Age and palynological characteristic of the Dongning Formation (Eastern Heilongjiang,China)

The taxonomic composition of the palynological assemblage of the Dongning Formation is supplemented. The palynological assemblage corresponds to those from the Lipovtsy Formation (Aptian) in the Razdolnaya Basin of Primorye (Russia) and the Muling Formation (Aptian) in the Jixi Basin of eastern Heilongjiang (China). It is found that the age of the Dongning Formation is Aptian. The palynological assemblage is characterized by dominance of spores of Gleicheniaceae; they are accompanied by spores of Cyatheaceae. The most important feature of the palynological assemblage of the Dongning Formation is the presence of angiosperm pollen (Tricolpites sp., T. micromunus, T. vulgaris, Retitricolpites georgiensis, R. vulgaris, Clavatipollenites hughesii, Quercites sparsus, Fraxiniopollenites variabilis).     

Contributions of zircon U–Pb geochronology to understanding the volcanic and sedimentary history of some Purāna basins,India

In this century, U–Pb ages of magmatic and detrital zircons, together with a few less accurate but fairly robust ages determined on monazite and baddeleyite, in the Purāna successions in India have established a few firm timelines that constrain the opening, closure, inversion, and provenance of the Purāna basins. The Cuddapah basin opened shortly before ca. 1900 Ma, the Vindhyan basin opened before ca. 1630 Ma, the Khariar basin likely opened ca. 1500 Ma, and the Chhattisgarh basin opened ca. 1400 Ma. The Marwar basin opened after ca. 750 Ma. The Chhattisgarh basin began to invert at ca. 1000 Ma and closed shortly thereafter. The Indravati and the Vindhyan basins closed ca. 1000 Ma. There are no other defensible geochronologic data to adequately constrain the opening and closure of other Purāna basins (e.g., Kaladgi, Badami, Bhima, Kurnool, Mallampalli, Albaka, Ampani, Sabari, and Kolhan). Neither the fossil record nor the biostratigraphy of these basins necessarily correspond to the chronology determined through radiometric measurements.The discovery of ca. 1000 Ma volcanic events in the Indravati and Chhattisgarh basins adds to the growing list of ca. 1000 Ma thermal disturbances in the Indian shield. Most of these events were likely the far field effects of the final assembly of Rodinia.     

Climate characteristics of the eastern Mongolian Plateau,China during the early Early Cretaceous (145–132 Ma): Palynological evidence from the Tongbomiao Formation in Well Hong-6, Hailar Basin

This study identified two palynological assemblages, namely Bayanhuasporites-Cycadopites-Protoconiferus and Cicatricosisporites-Cedripites-Perinopollenites, in the Tongbomiao Formation in the Hongqi Sag in the Hailar Basin, Inner Mongolia, China for the first time. The former is distributed in the lower part of the Tongbomiao Formation and is characterized by abundant gymnosperm pollen and diverse fern spores. Among them, the gymnosperm pollen is dominated by Paleoconifer (4.98%–31.62%) and Cycadopite (8.55%–25.23%) pollen grains and also includes other pollen grains such as Classopollis, Parcisporites, Erlianpollis, Callialasporites, and Jiaohepollis. The fern spores in the former palynological assemblage contain Bayanhuasporite (0–8.96%), Granulatisporites (0.93%–6.97%), and some important Cretaceous genera, such as Cicatricosisporites, Concavissimisporites, Densoisporites, Hsuisporites, Foraminisporis, and Leptolepidites. The Cicatricosisporites-Cedripites-Perinopollenites palynological assemblage is distributed in the upper part of the Tongbomiao Formation. Gymnosperm (77.30%), Pinaceae (31.9%), and Paleoconiferus (19.02%) pollen predominate this palynological assemblage, and Quadraeculina, Erlianpollis, and Jiaohepollis pollen are also common in this assemblage. The fern spores in this palynological assemblage include abundant Cicatricosisporites (4.29%). Besides, Concavissimisporites, Aequitriradites, and Leptolepidites are also common in this palynological assemblage. No angiosperm pollen has been found in both palynological assemblages. The identification of both palynological assemblages provides important evidence for the biostratigraphic correlation between the Hailar Basin and its adjacent areas. It also enables the reconstructions of the Berriasian-Valanginian (Early Cretaceous) vegetation and the paleoclimate on the eastern Mongolian Plateau during 141–132 Ma. The vegetation reconstructed on the palynological data of the represented by Hailar Basin in eastern Mongolian Plateau (141.6–141.4 Ma), form conifer forest or conifer broad-leaved mixed forest to conifer forest with shrubs and grassland, the climate belongs to warm temperate and warm-subtropicalt, the highest temperature is estimated to reach 35–38°C. Form 132.3 Ma, the vegetation type is conifer forest, and its paleoclimate is sub-humid warm temperate, the highest temperature is estimated to reach 24–29°C.©2022 China Geology Editorial Office.     

Conodont-based stratigraphy of the Mosolovian Regional Stage (Eifelian,Middle Devonian) of the Voronezh Anteclise

Conodonts from the deposits of the Mosolovian Regional Stage (Middle Devonian) of the Voronezh Anteclise are studied on the basis of the data from six boreholes. Three assemblages are recognized. The index species of the lowermost assemblage is Icriodus formosus Naz. The index species of the middle assemblage are I. formosus Naz. and Pseudobipennatus ziegleri Kon. et Kim, as well as I. arkonensis Stauff. and I. jejunus Naz. This assemblage is the most diverse and is recognized in all boreholes. The deposits containing this assemblage correspond to the maximum of the Nara transgression. The upper assemblage is also recognized in all the studied boreholes. Its index species is Ps. ziegleri Kon. et Kim. The Mosolovian Regional Stage is assigned to the kockelianus Zone. The distribution diagrams of species in the most diverse middle assemblage support the hypothesis that, in the southeastern regions of the Voronezh anteclise, the depth was greater than in its western part. The characteristic conodont species are shown in the plates.     

A neoselachian shark fauna from the Late Cretaceous of Senegal

A selachian fauna is described for the first time from the Late Cretaceous (Campanian–Maastrichtian) of Senegal. So far, the Campanian Paki Formation has only yielded a single tooth of Rhombodus sp. whereas the Cap de Naze Formation has yielded a more diverse fauna including juvenile Cretalamna cf. Cretalamna biauriculata, Serratolamna serrata, Carcharias cf. Carcharias heathi, ?Carcharias sp., Squalicorax pristodontus, Schizorhiza stromeri, Parapaleobates sp., Rhombodus binkhorsti and Rhombodus andriesi. Teeth of juvenile Cretalamna largely dominate the assemblage. Such an assemblage confirms a Late Maastrichtian age for the unit 3 in the Cap de Naze Formation. The assemblage, although composed of cosmopolitan taxa, is similar to the contemporaneous selachian assemblage from the phosphates of Morocco.     

Recent foraminifers from the inner shelf of the central West Coast,India: A reappraisal using factor analysis

Setty and Nigam (1980) had described 72 species of benthonic foraminifers from 25 inner shelf stations off central West Coast of India and the results showed somewhat patchy and anamolous distributions. This paper presents the results ofQ-mode factor analysis, which was applied to reduce the number of variables into assemblages. The analysis reveals 4 important foraminiferal assemblages.Ammonia beccarii—Ammonia annectens assemblage,Nonion boueanum—Florilus scaphum assemblage.Trochammina inflata assemblage andBulimina exilis assemblage. They can be related to freshwater run-off and organic matter contents of the sediment.     

Bolorian and Kubergandian stages of the Permian in the Sanandaj-Sirjan zone of Iran

Sections and fusulinids of the Bolorian (presumably) and Kubergandian (lower part) stages in the Sanandaj-Sirjan tectonic zone are described. Two fusulinid assemblages are distinguished in a most complete section near Sirjan. The lower one is represented by Skinnerella, Paraleeina, and relatively primitive Misellina forms, whereas Armenina, Kubergandella, and Yangchienia species appear in the upper assemblage and suggest its early Kubergandian age. Accordingly, the lower assemblage is attributed to the Bolorian Stage, although it is lacking fusulinids typical of this stage except for the primitive Misellina morphotypes. Fusulinids from the Tange-Darchaleh section near Qomsheh (Shahreza) are typical of the Kubergandian Stage. The described three new species of the genus Skinnerella are close to morphotypes known from younger (Murgabian) deposits and represent their ancestral forms most likely.     

Permian macro- and miofloral diversity,palynodating and palaeoclimate implications deduced from the coal-bearing sequences of Singrauli coalfield,Son–Mahanadi Basin,central India

The coal-bearing sequences of Barakar and Raniganj formations exposed in Bina and Jhingurdah open-cast collieries, respectively, are analysed for their macro- and miofloral content. The sediment successions primarily comprise of sandstones, shales, claystones and coal seams. In addition to the diverse glossopterid assemblage, four palynoassemblage zones, namely Zones I and II in Bina Colliery and Zones III and IV in Jhingurdah Colliery, have also been recorded in the present study. The megafossil assemblage from the Barakar strata of Bina Colliery comprises of three genera, namely Gangamopteris, Glossopteris and cf. Noeggerathiopsis. Palynoassemblage-I is characterised by the dominance of non-striate bisaccate pollen genus Scheuringipollenites and subdominance of striate bisaccate Faunipollenites and infers these strata to be of Early Permian (Artinskian) age (Lower Barakar Formation). The palynoassemblage has also yielded a large number of naked fossil spore tetrads, which is the first record of spore tetrads from any Artinskian strata in the world and has a significant bearing on the climatic conditions. The palynoassemblage-II is characterised with the dominance of Faunipollenites over Scheuringipollenites and is indicative of Kungurian age (Upper Barakar Formation). The megafossil assemblage from the Raniganj Formation of Jhingurdah Colliery comprises of five genera with 26 species representing four orders, viz., Equisetales, Cordaitales, Cycadales and Glossopteridales. The order Glossopteridales is highly diversified with 23 taxa and the genus Glossopteris, with 22 species, dominates the flora. The mioflora of this colliery is represented by two distinct palynoassemblages. The palynoassemblage-III is correlatable with the palynoflora of Early Permian (Artinskian) Lower Barakar Formation. The assemblage suggests the continuity of older biozones into the younger ones. The palynoassemblage-IV equates the beds with composition V: Striatopodocarpites–Faunipollenites–Gondisporites assemblage zone of Tiwari and Tripathi (1992) of Late Permian (Lopingian) Raniganj Formation in Damodar Basin. The FAD’s of Alisporites, Klausipollenites, Falcisporites, Arcuatipollenites pellucidus and Playfordiaspora cancellosa palynotaxa in this assemblage enhance the end Permian level of the Jhingurdah Top seam, as these elements are the key species to mark the transition of Permian into the Lower Triassic.     

Geothermobarometry of a stilpnomelane–garnet-bearing metapegmatite: P–T constraints on the Eo-Alpine metamorphic overprint of the Austroalpine nappes north of the Tauern Window

The Ordovician Kellerjochgneiss (Schwaz Augengneiss) is a polymetamorphic orthogneiss-bearing unit and is part of the Austroalpine basement nappes north of the Tauern Window. Within the Kellerjochgneiss a small, strongly deformed metapegmatite dike occurs. The pegmatite crosscuts the gneiss discordantly and contains the mineral assemblage muscovite 1,2+plagioclase+K-feldspar+chlorite+quartz+garnet 1 (Alm_67–76Andr_0.9–2Sps_17–28Prp_0.4–5)+garnet 2 (Grs_36–46Alm_24–32Andr_8–21Sps_15–17Prp_0–1)±stilpnomelane±biotite±clinozoisite. The magmatic protolith assemblage is comprised of relict K-feldspar, quartz and garnet 1. Textural observations indicate that biotite and muscovite cores (muscovite 1) are either part of the magmatic- or an earlier (Variscan?) metamorphic assemblage. Geothermobarometry of the metapegmatite was done on the latest-stage (Eo-Alpine) mineral assemblage garnet 2+muscovite 2+chlorite+stilpnomelane+plagioclase+quartz. Calculations of H₂O-absent intersections in the system [KCNFMAS] with the multi-equilibrium program THERMOCALC v.3.1 yielded P–T estimates of 4.4 to 6.7 kbar and 321°C to 376°C. Calculations of the P–T conditions by using the assemblage muscovite 2+chlorite+stilpnomelane+quartz yielded slightly higher pressures of 6.4 to 7.2 kbar at temperatures of 310–325°C. Correlating these P–T data with geochronological data from the neighbouring lithologies (Kellerjochgneiss, Innsbruck Quartzphyllite, Wildschönau Schists) and with structural investigations from these units indicate that the P–T estimates obtained in this investigation represent the Eo-Alpine metamorphic overprint. Hence, these unusual rocks provide important information on the Eo-Alpine P–T conditions since most samples studied from the investigated Austroalpine basement nappes north of the Tauern Window rarely contain mineral assemblages suitable for geothermobarometry.     

Smaller foraminifers from type sections of the Bolorian stage,the Lower Permian of Darvaz

Smaller foraminifers from Upper Yakhtashian and Bolorian deposits of the stratotype area (Pamir, Darvaz, Tajikistan) are investigated. Four assemblages are defined. The first assemblage is from Chalaroschwagerina vulgaris-Pamirina darvasica Zone. The second assemblage found in the transitional Yakhtashian-Bolorian beds includes Globivalvulina, Palaeotextulariidae, Hemigordiidae, and Glomospira, associated with the first Pachyphloia and Langella forms. Characteristic taxa of third assemblage from the Misellina (Brevaxina) dyhrenfurthi Zone are Geinitzinidae, Globivalvulina, Palaeotextulariidae, Glomospira, and rare Pachyphloia. The forth assemblage of Hemigordiidae, Pachyphloia, Palaeotextulariidae, Geinitzinidae, Pseudoagathammina is identified in the M. (Misellina) parvicostata Zone. The assemblages were compared with concurrent analogs from China, Japan, and Russia. New species and subspecies Glomospira paleograndis sp. nov., G. darvasica sp. nov., Agathammina darvasica sp. nov., Pachyphloia darvasica sp. nov., Nodosinelloides cubanicus elongatus subsp. nov., and Hemigordius saranensis darvasicus subsp. nov. are described.     

The Sinian microfossils from Jinning,Yunnan, south west China

A microfossil assemblage has been detected in phosphorite cherts from the Dengying and Meishucun Formations (610 Ma) of the Sinian System in Jinning County, Yunnan Province, southwest China.A unique type (mode) of microfossil preservation — collophanite or apatite, or even in rare cases, chert-replacement — is reported here for the first time.The size frequency statistics for the assemblage are consistent with the results obtained by Schopf. The assemblage includes 18 genera and 20 species, among which 4 genera and 10 species are new. The assemblage consists of solitary and colonial spheroidal cells and septate or non-septate filaments, mainly Cyanophyta, especially Chroococaceae, Oscillatoriaceae and Nostocaceae.The microfossils described are Bigeminococcus grandis n. sp., Myxococcoides kingii, Myxococcoides grandis, Palaeoanacystis vulgaris, Archaeophycus venustus n.g. et sp., Eomycetopsis robusta, Circulinema jinningense n.g. et sp., Circulinema muirae n.g. et sp., Cyanonema infliatum, Obruchevella parva, Oscillatoriopsis awramikii n. sp., Palaeolyngbya oehlerii n. sp., Palaeolyngbya spiralis n. sp., Palaeotubulus lamellosus n.g. et sp., Veteronostocale sp. 1, Veteronostocale sp. 2, Allantomorphus septatus n.g. et sp., Clonophycus vacus n. sp., Archaeotrichion contortum, Rhienonema antiquum, Micrhystridium setulerum and Protosphaeridium densum.     

A new assemblage of ray-finned fishes (Teleostei) from the Lower Oligocene “Schistes à Meletta” from the Glières plateau, Bornes Massif, eastern France

A recently discovered fish assemblage from the “Schistes à Meletta” facies (Lower Oligocene) of the Glières Plateau, Bornes Massif, Haute–Savoie, eastern France is described. The assemblage, comprising specimens ranging from fully-articulated skeletons to isolated scattered ossifications, is composed of Anenchelum cf. glarisianum, Pristigenys sp., Fistularia sp., Caranx cf. glarisianus alongside indeterminate teleosts. This new assemblage greatly increases the fish diversity previously known from the “Schistes à Meletta” of this area. The fauna shows biogeographic affinities with assemblages from the Peritethys domain, in particular from the Helvetic molassic basin and from the Paratethys. The genera identified at the Glières locality are represented today by species living in tropical nearshore environments together with species from the open sea. A possible explanation is that the environment of deposition was rather deep, and that shallow-water fishes were brought in by turbidity currents.     

New data on diatoms from the marine Paleogene of Western Kamchatka

Diatom assemblages from the marine Paleogene of Western Kamchatka (the Kovachina, Viventek, and Kuluven formations) are studied. The shallow-water neritic assemblage from the Kovachina Formation of the Pyatibratskii Cape section is tentatively attributed to the middle Eocene, as it is lacking zonal index species. A high abundance and great taxonomic diversity are characteristic of diatoms from the upper part of the Viventek Formation in the Kovran River section. The zonal index species Cavitatus rectus identified here among the diatoms suggests that the respective assemblage is macerated from the synonymous diatom zone of the middle Oligocene (29.6?28.2 Ma). According to its taxonomic composition and structure, this assemblage is indicative of transgression environments and the presumable influence of the middle Oligocene climatic optimum. The Rocella gelida (upper part) and Thalassiosira praefraga diatom zones of the Oligocene-Miocene transition are established in the Kuluven Formation sediments of the Mt. Uvuch section in Kovachina Bay.     

The shark-beds of the Eyam Limestone Formation (Lower Carboniferous,Viséan) of Steeplehouse Quarry,Wirksworth, Derbyshire,UK

The Eyam Limestone Formation of Steeplehouse Quarry, Wirksworth, Derbyshire, UK yields a diverse assemblage of Lower Carboniferous vertebrate remains. The assemblage is dominated by dermal denticles of the enigmatic selachian Petrodus patelliformis M’Coy, 1848, but also contains teeth of petalodonts, hybodonts and neoselachians. Actinopterygian remains also occur. The assemblage has yielded the earliest Neoselachian, Cooleyella fordi (Duffin and Ward, 1983) and the earliest British lonchidiid, Reesodus wirksworthensis (Duffin 1985). The first occurrence of the enigmatic spiny shark Acanthorhachis (Listracanthidae) is reported from the Viséan, extending its range back some 10 million years. Associated invertebrate remains and sedimentological data indicates a thriving fore-reef environment, deposited in a low energy off-reef setting. The vertebrate remains are well preserved with little abrasion, indicating short transport distances. Conodont elements indicating a late Brigantian age (Early Carboniferous, Viséan) have unusual and extensive euhedral apatite overgrowths.     

Palynological evidence for dating Jurassic-Cretaceous boundary sediments in the Bureya basin, Russian Far East

Palynological complexes from the coaliferous Talyndzhan and Dublikan formations of the Bureya sedimentary basin are analyzed. The palynological assemblage from the upper part of the Talyndzhan Formation is characterized by dominant gymnosperms largely close to Pinaceae and Ginkgocycadophytus. The content of ferns is insignificant against the background of their relatively high taxonomic diversity. The assemblage is marked by the last occurrence of Staplinisporites pocockii, Camptotriletes cerebriformis, C. nitida, and Cingulatisporites sanguinolentus spores typical of the Late Jurassic palynofloras. The palynological assemblage from the Dublikan Formation is dominated by Pteridophyts representing mainly by Cyathidites and Duplexisporites. In addition to the conifer, the role of Classopollis increased among the gymnosperms in this assemblage. It also includes the first-appearing Stereisporites bujargiensis, Neoraistrickia rotundiformis, Contignisporites dorsostriatus, Duplexisporites pseudotuberculatus, D. rotundatus, Appendicisporites tricostatus, and Concavissimisporites asper. These sporomorphs are characteristic of the Berriasian palynofloras. Thus, the Jurassic-Cretaceous boundary is most likely located between the Talyndzhan and Dublikan formations.     

U–Pb Zircon Ages of Early Cretaceous Volcanic Rocks in the Tethyan Himalaya at Yangzuoyong Co Lake,Nagarze, Southern Tibet,and Implications for the Jurassic/Cretaceous Boundary

Upper Jurassic–Lower Cretaceous transitional successions are widely distributed in the Tethyan Himalaya, southeast of Yangzuoyong Co Lake, southern Tibet. In ascending order, these include the Weimei (J₃, Tithonian), Sangxiu/Jiabula formations (K₁, Berriasian). The J/K boundary is located between the Weimei Formation and Sangxiu/Jiabula Formations. Ammonites found in J/K boundary sections in the research area have been classified into three assemblages: Valanginites–Phyllopachyceras assemblage zone (Valanginian), Spiticeras–Thurmanniceras assemblage zone (Berriasian) and Haplophylloceras–Blanfordiceras–Himalayites assemblage zone (Tithonian). Six nannofossil zones: Calcicalathina oblongata assemblage zone, Speetonia colligate zone, N. st. steinmannii zone, N. st. minor zone, P. beckmanni–N. st. minor interval zone, Conusphaera–Polycostella–Nannoconus–Watznaueria assemblage zone were recognized as well.On the basis of lithology, biostratigraphy and geochronology of the J/K transitional deposition succession, this study suggests that the J/K boundary, in southern Tibet, is located on the bottom of P. beckmanni–N. st. minor interval zone, which is further definited as and disappear of Polycostella beckmanni. To address the paucity of previously reported reliable ages for the J/K boundary, this study reports four U–Pb zircon ages (140–142 Ma) obtained with Secondary Ion Mass Spectrometry (SIMS) from the volcanic rocks interbedded in the lower Sangxiu Formation, which is expected to provides a direct date reference for the J/K boundary in the Tethyan Himalaya, southern Tibet. From integration of our new (SIMS) U–Pb zircon ages with calcareous nannofossils and ammonites, the age of the N. st. minor zone (NK-D) directly above the P. beckmanni-N. st. minor interval zone (NJK-C) of the basal Berriasian in the Tethyan realm is estimated to be 141–142 Ma. This research is not only helpful to improve the isotopic determination of absolute age for the J/K boundary, but also implies that the Tethyan Himalaya of southern Tibet may be an ideal location in which to explore the J/K boundary in both biostratigraphy and geochronology in future.     

Calculation of pH and mineral equilibria in hydrothermal waters with application to geothermometry and studies of boiling and dilution

Using chemical analyses and 25° pH measurements of quenched high-temperature waters, we calculate in situ pH and distribution of aqueous species at high temperature. This is accomplished by solving simultaneous mass action equations for complexes and redox equilibria and mass balance equations, on all components, including a H⁺ equation with as many as 60 terms (depending on water composition). This calculation provides accurate values for the activities of aqueous ions in a given water at high temperature, which are used to calculate an ion activity product (Q) for each of more than 100 minerals. The value of log(Q/K) for each mineral, where K is the equilibrium constant, provides a measure of proximity of the aqueous solution to equilibrium with the mineral. By plotting log Q/Kvs. T for natural waters, it is possible to determine: a) whether the water was in equilibrium with a host rock mineral assemblage, b) probable minerals in the equilibrium assemblage and c) the temperature of equilibrium. In cases where the fluid departs from equilibrium with a host rock assemblage, it is possible to determine whether this may result from boiling or dilution, and an estimate of amount of lost gas or diluting water can be determined.The calculation is illustrated by application to geothermal waters from Iceland, Broadlands, and Sulphur Bank, hot spring waters from Jemez, Yellowstone and Blackfoot Reservoir (Idaho) and fluid inclusions from the Sunnyside Mine, Colorado. It is shown that most geothermal waters approach equilibrium with a subsurface mineral assemblage at a temperature close to measured temperatures and that some hot springs also approach equilibrium with the host rock at temperatures above outlet temperatures but commonly below the Na-K-Ca temperatures. The log Q/K plots show that some discrepancies between Na-K-Ca temperatures on spring waters and actual temperatures result from a failure of alkali feldspars to equilibrate with the fluid and with each other.Calculations on Sulphur Bank fluids show that boiling probably caused cinnabar precipitation near 150°C and that the boiled fluids equilibrated with secondary minerals near 150° even though temperatures up to 185° have been measured at depth. For the fluid inclusions, the measured bubble temperatures are close to those calculated for equilibration of the fluid with the observed sulfide mineral assemblage.New estimates of stability constants for aluminum hydroxide complexes are included at the end of the paper.     

Recognizing cryptic environmental changes by using paleoecology and taphonomy of Pleistocene bivalve assemblages in the Oga Peninsula,northern Japan

Multivariate analyses applied to Pleistocene bivalve assemblages from the Oga Peninsula (northern Japan) discriminate three distinct assemblages. The assemblages and their taphonomy were used to recognize environmental settings and changes. The Astarte–Cyclocardia–Glycymeris assemblage indicates shelf environment (below the storm wave base) where gravels and shells are transported from shallower settings. Supply of the exotic coarse sediment probably enabled epifaunal bivalves to inhabit the sea floor. The Glycymeris assemblage is characterized by dominance of G. yessoensis and represents current-swept shoreface environment (above the storm wave base). The Moerella assemblage is characterized by bivalves inhabiting bay to open-marine conditions and diverse deposit-feeders, indicating a moderately land-locked environment, such as an open bay or a bay mouth. Fine-grained substrata rich in organic matters in the bay were probably suitable for the deposit-feeders. Ordination also shows the assemblages along two environmental gradients, a bathymetrical one and the other related to open-marine and bay conditions. The environmental changes are explained mainly by glacio-eustatic sea-level changes and alternation of coastal geomorphology caused by local crustal movements. This study also suggests that fossil assemblages can be a powerful tool to reconstruct environments and depositional dynamics even in intensely bioturbated sedimentary facies.     

Dinosaur eggshells from the Santonian Milk River Formation of Alberta,Canada

The North American fossil record of dinosaur eggshells for the Cretaceous is primarily restricted to formations of the middle (Albian–Cenomanian) and uppermost (Campanian–Maastrichtian) stages, with a large gap in the record for intermediate stages. Here we describe a dinosaur eggshell assemblage from a formation that represents an intermediate and poorly fossiliferous stage of the Upper Cretaceous, the Santonian Milk River Formation of southern Alberta, Canada. The Milk River eggshell assemblage contains five eggshell taxa: Continuoolithus, Porituberoolithus, Prismatoolithus, Spheroolithus, and Triprismatoolithus. These ootaxa are most similar to those reported from younger Campanian–Maastrichtian formations of the northern Western Interior than they are to ootaxa reported from older middle Cretaceous formations (i.e., predominantly Macroelongatoolithus). Characteristics of the Milk River ootaxa indicate that they are ascribable to at least one ornithopod and four small theropod species. The taxonomic affinity of the eggshell assemblage is consistent with the dinosaur fauna known based on isolated teeth and fragmentary skeletal remains from the formation, although most ornithischians and large theropods are not represented by eggshell. Relative to the Milk River Formation eggshell, similar oospecies occurring in younger Cretaceous deposits tend to be somewhat thicker, which may reflect an increase in body size of various dinosaur lineages during the Late Cretaceous.