The Sima de los Huesos (Burgos,northern Spain): palaeoenvironment and habitats of Homo heidelbergensis during the Middle Pleistocene

Interpreting how environmental dynamics respond to global climate change and how this has affected human evolution and dispersal is an on-going topic of debate. During the early Middle Pleistocene (～0.6–0.4 Ma), as compared to earlier, environmental conditions were relatively more stable, with longer climatic cycles alternating between open and forested landscapes. During this interval, humans spread successfully providing an important number of fossil sites where fossils or tools are reported. The Atapuerca-Sima de los Huesos (Burgos, northern Spain) site (Atapuerca-SH) is one of the earliest localities with hominin evidence in the European Middle Pleistocene, with the most important accumulation of Homo heidelbergensis so far. We have analyzed the abundant faunal record from Sima de los Huesos, which is mainly comprised of carnivores, in order to approach an interpretation of the palaeoenvironmental circumstances where these hominids inhabited within the Sierra. Other sites from Sierra de Atapuerca referred to the same Faunal Unit (FU 6), are roughly contemporaneous, and include important ungulates, which are here analyzed with Atapuerca-SH. Additional information provided by isotopic analysis helps elucidate the ancient ecology of taxa present in Sima de los Huesos allowing for an accurate portrayal of the setting in which humans lived. The timing of the spread of Homo heidelbergensis is dominated by a relative climatic and environmental stability and points to a landscape dominated by savannah-like open woodland.     

Palaeomagnetism of Pliocene-Pleistocene sediments and lava flows on Tjörnes and Flatey, North Iceland

Palaeomagnetic measurements have been carried out on lava flows and volcaniclastic sedimentary rocks of Upper Pliocene and Pleistocene age from sections on the Tjömes Peninsula and a core section from the island of Flatey off North Iceland. Earlier palaeomagnetic work in the area is reviewed, and new data on the stratigraphy of the Hoskuldsvik Group on Tjörnes are presented. A revised polarity column based on our resultsisset upfortheoutcropson Tjörnes, including twoadditionalnormalsubchronswithin the Matuyama. Our palaeomagnetic results indicate that the rocks penetrated by the Flatey drillhole accumulated during the Matuyama. Age estimates for the Upper Tjömes sequence are reviewed, and correlation alternatives indicate that the earliest glaciation of Tjörnes took place either at c.2 Ma or near the Gauss/Matuyama boundary at c. 2.5 Ma. Correlation of individual Matuyama subchrons remains uncertain. A comparison of the climatic evidence from the Tjörnes area with data on oxygen isotope and carbonate variations at DSDP site 552A is used to evaluate correlation alternatives.     

The Upper Pleistocene to Holocene sediments on the Mediterranean island of Lampedusa (Italy)

The island of Lampedusa lies on the northern edge of the African continental shelf, but during some Quaternary marine lowstands it was joined to the African continent. The study and dating of the aeolian, alluvial, detrital sediments, calcareous crusts and speleothems have established that the climatic–environmental variations recorded on the island can be related chronologically to those known for northern Libya, Tunisia and the Italian peninsula. During the Last Glacial Maximum, phases of Saharan dust accumulation on Lampedusa occurred, and were coeval with dust accumulation in crater lakes and on high mountains in central‐southern Italy, and with phases of glacial advance in the Apennines and in the Alps. During the late Holocene, accumulation of Saharan dust on Lampedusa occurred but there was little accumulation of dust on the northern side of the Mediterranean Sea. With the new data from Lampedusa, it is possible to envisage two different scenarios of atmospheric circulation relating to the Last Glacial Maximum and to the late Holocene. During the Last Glacial Maximum, southerly atmospheric circulation brought rainfall to the southern slopes of the Alps and to the Apennines. During the late Holocene, a prevalent westerly atmospheric circulation became established in the northern Mediterranean. Copyright © 2004 John Wiley & Sons, Ltd.     

Dinoflagellate cyst stratigraphy and palaeoenvironment of the marginal marine Middle and Upper Miocene of the eastern Campine area,northern Belgium (southern North Sea Basin)

A stratigraphical and palaeoenvironmental analysis with organic‐walled palynomorphs of the Bolderberg and Diest formations provides new insights in the depositional history during Miocene times at the southern border of the North Sea Basin. The Neogene transgression invaded Belgium from a north–northwestern direction and fully marine sediments were deposited in the northern part of Belgium. The age and the palaeoenvironment of the deposits at the very border of the southern North Sea Basin remained till a few decades ago incomplete. The recovered dinoflagellate cysts, acritarchs and green algae from the Bolderberg Formation in the Wijshagen Borehole indicate a marginal marine depositional environment during late Burdigalian and Langhian times in the eastern Campine area, in contrast to the deeper marine conditions prevailing to the north–northwest. The relative dating of the Bolderberg Formation confirms that maximum flooding occurred during Langhian to early Serravallian times. Deposition apparently took place during the Middle Miocene Climatic Optimum, and ended when the climatic deterioration set in at around 14 Ma. A hiatus spanning ca. 2 Ma separates the Middle Miocene Bolderberg Formation from the Upper Miocene Diest Formation in the eastern Campine area at the border of the North Sea Basin. Copyright © 2007 John Wiley & Sons, Ltd.     

The palaeoenvironment associated with a partial Iguanodon skeleton from the Upper Weald Clay (Barremian, Early Cretaceous) at Smokejacks Brickworks (Ockley, Surrey, UK), based on palynomorphs and ostracods

In 2001 a partial skeleton of an Iguanodon was discovered in the Upper Weald Clay (Barremian, Early Cretaceous) at Smokejacks Brickworks near Ockley, Surrey, UK. When the dinosaur was excavated, a detailed stratigraphic section was logged and 25 samples taken for palynological and micropalaeontological (ostracod and megaspore) analysis, including a detailed sample set of the dinosaur bed itself. Qualitative and quantitative analysis of the palynoflora revealed rich and well-preserved non-marine assemblages of pollen and spores, including early angiosperms, and freshwater green algae. Four types of angiosperm pollen are described and assigned to the genus Retimonocolpites Pierce, 1961, but left in open nomenclature. Some marine elements such as dinoflagellate cysts are identified as the result of reworking of Middle and Upper Jurassic sediments. The pollen/spore assemblages depict a vegetational change from principally gymnosperm-dominated assemblages at the base to principally pteridophyte-dominated assemblages at the top of the section. The dinosaur bed shows a pteridophyte-dominated assemblage, with a significantly high amount of the freshwater green alga Scenedesmus novilunaris He Cheng-quan et al., 1992. Samples close to the dinosaur bed yielded the first useful ostracod finds from Smokejacks Brickworks: well-preserved assemblages containing Cypridea clavata (Anderson, 1939), Damonella cf. pygmaea (Anderson, 1941), Stenestroemia cf. cressida Anderson, 1971 and Stenestroemia sp. A, and fragments and damaged valves of a thin-shelled ostracod, possibly belonging to Mantelliana Anderson, 1966. Those identified as Cypridea clavata show a wide range of morphological variety and in opposition to Anderson's (1967, 1985) taxonomic scheme, which would assign them to up to five different taxa, they are considered to be intraspecific variants of a single species. The possibilities and limitations of age determination of the Wealden sediments using palynomorphs and ostracods are discussed; distinct forms of early angiosperm pollen, together with the ostracod fauna, are consistent with an early Barremian age. Pollen and spores are discussed in terms of their parent plants and the reconstruction of vegetation and palaeoclimate. Palynology and ostracods give evidence for temporary freshwater conditions at the time when the Iguanodon died and the carcase was buried.     

Late Quaternary palaeoenvironment of northern Guatemala: evidence from deep drill cores and seismic stratigraphy of Lake Petén Itzá

Long sediment cores were collected in spring 2006 from Lake Petén Itzá, northern Guatemala, in water depths ranging from 30 to 150 m, as part of an International Continental Scientific Drilling Program project. The sediment records from deep water consist mainly of alternating clay, gypsum and carbonate units and, in at least two drill sites, extend back >200 kyr. Most of the lithostratigraphic units are traceable throughout the basin along seismic reflections that serve as seismic stratigraphic boundaries and suggest that the lithostratigraphy can be used to infer regional palaeoenvironmental changes. A revised seismic stratigraphy was established on the basis of integrated lithological and seismic reflection data from the basin. From ca 200 to ca 85 ka, sediments are dominated by carbonate‐clay silt, often interbedded with sandy turbidites, indicating a sediment regime dominated by detrital sedimentation in a relatively humid climate. At ca 85 ka, an exposure horizon consisting of gravels, coarse sand and terrestrial gastropods marks a lake lowstand or partial basin desiccation, indicating dry climate conditions. From ca 85 to ca 48 ka, transgressive carbonate‐clay sediments, overlain by deep‐water clays, suggest a lake level rise and subsequent stabilization at high stage. From ca 48 ka to present, the lithology is characterized by alternating clay and gypsum units. Gypsum deposition correlates with Heinrich Events (i.e. dry climate), whereas clay units coincide with more humid interstadials.     

New data on stratigraphy (palynomorphs,ostracods, paleomagnetism) of Cenozoic continental deposits of the Ishim plain,Western Siberia

New micropaleontological and paleomagnetic data were obtained by studying core samples of Cenozoic continental deposits from two boreholes drilled in the south of Tyumen oblast (Western Siberia). Palynological assemblages in deposits of the Tavda (upper part), Novomikhailovka, Turtas, Abrosimovka, Tobolsk, Smirnovka, and Suzgun formations were described. Deposits of these formations are enriched in spore-pollen assemblages, which can be correlated with assemblages of regional palynozones of the West Siberian Plain. Ostracods were described in Quaternary deposits. On the basis of biostratigraphic and paleomagnetic data, the Late Eocene (Priabonian)–Holocene age of deposits was substantiated. For the first time, beds with dinocysts of genus Pseudokomewuia were identified in deposits of the Turtas Formation (Upper Oligocene) of the Ishim lithofacial area. In total, nine regional magnetozones were distinguished in the paleomagnetic section. On the basis of palynological and paleomagnetic data, sections of two boreholes were correlated, and hiatuses in sedimentation were revealed. A large hiatus is at the Eocene-Oligocene boundary (Western Siberia): the Lower Oligocene Atlym Horizon and Miocene–Pliocene and Eopleistocene sediments are missing. The Oligocene interval of the section is represented in a reduced volume.     

Late Pleistocene stratigraphy, sedimentology and paleoenvironmental evolution of the Tarija-Padcaya basin (Bolivian Andes)

We illustrate the results of geomorphological, stratigraphical and sedimentological analysis of the Tarija-Padcaya basin, a wide depression in the eastern side of the Bolivian Cordillera. The basin is well known for the rich mammal fauna discovered since the beginning of the 19th century. The sedimentary infilling belongs to the Tolomosa Formation, corresponding to a major synthem subdivided into three main sub-synthems, mostly made of fluvial and alluvial fan sediments locally weathered by paleoalfisols (Ancon Grande sub-synthem), glacial and fluvio-glacial sediments (Puente Phayo sub-synthem) and finally alluvial fan and alluvial plain sediments (San Jacinto sub-synthem). Radiocarbon dating provides a chronology for the last sub-synthem and testifies that the sequence encompasses the Last Interglacial-Glacial cycle and constitutes a good proxy record for Late Pleistocene climatic changes. The occurrence of glacial deposits in the deeper part of the sedimentary filling suggests a major ice advance during MIS 4 and, together with glacial geomorphological evidence, points to further glacial erosion during the Last Glacial Maximum (LGM). The importance of glacial deposition and erosion opens the question of correlation with the events reported in the nearby Altiplano, where glacial deposits have been recognized only along the slopes of the higher volcanoes. In the Altiplano the LGM has been claimed to be characterized by an absence of deposition or deep erosion, due to extreme dryness, but the Tarija record suggests an erosional event of a scale that would imply the occurrence of a large ice cap.     

Magnetic polarity stratigraphy of the middle Pleistocene (Ensenadan) Tarija Formation of southern Bolivia

The Tarija Formation of southern Bolivia, which is well known for its classic vertebrate faunas, is of prime importance in understanding of the chronology of the Ensenadan Land Mammal Age. This formation consists of well-exposed and relatively fossiliferous sections of clays, clayey silts, sands, gravels, and tuffs which were deposited in a predominately fluviatile regime in a Pleistocene structural basin. Four stratigraphic sections, each measuring 110 m or less, were studied to establish a magnetic polarity stratigraphy. Paleomagnetic samples were collected from the finer-grained sediments at 100 sites spaced at stratigraphic intervals of 5 m or less. All paleomagnetic specimens were demagnetized in alternating fields of least 250 oersteds (oe). Some specimens were also thermally demagnetized at 200°C or more. Of the 100 sites, 77 were ultimately used to determine the magnetic polarity zonation. Based on the four sections sampled, the Tarija Formation spans a time interval from about 1 my to about 0.7 my B.P. or perhaps younger. The lower half of the composite section is of reversed polarity punctuated by a short normal event. This sequence probably represents the late Matuyama chron with the Jaramillo subchron. The upper part of the section is of normal polarity and represents early Brunhes time. A tuffaceous unit 43 m above the Brunhes-Matuyama boundary yielded a fission track (zircon) age of 0.7 ± 0.2 by B.P. These data indicate that the classic Tarija fauna is middle Pleistocene Ensendan in age.     

Integrated stratigraphy and palaeoenvironment of the P/E boundary interval, Rakhi Nala section, Indus Basin (Pakistan)

Marine sedimentary section across the Paleocene/Eocene (P/E) boundary interval is preserved in the Dungan Formation (Lower Indus Basin), Pakistan. Four dinoflagellate zones in the P/E interval of the Rakhi Nala section (Lower Indus Basin) are identified and correlated. The quantitative analysis of the dinoflagellate cyst assemblages together with geochemical data are used to reconstruct the palaeoenvironment across the P/E interval. The dinocyst assemblages allow the local correlation of the Dungan Formation (part) of the Sulaiman Range with the Patala Formation (part) of the Upper Indus Basin and global correlation of the Zone Pak-DV with the Apectodinium acme Zone of the Northern and Southern hemispheres. The onset of the carbon isotopic excursion (CIE) associated with Paleocene Eocene Thermal Maximum (PETM) is used globally to identify the P/E boundary. The CIE for the total organic carbon (fine fraction) δ¹³C_FF is of a magnitude of ?1.7‰ is recorded for the first time in the Indus Basin. The Apectodinium acme precedes and straddles the onset of the CIE in the Indus Basin. This Apectodinium acme is also accompanied by a planktonic and benthonic foraminifera “barren zone.” The CIE in the Indus Basin, coupled with the changes in the dinocyst distribution and the benthonic and planktonic foraminifera assemblages, provides evidence of the changes associated with the PETM in this little-known part of the world. The benthonic foraminiferal assemblage indicates bathyal environment of deposition at the time of P/E boundary interval; the presence of dominantly open marine dinoflagellates and high planktonic foraminiferal ratio suggest that the water column at this site was well connected with the rest of the Tethys.     

Genetic sequence stratigraphy of cool water slope carbonates (Pleistocene Eucla Shelf, southern Australia)

The south Australian Eucla Shelf belongs to the world's largest cool-water carbonate sedimentary system. During the Pleistocene, it exported large amounts of sediment to the shelf edge and upper slope resulting in an expanded sedimentary wedge. Wedge-internal clinoforming seismic reflectors suggest a stacking of the deposits into genetic sequences. High-resolution stable oxygen and carbon isotope, point counting, grain size, and carbonate mineralogical XRD analyses were carried out to characterize these genetic sequences along a dip-parallel transect of three ODP Leg 182 drill holes located between the shelf edge and upper slope. Oxygen and carbon isotope fluctuations show that the genetic sequences formed as a response to sea level fluctuations. Within the genetic sequences, facies differentiation and sediment volume partitioning occur along the transect. Lowstand deposits are fine grained and contain more sponge spicules and micrite. Highstand deposits are coarse grained with tunicate spicules, brown bioclasts, as well as bryozoan and corallinacean debris. Boundaries separating highstand and lowstand deposits are triggered by sea level fall, and are expressed as abrupt grain size changes or as turning points in grain-size trends. Analyzed components vary in abundance along the transect. Genetic sequences show dip-parallel variations in thickness combined with changing relative proportions of lowstand versus highstand deposits.     

Sequence stratigraphy in the upper cretaceous of the Basco-Cantabrian Basin (northern Spain)

The sedimentary cycles of the Cenomanian to Maastrichtian were investigated in the Basco-Cantabrian Basin (BCB) in northern Spain (Provinces of Alava, Vizcaya and Burgos). The depositional area was a distally steepened carbonate ramp which extended from Catalonia northwestwards to the Basque country. The investigated sediments range from calciturbidites and pelagic marls to marl-limestone alternations deposited on a distal carbonate ramp. Shallow marine limestones, marls and intertidal clastics and carbonates were deposited on the proximal part of the carbonate ramp. The establishment of a regional sequence analysis is based on the investigation of seismic profiles, well logs and outcrop sections. Examples of outcrop sections are interpreted in terms of sequence stratigraphy (unconformities of third- and second-order cycles, depositional geometries, systems tracts). The sequence stratigraphic interpretation of outcrop sections is based on facies analysis, interpretation of observed depositional geometries and correlation of unconformities and marine flooding surfaces through the basin. A biostratigraphic framework is established based on ammonites, inoceramids, planktonic and benthic foraminifera. As a result, a regional sequence stratigraphic cycle chart is presented and compared with published global cycle charts. The correlation of the regional cycle chart with published cycle charts is good. In the Cenomanian and Turonian, several sequence boundaries in the BCB are shifted by up to one biozone compared with the global chart. Some type 1 boundaries of the standard chart are only type 2 in the BCB. Important type 1 boundaries in the BCB are: top Geslinianum Zone with a 100 m lowstand wedge at the basis of the sequence (sequence boundary 92.2) ; base Petrocoriense Zone with a 250 m shallowing-upwards lowstand wedge at the basis (sequence boundary 89.2); and within the Syrtale Zone (sequence boundary 85.0).The Campanian-Maastrichtian sequence record is strongly disturbed by local compressive tectonics. Several sequences are recognizable and can be correlated with the global cycle chart. Correlation is hampered by the low biostratigraphic resolution in the western basin part. Subsidence analysis of several sections of the Upper Cretaceous of the BCB and its interpretation in the regional tectonic context leads to a discussion of the causes of the observed cyclicity. A regional eustatic curve is presented for the Upper Cretaceous of the BCB. Stage and substage names were used according Code-Committee (1977). Correspondence to: K.-U. Gräfe     

Tectonic analysis of an oceanic transform fault zone based on fault-slip data and earthquake focal mechanisms: the Húsavík–Flatey Fault zone, Iceland

The Húsavík–Flatey Fault (HFF) is an oblique dextral transform fault, part of the Tjörnes Fracture Zone (TFZ), that connects the North Volcanic Zone of Iceland and the Kolbeinsey Ridge. We carry out stress inversion to reconstruct the paleostress fields and present-day stress fields along the Húsavík–Flatey Fault, analysing 2700 brittle tectonic data measured on the field and about 700 earthquake focal mechanisms calculated by the Icelandic Meteorological Office. This allows us to discuss the Latest Cenozoic finite deformations (from the tectonic data) as well as the present-day deformations (from the earthquake mechanisms). In both these cases, different tectonic groups are reconstructed and each of them includes several distinct stress states characterised by normal or strike-slip faulting. The stress states of a same tectonic group are related through stress permutations (σ₁−σ₂ and σ₂−σ₃ permutations as well as σ₁−σ₃ reversals). They do not reflect separate tectonic episodes. The tectonic groups derived from the geological data and the earthquake data have striking similarity and are considered to be related. The obliquity of the Húsavík–Flatey Fault implies geometric accommodation in the transform zone, resulting mainly from a dextral transtension along an ENE–WSW trend. This overall mechanism is subject to slip partitioning into two stress states: a Húsavík–Flatey Fault-perpendicular, NE–SW trending extension and a Húsavík–Flatey Fault-parallel, NW–SE trending extension. These three regimes occur in various local tectonic successions and not as a regional definite succession of tectonic events. The largest magnitude earthquakes reveal a regional stress field tightly related to the transform motion, whereas the lowest magnitude earthquakes depend on the local stress fields. The field data also reveal an early extension trending similar to the spreading vector. The focal mechanism data do not reflect this extension, which occurred earlier in the evolution of the HFF and is interpreted as a stage of structural development dominated by the rifting process.     

Climate control on stacked paleosols in the Pleistocene of the Po Basin (northern Italy)

Paleosols are recurrent features in alluvial successions and provide information about past sedimentary dynamics and climate change. Through sedimentological analysis on six sediment cores, the mud-dominated succession beneath the medieval ‘Two Towers’ of Bologna was investigated down to 100 m depth. A succession of weakly developed paleosols (Inceptisols) was identified. Four paleosols (P1, P2, P3 and PH) were radiocarbon-dated to 40–10 cal ka bp . Organic matter and CaCO₃ determinations indicate low groundwater levels during soil development, which spanned periods < 5 ka. The development and burial of soils, which occurred synchronously in the Bologna region and in other sectors of the Po Plain, are interpreted to reflect climatic and eustatic variations. Climatic oscillations, at the scale of the Bond cycles, controlled soil development and burial during Marine Isotope Stage (MIS) 3 (P1 and P2). Rapid sea-level oscillations probably induced soil development at the MIS 3/2 transition (P3) and favored burial of PH after 10 ka bp . Weakly developed paleosols in alluvial successions can provide clues to millennial-scale climatic and environmental variations. In particular, the paleosol-bearing succession of the Po Plain represents an unprecedent record of environmental changes across the Late Pleistocene (MIS 3 and 2) in the Mediterranean region.     

Early to middle Holocene palaeoenvironmental reconstruction of the Beagle Channel (southernmost Argentina) based on terrestrial and marine palynomorphs

This paper presents the main palaeoenvironmental results obtained from a site located in the western sector of the Beagle Channel, Isla Grande de Tierra del Fuego, southernmost Argentina. The palynological analysis performed on marine sediments from Aserradero‐Lapataia 2 (latitude 54°51′22.7″S, longitude 68°34′22.8″W) allowed us to document the evolution of palaeoenvironmental conditions during the beginning of the Holocene marine transgression into the Beagle Channel. The results indicate that, prior to the marine incursion, the coastal areas were characterized by the presence of open‐grown shrubs and herbs along with woodland palaeocommunities. The aquatic environments were dominated exclusively by freshwater taxa. Around 8300 cal. a BP, seawater gradually flooded the channel while cold and high effective moisture conditions favoured the development of an arboreal vegetation with dominance of Nothofagus forest and scarcity of shrub and herbaceous communities. Gradually increasing salinities allowed the development of freshwater/marine transitional environments indicated by aquatic palynomorphs able to tolerate stressed conditions under fluctuating salinities. The increasing dinocyst diversification with dominance of heterotrophic taxa corroborates the establishment of a fully marine environment during the middle–late Holocene in the Beagle Channel.     

Problems of Oxfordian and Kimmeridgian stratigraphy in northern Central Siberia (Nordvik Peninsula section)

The Oxfordian–Lower Hauterivian section of the Nordvik Peninsula (northern Central Siberia) is a reference for developing zonal scales for various fossil groups and improving the Boreal zonal standard. In the middle 1950s–late 1980s, it was studied extensively by geologists, stratigraphers, lithologists, and experts on various fossil groups. These studies yielded rich fossil and microfossil collections and a set of parallel zonal scales for various faunal groups. Recently, a new detailed ammonite zonation of the Oxfordian and Kimmeridgian units of this section has been proposed. These results contradict the previous biostratigraphic data on ammonites, foraminifers, and palynomorphs. In the present paper, all the biostratigraphic data on the Oxfordian and Kimmeridgian units of the Nordvik Peninsula (Cape Urdyuk-Khaya) and northern Central Siberia undergo a comprehensive analysis and comparison with those on the Boreal Realm. The ammonite-constrained stratigraphic position of the lower Upper Jurassic in the Cape Urdyuk-Khaya section is interpreted as Upper Oxfordian or Middle Oxfordian. In our view, this difference in the understanding is due to the misidentification of some Oxfordian ammonite forms. The zones based on other fossil groups (foraminifers, dinocysts) which were distinguished in the Upper Oxfordian and Kimmeridgian sections of the Nordvik Peninsula are well traceable circumarctically. Their stratigraphic position in various regions of the Northern Hemisphere is constrained by ammonites and bivalves. However, if we use the last alternative ammonite zonation of this section part, hardly explicable contradictions will appear in interregional foraminiferal and dinocyst correlations.