch Sands are described and interpreted for their chronostratigraphic, palaeoenvironmental and palaeoclimatic significance.     

Saurischian track assemblages from the Lower Cretaceous Shenhuangshan Formation in the Yuanma Basin,Southern China

Cretaceous dinosaur tracks from Hunan Province are historically significant as the basis for three ichnotaxa: Xiangxipus chenxiensis, Xiangxipus youngi, and Hunanpus jiuquwanensis all representing theropodan tracks, described from a single site in 1982. Although the type locality has since been destroyed, the type specimen and replicas remained available for restudy in 2006, when paratype Hunanpus tracks and sauropod footprints were described from a second, nearby locality. Material from both localities is here re-described in detail. It is proposed that while Xiangxipus chenxiensis can be regarded as a distinct ichnospecies, probably representing an ornithomimid trackmaker, Xiangxipus youngi cannot be accommodated in the same ichnogenus. We consider it similar to the ichnogenus Wupus from the Lower Cretaceous of Sichuan Province, and therefore of possible avian affinity. We also find no justification for regarding Hunanpus jiuquwanensis as generically distinct from the widespread ichnogenus Grallator, and therefore rename it Grallator jiuquwanensis comb. nov. The Hunan track record is generally consistent with the sparse record of theropod skeletal remains in the province, but adds evidence of sauropods that was previously lacking.     

鸡西盆地下白垩统含煤层系低孔低渗砂岩的成因探讨

黑龙江鸡西盆地下白垩统城子河组和穆棱组是主要的含煤系地层,岩石类型为长石岩屑砂岩和岩屑长石砂岩,成岩作用主要为压实作用、胶结作用、溶蚀溶解作用及交代作用等,露头砂岩经历了表生成岩作用。压实作用和胶结作用是储层砂岩物性差的重要原因,二者相比,前者对该区孔隙度减少贡献更大。对比露头砂岩与岩芯砂岩,露头砂岩的储层物性更好。     

A tiny maniraptoran dinosaur in the Lower Cretaceous Hastings Group: Evidence from a new vertebrate-bearing locality in south-east England

In contrast to the Barremian Wessex Formation on the Isle of Wight, the remains of small theropods are rare in the Berriasian-Valanginian Hastings Group of the English mainland. Both units are part of the dinosaur-rich Wealden Supergroup (Berriasian-Aptian) of southern Britain. Here we report the cervical vertebra of a small dinosaur from the Pevensey Pit at Ashdown Brickworks, a site located northwest of Bexhill, East Sussex. The pit yields a rich assemblage of vertebrate fossils from the Valanginian Wadhurst Clay Formation of the Hastings Group. The new specimen, a near-complete but water-worn posterior cervical vertebra, is tiny (total centrum length = 7.1 mm) but evidently from an adult theropod. Its large hypapophysis, X-shaped neural arch and amphicoelous centrum suggest referral to Maniraptora, and the subparallel anterior and posterior articular surfaces imply that it does not belong to a deinonychosaur. The X-shaped neural arch recalls a similar condition seen in oviraptorosaurs while the high neural canal/articular surface ratio (0.70) is bird-like. The specimen is significant in representing the first maniraptoran to be reported from the Hastings Group but is otherwise indeterminate. By comparing the specimen to better known maniraptorans and estimating the proportions of the animal to which it belongs, we suggest that the total skeletal length of this maniraptoran was somewhere between 16 and 40 cm. It may therefore have been among the smallest of known Mesozoic dinosaurs.