Fossil pollen from Los Toldos locality: A record of the Late-glacial transition in the Extra-Andean Patagonia

Paleoenvironmental changes during the Late-glacial transition are interpreted from a pollen record from two caves at Los Toldos (47°22′S; 68°58′W) in Extra-Andean Patagonia in Argentina. The paleoenvironmental interpretation is based on changes in the ratio between shrub and grass steppe taxa and on comparison with other pollen records from the region. Between 12,600 and 8750 yr BP two vegetational changes occurred thought to reflect the establishment of Holocene-type atmospheric circulation patterns. The first change is at ca. 11,000 yr BP expressed by replacement of shrub steppe with Ephedra by a grass steppe. The second change occurred ca. 10,000 yr BP when the grass steppe was replaced by a shrub steppe dominated by Asteraceae. Before ca. 11,000 yr BP the environmental conditions were extremely arid with precipitation lower than 200 mm. Between ca. 11,000 and ca. 10,000 yr BP effective moisture increased, probably related to an increase in precipitation to about 200 mm under cold conditions. A grass steppe extended through Patagonia and Tierra del Fuego, at least as far north as 47°S. It is difficult to find modern analogues for pollen associations south of 47°S older than 10,000 yr BP. Possibly at that time climate patterns were markedly different from today. At about this time of environmental changes Level 11 industry associated with extinct grazing herbivores developed. Starting at ca. 10,000 yr BP, with expansion of the shrub steppe of Asteraceae east of the Andes at 47° to 52°S and forest at 51°–54°S west of the Andes, temperature increased while water availability decreased. Precipitation probably was similar to the present, which suggests that the modern climate patterns were established at the beginning of the Holocene. The Toldense Industry, which is associated with remains of modern fauna as well as with the last remains of the Pleistocene fauna, developed in this period. The beginning of this development coincides with the extinction of Pleistocene fauna.     

Mammoth tracks indicate a declining Late Pleistocene population in southwestern Alberta,Canada

Much debate has raged over the role that early humans played in this most recent large extinction. Fossil mammoth (Mammuthus primigenius) footprints were discovered at the St. Mary Reservoir in southwestern Canada (Wally's Beach DhPg-8). They are located in aeolian sediment dated at 11,300–11,000 years BP. By comparing the size distribution of these tracks with those of modern African elephants (Loxodonta africana), the age distribution of this mammoth population was determined. Containing far fewer juveniles than would be expected for an expanding or stable population, these tracks provide the first evidence that a living mammoth population, coexisting with human inhabitants, was in decline. Additionally, the same site provides corroborating evidence of humans hunting megafauna (horse and bovids). This suggests that humans, in addition to climate change, played a role in the end Pleistocene extinctions in North America.     

Paleoecological perspectives on human adaptation in central Panama. I. The Pleistocene

Paleoecological data from two sites in central Pacific Panama have allowed the reconstruction of Late Pleistocene climate and vegetation in lowland areas, the timing of important environmental changes, and the generation of predictions concerning Paleoindian settlement and subsistence. The last 9000 years of the glacial period, from 20,000 B.P. until 11,050 B.P. were marked by climates cooler and drier than today's. The period from ca. 16,000 B.P. until 11,000 B.P. appears to have been the coolest and driest. We postulate that the major effects of these conditions were to have brought montane vegetation 900 m lower than its present range and to create tracts of open landscape along the Pacific coast. Forests, however, were widespread in extent and many lowland forest taxa apparently persisted, creating the basis for the rejection of refugial theory as an explanatory model for early human occupation. Neither the admittedly limited archaeological or paleoecological data indicated the presence of pre-Clovis populations in Panama, and we chose to view Clovis as the first human expression on the Isthmus. Correlation of reconstructed environmental setting with Paleoindian site location suggests that Clovis adaptations were fluid and flexible, and utilized a wide variety of vegetation types. Occupation and modification of tropical forest appear to have been integral parts of Clovis subsistence and settlement strategies.     

The Isthmus of Panama as a Crossroad for Prehistoric Migration of Domesticated Plants

For thousands of years, Indian tribes have gathered, selected, domesticated and spread useful plants over the whole American continent while mostly practicing shifting forms of horti-agriculture. It has been argued that origin and domestication of New World crops could be traced back to essentially two independent, narrowly circumscribed core regions, (i.e. Vavilov's genecentres) one in the highlands of Mexico and the other in the Andes of Peru. Those alleged centers of origin and genetic diversity have been synchronized with the main cradles of American agriculture. Based on long periods of research work in South America, we disagree with this opinion which is maintained primarily by anthropologists, sociologists and ethnologists. Except for marginal desert and mountain environments, there are no real biogeographical restrictions for plant domestication. Apparent geographical barriers, like the Darien gap, did not represent insurmountable hindrances for the diffusion and migration of useful species in the hands of indigenous peoples.     

Investigating human and megafauna co-occurrence in Australian prehistory: Mode and causality in fossil accumulations at Cuddie Springs

Human arrival in Sahul – Pleistocene Australia and New Guinea – has long been argued as the catalyst in the decline and disappearance of a suite of extinct animals referred to as megafauna. The debate concerning causality in Sahul is highly polarised, with climate change often cited as the alternative explanatory model. On continental Australia, there are few datasets available with which to explore the likely processes leading to the extinction events. At the present time, there is one site in New Guinea (Nombe Rockshelter) and one on continental Australia (Cuddie Springs) where the coexistence and temporal overlap of humans and megafauna has been identified. The Cuddie Springs Pleistocene archaeological site in southeastern Australia contains an association of fossil extinct and extant fauna with an archaeological record through two sequential stratigraphic units dating from c. 36 to c. 30 ka ago. A taphonomic study of the fossil fauna has revealed an accumulation of bone in a primary depositional context, consistent with a waterhole death assemblage. Overall the faunal assemblage studied here (n: 8146; NISP: 1355) has yielded little direct evidence of carnivore damage or human activities. Post depositional factors such as physical destruction incurred by trampling, compaction of sediments, and/or the hydrological status of the lake at that time have played important roles. As the only known site on continental Australia where megafauna and humans co-occur, the Cuddie Springs faunal assemblage yields equivocal evidence for a significant human role in the accumulation of the fauna here. At the present time there is no evidential basis to the argument that humans had a primary role in the extinction of the Australian megafauna. The first colonisers are likely to have preyed upon those few species known to have persisted to this time, but their impact may have been restricted to the tail end of a process that had been underway for millennia prior to human arrival.     

Paleoclimatic implications of late quaternary lacustrine sediments in Western Nubia, Sudan

Owing to the hypercontinental location of Western Nubia, secular fluctuations of climate have been filtered and wet phases can be considered as representative of conditions throughout the southeastern Sahara. The study area is crossed by the 20-mm isohyet; between 9300 and about 4000 yr B.P., however, there were widespread lake and swamp environments with freshwater molluscs, ostracods, and diatoms, and a species-rich savanna mammal fauna. The center of the West Nubian Basin (approx. 18°N), an area of about 20,000 km², was occupied by a semiaquatic landscape which was situated at the same latitude as Paleolake Chad. From extensive lake carbonates up to about 4 m thick, a long-term rise of the groudwater table is inferred. Environments developed that now exist at about latitude 13°N. Radiocarbon dates from lake sediment sequences cluster between 30,000 and 21,000 yr B.P., indicating a Pleistocene wet phase. A gap in radiocarbon dates between 21,000 and 11,000 yr B.P. signals a phase of hyperaridity, similar to the present hyperarid phase, with eolian deflation and deposits of sand being the dominant forms of erosion and accumulation.     

Extinction of the woolly mammoth (Mammuthus primigenius) and woolly rhinoceros (Coelodonta antiquitatis) in Eurasia: Review of chronological and environmental issues

Kuzmin, Y. V. 2009: Extinction of the woolly mammoth (Mammuthus primigenius) and woolly rhinoceros (Coelodonta antiquitatis) in Eurasia: Review of chronological and environmental issues. Boreas, 10.1111/j.1502‐3885.2009.00122.x. ISSN 0300‐9483. The current evidence for date and environmental preferences of the extinction of two middle–late Pleistocene megafaunal species, the woolly mammoth (Mammuthus primigenius Blum.) and woolly rhinoceros (Coelodonta antiquitatis Blum.), is presented in this review. It is suggested that extinction of these large herbivores in Eurasia was closely related to landscape changes near the Pleistocene–Holocene boundary (c. 12 000–9000 uncalibrated radiocarbon years ago, yr BP), mainly involving the widespread forest formations in the temperate and arctic regions of northern Eurasia and the loss of grasslands crucial to the existence of woolly mammoth and rhinoceros. However, some woolly mammoth populations survived well into the Holocene (up to c. 3700 yr BP), showing that the process of final extinction was fairly complex, with delays in some regions of up to several millennia. The possible role of Palaeolithic humans in the extinction of Late Pleistocene megafauna is also considered.     

A late Pleistocene human presence at Huaca Prieta,Peru, and early Pacific Coastal adaptations

Archaeological excavations in deep pre-mound levels at Huaca Prieta in northern Peru have yielded new evidence of late Pleistocene cultural deposits that shed insights into the early human occupation of the Pacific coast of South America. Radiocarbon dates place this occupation between ~ 14,200 and 13,300 cal yr BP. The cultural evidence shares certain basic technological and subsistence traits, including maritime resources and simple flake tools, with previously discovered late Pleistocene sites along the Pacific coast of Peru and Chile. The results help to expand our knowledge of early maritime societies and human adaption to changing coastal environments.     

Quaternary climate,environment, and the human occupation of the south-central Andes

This paper reviews the paleoenvironmental context for Paleoindian and Early Archaic settlement of the south-central Andes. I attempt to reconcile proposals for late-glacial and early Holocene environmental changes that have originated in several Quaternary sciences. Most useful are the records of changing lake levels, geomorphological indicators of glacial advance and retreat, changes in vegetation as seen through pollen stratigraphy, the record of large mammal extinctions, and the archaeological settlement pattern itself. High lake stands in the central and south-central Andes, c. 12,500 to 11,000 B.P., seem to correspond to glacial retreat. Palynological studies also suggest higher temperatures, coincident with greater summer precipitation from 11,500 B.P. to perhaps 10,000 B.P., followed by some reduction in temperatures and then widespread aridity from about 8500 to 5000 B.P. Environmental change at the Pleistocene/Holocene boundary was at least a major contributing factor to the extinction of horse and sloth around 10,000 B.P. Archaeology of the salt puna and Punta Negra region conforms well with the environmental record, showing human entry by the Early Archaic (11,000-8000 B.P.), subsequent depopulation and perhaps abandonment, then readaptation to more localized resources in the Late Archaic (5500-4000 B.P.).     

The Permian–Triassic mass extinction: Ostracods (Crustacea) and microbialites

The end-Permian mass extinction (EPE), about 252 Myr ago, eradicated more than 90% of marine species. Following this event, microbial formations colonised the space left vacant after extinction of skeletonised metazoans. These post-extinction microbialites dominated shallow marine environments and were usually considered as devoid of associated fauna. Recently, several fossil groups were discovered together with these deposits and allow discussing the palaeoenvironmental conditions following the EPE. At the very base of the Triassic, abundant Ostracods (Crustacea) are systematically present, only in association with microbialites. Bacterial communities building the microbial mats should have served as an unlimited food supply. Photosynthetic cyanobacteria may also have locally provided oxygen to the supposedly anoxic environment: microbialites would have been refuges in the immediate aftermath of the EPE. Ostracods temporarily disappear together with microbialites during the Griesbachian.     

Chronological overlap between humans and megafauna in Sahul (Pleistocene Australia–New Guinea): A review of the evidence

Over 60 faunal species disappeared from the Australian continent during the Middle–Late Pleistocene. Most of these animals were large to gigantic marsupials, birds and reptiles. A terminal extinction date of 46.4 kyr has been proposed for the megafauna, with all sites containing younger fossil megafauna dismissed by some researchers because of questions over stratigraphic integrity or chronologies. The timing of the extinctions is argued to be broadly coincident with estimates of first colonization of the continent by modern humans, and explanatory extinction models involving humans have subsequently gained currency. However there is considerable evidence to suggest that in some parts of the continent, people and some species of megafauna may have co-existed well beyond 46.4 kyr. In other places, such as Tasmania and the north of the continent, there is no known record of a human–megafauna temporal overlap. A review of the available evidence indicates that only 13 species of megafauna were extant on human arrival in Australia. The archaeology of this period indicates that rather than a focus on big game hunting or ‘firestick farming’, it was characterized by regional variability in subsistence strategies consistent with the range of environmental zones. At the present time there is no substantive argument for a terminal extinction date of 46.4 kyr, the current evidence indicating that there is no specific time period that correlates to any single mass extinction event. On the basis of available evidence arguments for either human or climatic causation are entirely circumstantial and implicitly require acceptance of many unproven assumptions. Claims to have eliminated climate as a primary driver are premature and the recent focus on delivering ‘proof’ of human causation in Pleistocene faunal extinctions diverts attention from achieving a better understanding of the differential impacts of climate change and short term climatic flux in a land of environmental extremes.     

双河洞中大熊猫-剑齿象动物群化石的发现

对贵州省境内的双河洞历次科考中发现的第四纪动物群化石进行初步研究和总结,认为目前对双河洞第四纪动物群研究尚处于起步阶段,化石的发现者都是洞穴科考专家,古生物知识相对欠缺且重视不够是洞穴化石研究进展缓慢的原因。为了加快双河洞第四纪动物群的研究进程,以今年中法联合科考过程中发现的大熊猫-剑齿象动物群化石为样本,采用ICP-MS铀系定年方法对这批大熊猫骨骼化石碳酸盐岩包裹物的年龄进行测试,结果表明该化石群化石距今6.75万年,初步判定此动物群可能生存于中更新世-晚更新世早期。这批哺乳动物化石及其地层信息将为贵州（尤其是黔北）第四纪的研究提供重要资料。      

北京环境考古

人类居住场所和生活方式受环境的显著影响。30,000年以来北京环境演变可分作6个阶段。旧石器晚期在干寒的环境下,古人类依洞穴而居。到新石器时代之初,气候变暖,有人类活动于山间谷地。后来,直到商、周时期,气候暖湿,人类多在山前台地上生活。到战国至西汉,气候明显变干变凉,人类向泛滥平原转移。北京的先民居住场所迁移的总趋势是,由山地而平原,由台地而低谷。农业兴起于新石器时代早、中期的山前二级台地上,发达于历史时期的低谷地带。北京城的兴起,地利起了重要作用。     

我国东北地区晚更新世中晚期环境变化与猛犸象—披毛犀动物群绝灭研究综述

通过对文献资料的总结、统计分析,结合对我国东北地区的实地考察和初步研究结果,论述了我国东北地区晚更新世中晚期以来环境变化与猛犸象—披毛犀动物群演化、灭绝的研究现状和存在问题及其与国际对相应时段猛犸象研究之间的差距。提出加强对我国东北地区晚更新世气候环境的重建、认识气候变化的区域特点和突变性,系统而可靠地建立以猛犸象—披毛犀为主的化石动物年代序列、区分主要类型化石动物年代是研究的关键。以独立的气候环境变化和化石动物的年代序列为参照系,在区域环境变化的框架内讨论化石动物群生存的气候环境、生态特征和灭绝过程,进而探讨动物群演化及灭绝与气候环境变化的关系。这将为认识区域环境突变、动植物对环境变化的响应、我国东北地区晚更新世猛犸象—披毛犀动物群与欧亚大陆其它地区猛犸象之间的内在联系,认识人类演化与生存环境的适应、动植物演替与绝灭的临界条件等一系列重大科学问题提供资料。     

Paleoecology of late-glacial peats from the bering land bridge, Chukchi Sea shelf region, northwestern Alaska

Insect fossils and pollen from late Pleistocene nonmarine peat layers were recovered from cores from the shelf region of the Chukchi Sea at depths of about 50 m below sea level. The peats date to 11,300−11,000 yr B.P. and provide a limiting age for the regional Pleistocene-Holocene marine transgression. The insect fossils are indicative of arctic coastal habitats like those of the Mackenzie Delta region (mean July TEMPERATURES = 10.6–14°C) suggesting that 11,000 yr ago the exposed Chukchi Sea shelf had a climate substantially warmer than modern coastal regions of the Alaskan north slope. The pollen spectra are consistent with the age assignment to the Birch Interval (14,000–9000 yr B.P.). The data suggest a meadow-like graminoid tundra with birch shrubs and some willow shrubs growing in sheltered areas.     

Paleoenvironments and cultural diversity in late Pleistocene South America

After a summary assessment of certain selected early man sites in various parts of America, the environment of the Venezuelan coastal plain is discussed in order to evaluate the stratigraphy and radiometric dating of the Taima-Taima site, near Coro, Venezuela, where mammals, many now extinct, were killed by people making El Jobo points about 13,000 years ago.Potentially important areas in Peru, Chile and Brazil are mentioned. Certain problems in the models and interpretation of South American paleoclimates are pointed out.The presence of at least four different bifacially flaked stone projectile point traditions in widely separated and environmentally diverse parts of America between 11,000 and 13,000 yr ago suggests that the immediate cultural antecedents of these traditions were essentially independent of one another. From this it is argued that several early American flaked-stone point traditions developed indigenously in America from technological bases which were present in the Old World Middle Paleolithic.     

Glacial history of southernmost South America

In southernmost South America, an incomplete radiometrically dated glacial chronology has been obtained by KAr dating for the interval 3.5-1 MY ago, and a more detailed chronology by C-14 dating for the last 25,000 years, with some older minimal ages. The first major glaciation was about 3.5 MY ago during the middle Pliocene. Little is yet known about glacial fluctuations during the interval 3.5-2.1 MY ago. Between 2.1 and 1 MY ago many glaciations occurred, probably including the greatest of late Cenozoic time which took place after 1.2 MY and, according to inconclusive evidence, before 1 MY ago. The Patagonian Gravel in its type area is mid-Pliocene to early Pleistocene glacial outwash that accumulated from the first to the greatest glaciations. During the late Pleistocene several glaciations occurred, but only the most recent has been radiometrically dated. During the last glaciation the glaciers were most extensive before 56,000 BP. Successively smaller advances that culminated about 19,500 BP and, probably, about 13,000 BP were separated by an interstade when glaciers shrank by more than half. The glaciers receded rapidly after 13,000 BP and were within their present borders by 11,000 BP; they remained so during the European Younger Dryas Stade 11,000-10,000 BP. Neoglacial regional readvances culminated 4600-4200 BP, probably 2700-2000 BP, and during the last three centuries; most glaciers reached their Neoglacial maxima during the first episode. Between readvances, the glaciers shrank within their present borders.     

On the character of Black Sea level rise during the Holocene

The character of changes in the mollusk fauna in the Late Pleistocene and Holocene sediments within the Bulgarian, Northwestern, Crimean, and Kerch segments of the Black Sea shelf has been examined. A bed containing fossil remains of brackish-water and marine mollusks was recognized in these sediments; the bed accumulated, based on results of radiocarbon datings, 500–1000 years ago. The obtained data do not support the existing notion of a catastrophic fill-up of the Black Sea Basin by marine Mediterranean water [19].     

Co-existence of scimitar-toothed cats,lions and hominins in the European Pleistocene. Implications of the post-cranial anatomy of Homotherium latidens (Owen) for comparative palaeoecology

Human occupants of Europe shared food resources with a number of larger Carnivora, and their coexistence with two lion-sized felids, the lion and the scimitar-toothed machairodont Homotherium latidens, poses intriguing paleoecological problems. We investigate the ecology of Homotherium latidens using an exceptional sample of postcrania from the Spanish Early Pleistocene site of Incarcal, making comparisons with modern cats and with other machairodont species. Evidence of cursorial adaptations in Homotherium suggests a hunting technique different from modern cats or smilodontine sabre-tooths. Some, like reduction of the claws, would have limited the ability of individual homotheres to bring down large prey, implying group action. Homotherium would also have been disadvantaged in direct confrontation with Pleistocene lions by smaller body mass, reduced forepaw muscle strength, smaller claws and more fragile dentition. Its hunting technique would have worked best in more open habitats, but competition from lions would have forced it to seek moderate cover. Among factors that could de-stabilise coexistence of the two big cat species in Pleistocene Europe we invoke a decrease in environmental mosaicism associated with stepped climatic change over the last million years, and the increased importance of humans within the larger predator guild.     

Age profiles in elephant and mammoth bone assemblages

Age profiles of modern African elephant (Loxodonta africana) populations are significantly affected by drought conditions that cause local die-offs. Subadult animals die in proportions that may be nearly twice what is recorded in live populations. Such biasing of death sample age profiles might also have occurred during late Pleistocene die-offs of Mammuthus. This comparative study of modern and fossil proboscidean age structures supports a tentative interpretation that late Pleistocene extinction of Mammuthus (at least in the southwestern United States) resulted from severe drought conditions, at which Clovis hunters were witnesses, but not necessarily frequent participants.