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1.
Holocene fire disturbance and vegetation history were reconstructed using macroscopic charcoal and pollen accumulation rates from two lake sediment records (Holtjärnen and Klotjärnen) collected in the boreal forest of central Sweden. The records were used to examine the potential drivers associated with changes in fire regime. Climate, vegetation and human activity were all identified as factors variously influencing the fire regime. In the early Holocene, near bicentennial fire return intervals were regionally widespread, suggesting that fire disturbance was largely regulated by climate at that time. In the mid‐ and late Holocene, vegetation exerted an important control on the fire regime. During the mid‐Holocene, the expansion of thermophilous broadleaf vegetation offset the influence of warmer climate by altering the local microclimate and by changing the structure and flammability of the available fuels. During the transition to the late Holocene, thermophilous vegetation decreased in abundance and Pinus increased, resulting in a more flammable forest even though the climate was cooling and moistening. Fire disturbance correspondingly increased. The modern boreal forest was established in the late Holocene as Picea expanded regionally as the climate cooled, moistened, and became increasingly continental. Although no change in the frequency of fire was apparent at this time, increased stand densities likely facilitated greater fuel consumption in subsequent fires. Within the last millennium, human action markedly modified the forested landscape, altering the fire regime.  相似文献   

2.
《Quaternary Science Reviews》2007,26(17-18):2167-2184
The northwestern Great Basin lies in the transition zone between the mesic Pacific Northwest and xeric intermountain West. The paleoenvironmental history based on pollen, macroscopic charcoal, and plant macrofossils from three sites in the northwestern Great Basin was examined to understand the relationships among the modern vegetation, fire disturbance and climate. The vegetation history suggests that steppe and open forest communities were present at high elevations from ca 11,000 to 7000 cal yr BP, and were replaced by forests composed of white fir, western white pine, and whitebark pine in the late Holocene. Over the last 11,000 years, fires were more frequent in mid-elevation forests (10–25 fire episodes/1000 years) and rare in high-elevation forests (2–5 fire episodes/1000 years). Applying modern pollen–climate relationships to the fossil pollen spectra provided a means to interpret past climate changes in this region. In the past 9000 years summer temperatures decreased from 1 to 4 °C, and annual precipitation has increased 7–15%. These results indicate that the millennial-scale climate forcing driving vegetation changes can be quantified within the intermountain West in general and northwestern Great Basin in particular. In addition, fire can be considered an important component of these ecosystems, but it does not appear to be a forcing mechanism for vegetation change at the resolution of these records.  相似文献   

3.
The objective of this study was to investigate the possible links between regional climate, fire and vegetation at the small spatial scale during the early and mid Holocene in southern Sweden using pollen, plant macrofossil and charcoal records from a small bog. The fire history was compared with climate reconstructions inferred from various proxy records in the study region. High fire activity is related to dry and warm climate around 8550, 7600, 5500–5100 and 4500 cal. a BC. Low fire activity ca. 6500–6000 and 4750 BC may correspond to the widespread ‘8.2 k event’ (ca. 6200 BC) recorded across the North Atlantic region, and a later, brief period of increased precipitation, respectively. The decrease in broadleaved trees culminating ca. 6500–6000 BC correlates with the ‘8.2 k event’. A long mid Holocene period with low fire activity (ca. 4350–1000 BC) agrees with the pattern emerging for Europe from the global charcoal database, and may correspond to generally wetter and cooler conditions. High fire activity ca. 8550 BC probably triggered the local establishment of Corylus. Warmer and drier conditions (and high fire activity) ca. 7600 BC might have favoured the establishment of Alnus, Quercus and Tilia. The fire‐adapted Pinus maintained important populations throughout the early and mid Holocene. Copyright © 2008 John Wiley & Sons, Ltd.  相似文献   

4.
Research on abrupt paleoclimatic and paleoenvironmental change provides a scientific basis for evaluating future climate. Because of spatial variability in monsoonal rainfall, our knowledge about climate change during the mid-to lateHolocene in southern China is still limited. We present a multi-proxy record of paleoclimatic change in a crater lake, Lake Shuangchi. Based on the age-depth model from 210 Pb, 137 Cs and AMS14 C data, high-resolution mid-to late-Holocene climatic and environmental r...  相似文献   

5.
Sillasoo, Ü., Mauquoy, D., Blundell, A., Charman, D., Blaauw, M., Daniell, J. R. G., Toms, P., Newberry, J., Chambers, F M. & Karofeld, E. 2007 (January): Peat multi‐proxy data from Männikjärve bog as indicators of late Holocene climate changes in Estonia. Boreas, Vol. 36, pp. 20–37. Oslo. ISSN 0300–9483. As part of a wider project on European climate change over the past 4500 years, a 4.5‐m peat core was taken from a lawn microform on Männikjärve bog, Estonia. Several methods were used to yield proxy‐climate data: (i) a quadrat and leaf‐count method for plant macrofossil data, (ii) testate amoebae analysis, and (iii) colorimetric determination of peat humification. These data are provided with an exceptionally high resolution and precise chronology. Changes in bog surface wetness were inferred using Detrended Correspondence Analysis (DCA) and zonation of macrofossil data, particularly concerning the occurrence of Sphagnum balticum, and a transfer function for water‐table depth for testate amoebae data. Based on the results, periods of high bog surface wetness appear to have occurred at c. 3100,3010–2990,2300, 1750–1610, 1510, 1410, 1110, 540 and 310 cal. yr BP, during four longer periods between c. 3170 and 2850 cal. yr BP, 2450 and 2000 cal. yr BP, 1770 and 1530 cal. yr BP and in the period from 880 cal. yr BP until the present. In the period between 1770 and 1530 cal. yr BP, the extension or initiation of a hollow microtope occurred, which corresponds with other research results from Mannikjarve bog. This and other changes towards increasing bog surface wetness may be the responses to colder temperatures and the predominance of a more continental climate in the region, which favoured the development of bog micro‐depressions and a complex bog microtopography. Located in the border zone of oceanic and continental climatic sectors, in an area almost without land uplift, this study site may provide valuable information about changes in palaeohydrological and palaeoclimatological conditions in the northern parts of the eastern Baltic Sea region.  相似文献   

6.
Located on a mountain pass in the west-central Pyrenees, the Col d'Ech peat bog provides a Holocene fire and vegetation record based upon nine 14C (AMS) dates. We aim to compare climate-driven versus human-driven fire regimes in terms of frequency, fire episodes distribution, and impact on vegetation. Our results show the mid-Holocene (8500–5500 cal yr BP) to be characterized by high fire frequency linked with drier and warmer conditions. However, fire occurrences appear to have been rather stochastic as underlined by a scattered chronological distribution. Wetter and colder conditions at the mid-to-late Holocene transition (4000–3000 cal yr BP) led to a decrease in fire frequency, probably driven by both climate and a subsequent reduction in human land use. On the contrary, from 3000 cal yr BP, fire frequency seems to be driven by agro-pastoral activities with a very regular distribution of events. During this period fire was used as a prominent agent of landscape management.  相似文献   

7.
Climate change is allowing fire to expand into previously unburnt ecosystems and regions. While management policies such as fire suppression have significantly altered their frequency and intensity. To prevent future biodiversity/ecosystem services loss, and the large financial burden of wildfires, management plans will be required to adapt to future climate and land use changes. Long-term ecological data offer a unique perspective to assess fire variability under different climate and land-use conditions. In this study, we focus on Killarney National Park, Ireland. An area which today is under threat from an increase in fire activity. Comparing palaeoecological and archaeological records, we reconstruct the past fire dynamic and its impact on the landscape, and evaluate the role of climate vs humans in influencing the natural fire regime over the millennial time-scale. Our results indicate that fire has been present in the landscape since the beginning of the Holocene, with fire in the early Holocene being largely controlled by climate and microsite conditions, and in the late Holocene being increasingly influenced by human activity. The knowledge of past fire regimes can help inform future management in order to protect the semi-natural native woodland. The park's present landscape mosaic, could be preserved by limiting forest encroachment through moderate grazing and burning, while also protecting any fragmented forest from excessive grazing and large/intense fires, via traditional fire management strategies such as fuel load management. However, a fire management strategy should only be implemented following careful consideration of all ecosystem factors and controls.  相似文献   

8.
Two new peat‐based climate records from Ireland covering the late Holocene are presented. The sequences are dated by a strong chronological framework formed by AMS radiocarbon dates and SCPs. Three proxy indicators (testate amoebae, macrofossils and humification) have been determined allowing the limitations and strengths of each to be identified and utilised to provide a bog surface wetness (BSW) record for both sites. Age–depth models take into account the potential for accumulation rates to vary with bog vegetation. The records from each site have been used to derive a combined BSW record that displays changes to a wetter/cooler climate from ca. AD 30 (1920 BP), ca. AD 310 (1640 BP), ca. AD 805 (1145 BP), ca. AD 1040 (910 BP) and ca. AD 1300 (650 BP). Changes follow closely those identified in a northern Britain composite BSW record and largely correspond with lake‐level data in central France suggesting the main changes in water balance were coherent over a large region. Correspondence with increases in IRD and slower Iceland‐Scotland Overflow Water (ISOW) suggests that these changes were related to oceanic forcing influencing the track of dominant westerly air flow over Ireland. Copyright © 2007 John Wiley & Sons, Ltd.  相似文献   

9.
A 13,100-year-long high-resolution pollen and charcoal record from Foy Lake in western Montana is compared with a network of vegetation and fire-history records from the Northern Rocky Mountains. New and previously published results were stratified by elevation into upper and lower and tree line to explore the role of Holocene climate variability on vegetation dynamics and fire regimes. During the cooler and drier Lateglacial period, ca 13,000 cal yr BP, sparsely vegetated Picea parkland occupied Foy Lake as well as other low- and high-elevations with a low incidence of fire. During the warmer early Holocene, from ca 11,000–7500 cal yr BP, low-elevation records, including Foy, indicate significant restructuring of regional vegetation as Lateglacial Picea parkland gave way to a mixed forest of Pinus-Pseudotsuga-Larix. In contrast, upper tree line sites (ca >2000 m) supported Pinus albicaulis and/or P. monticola-Abies-Picea forests in the Lateglacial and early Holocene. Regionally, biomass burning gradually increased from the Lateglacial times through the middle Holocene. However, upper tree line fire-history records suggest several climate-driven decreases in biomass burning centered at 11,500, 8500, 4000, 1600 and 500 cal yr BP. In contrast, lower tree line records generally experienced a gradual increase in biomass burning from the Lateglacial to ca 8000 cal yr BP, then reduced fire activity until a late Holocene maximum at 1800 cal yr BP, as structurally complex mesophytic forests at Foy Lake and other sites supported mixed-severity fire regimes. During the last two millennia, fire activity decreased at low elevations as modern forests developed and the climate became cooler and wetter than before. Embedded within these long-term trends are high amplitude variations in both vegetation dynamics and biomass burning. High-elevation paleoecological reconstructions tend to be more responsive to long-term changes in climate forcing related to growing-season temperature. Low-elevation records in the NRM have responded more abruptly to changes in effective precipitation during the late Holocene. Prolonged droughts, including those between 1200 and 800 cal yr BP, and climatic cooling during the last few centuries continues to influence vegetation and fire regimes at low elevation while increasing temperature has increased biomass burning in high elevations.  相似文献   

10.
A high‐resolution record, covering 9.3–0.2 ka BP, from the sub‐arctic Stjernsund (70°N) was studied for benthic foraminiferal faunas and stable isotopes, revealing three informally named main phases during the Holocene. The Early‐ to Mid‐Holocene (9.3–5.0 ka BP) was characterized by the strong influence of the North Atlantic Current (NAC), which prevented the reflection of the Holocene Climatic Optimum (HCO) in the bottom‐water temperature. During the Mid‐Holocene Transition (5.0–2.5 ka BP), a turnover of benthic foraminiferal faunas occurred, Atlantic Water species decreased while Arctic‐Polar species increased, and the oxygen isotope record showed larger fluctuations. Those variations correspond to a period of global climate change, to spatially more heterogeneous benthic foraminiferal faunas in the Nordic Seas region, and to regionally diverging terrestrial temperatures. The Cool Late Holocene (2.5–0.2 ka BP) was characterized by increased abundances of Arctic‐Polar species and a steady cooling trend reflected in the oxygen isotopes. In this period, our record differs considerably from those on the SW Barents Sea shelf and locations farther south. Therefore, we argue that regional atmospheric cooling triggered the late Holocene cooling trend. Several cold episodes centred at 8.3, 7.8, 6.5, 4.9, 3.9 and 3.3 ka BP were identified from the benthic foraminiferal faunas and the δ18O record, which correlated with marine and atmospherically driven proxy records. This suggests that short‐term cold events may result from reduced heat advection via the NAC or from colder air temperatures.  相似文献   

11.
The paucity of low- and middle-elevation paleoecologic records in the Northern Rocky Mountains limits our ability to assess current environmental change in light of past conditions. A 10,500-yr-long vegetation, fire and climate history from Lower Decker Lake in the Sawtooth Range provides information from a new region. Initial forests dominated by pine and Douglas-fir were replaced by open Douglas-fir forest at 8420 cal yr BP, marking the onset of warmer conditions than present. Presence of closed Douglas-fir forest between 6000 and 2650 cal yr BP suggests heightened summer drought in the middle Holocene. Closed lodgepole pine forest developed at 2650 cal yr BP and fires became more frequent after 1450 cal yr BP. This shift from Douglas-fir to lodgepole pine forest was probably facilitated by a combination of cooler summers, cold winters, and more severe fires than before. Five drought episodes, including those at 8200 cal yr BP and during the Medieval Climate Anomaly, were registered by brief intervals of lodgepole pine decline, an increase in fire activity, and mistletoe infestation. The importance of a Holocene perspective when assessing the historical range of variability is illustrated by the striking difference between the modern forest and that which existed 3000 yr ago.  相似文献   

12.
In order to study the stability and dynamics of mountain rainforest and paramo ecosystems, including the biodiversity of these ecosystems, the Holocene and late Pleistocene climate and fire variability, and human impact in the southeastern Ecuadorian Andes, we present a high‐resolution pollen record from El Tiro Pass (2810 m elevation), Podocarpus National Park. Palaeoenvironmental changes, investigated by pollen, spores and charcoal analysis, inferred from a 127 cm long core spanning the last ca. 21 000 cal. yr BP, indicate that grass‐paramo was the main vegetation type at the El Tiro Pass during the late Pleistocene period. The grass‐paramo was rich in Poaceae, Plantago rigida and Plantago australis, reflecting cold and moist climatic conditions. During the early Holocene, from 11 200 to 8900 cal. yr BP, subparamo and upper mountain rainforest vegetation expanded slightly, indicating a slow warming of climatic conditions during this period. From 8900 to 3300 cal. yr BP an upper mountain rainforest developed at the study site, indicated by an increase in Hedyosmun, Podocarpaceae, Myrsine and Ilex. This suggests a warmer climate than the present day at this elevation. The modern subparamo vegetation became established since 3300 cal. yr BP at El Tiro Pass. Fires, probably anthropogenic origin, were very rare during the late Pleistocene but became frequent after 8000 cal. yr BP. Copyright © 2007 John Wiley & Sons, Ltd.  相似文献   

13.
Current high‐resolution palaeoenvironmental records reveal short‐term Holocene coolings. One of these major Holocene rapid climate changes occurred between 3.2 and 2.7 cal. ka BP. The sensitivity of river systems vis‐à‐vis slight and short‐term Holocene climatic variations is a subject of controversy in the scientific community. In this paper, we present a 4.0 to 1.4 cal ka BP palaeoflood record from the Lower Moulouya River (northeastern Morocco) to demonstrate the high sensitivity of semiarid rivers in the southwestern Mediterranean towards Holocene environmental changes. The Lower Moulouya flood deposits are characterised by thick, well‐stratified, predominantly clayey to silty overbank fine sediments. These cohesive sediments show evidence of excellent preservation conditions against fluvial erosion and contain a continuous record of mid to late Holocene flood sequences. The Moulouya palaeoflood record can be interpreted in the context of regional and global high‐resolution proxy data, revealing a strong coupling with Holocene rapid climate changes. The centennial‐scale Moulouya palaeohydrological history will be discussed with palaeoenvironmental data from the same record (palaeomagnetics, sedimentary charcoal record, anthracological analyses, snail analyses) to generate new ideas about the mid to late Holocene hydrological cycle in the southwestern Mediterranean. The deduced features of pronounced Lower Moulouya flooding and the decreased fire recurrence during Holocene cooling remain somewhat inconsistent with the interpretation of other palaeohydrological and paleaoecological records from the southwestern Mediterranean. However, enhanced Lower Moulouya flood frequencies between 3.2 and 2.7 cal. ka BP agree with increased floodplain aggradation in other major river systems of Mediterranean North Africa. Copyright © 2009 John Wiley & Sons, Ltd.  相似文献   

14.
Holocene fire-climate-vegetation linkages are mostly understood at individual sites by comparing charcoal and pollen records with other paleoenvironmental proxy and model simulations. This scale of reconstruction often obscures detection of large-scale patterns in past fire activity that are related to changes in regional climate and vegetation. A network of 31 charcoal records from southern South America was examined to assess fire history along a transect from subtropic to subantarctic biomes. The charcoal data indicate that fire activity was greater than present at ca. 12,000 cal yr BP and increased further and was widespread at 9500 cal yr BP. Fire activity decreased and became more spatially variable by 6000 cal yr BP, and this trend continued to present. Atmospheric circulation anomalies during recent high-fire years show a southward shift in westerlies, and paleoclimate model simulations and data syntheses suggest that such conditions may have prevailed for millennia in the early Holocene when the pole-to-equator temperature gradients were weaker and annual temperatures were higher than present, in response to orbital-time-scale insolation changes.  相似文献   

15.
High-resolution macroscopic charcoal analysis was used to reconstruct a 14,300-year-long fire history record from the lower Columbia River Valley in southwestern Washington, which was compared to a previous vegetation reconstruction for the site. In the late-glacial period (ca. 14,300-13,100 cal yr BP), Pinus/Picea-dominated parkland supported little to no fire activity. From the late-glacial to the early Holocene (ca. 13,100-10,800 cal yr BP), Pseudotsuga/Abies-dominated forest featured more frequent fire episodes that burned mostly woody vegetation. In the early to middle Holocene (ca. 10,800-5200 cal yr BP), Quercus-dominated savanna was associated with frequent fire episodes of low-to-moderate severity, with an increased herbaceous (i.e., grass) charcoal content. From the middle to late Holocene (ca. 5200 cal yr BP to present), forest dominated by Pseudotsuga, Thuja-type, and Tsuga heterophylla supported less frequent, but mostly large or high-severity fire episodes. Fire episodes were least frequent, but were largest or most severe, after ca. 2500 cal yr BP. The fire history at Battle Ground Lake was apparently driven by climate, directly through the length and severity of the fire season, and indirectly through climate-driven vegetation shifts, which affected available fuel biomass.  相似文献   

16.
Paleoenvironmental reconstructions are important for understanding the influence of long-term climate variability on ecosystems and landscape disturbance dynamics. In this paper we explore the linkages among past climate, vegetation, and fire regimes using a high-resolution pollen and charcoal reconstruction from Morris Pond located on the Markagunt Plateau in southwestern Utah, USA. A regime shift detection algorithm was applied to background charcoal accumulation to define where statistically significant shifts in fire regimes occurred. The early Holocene was characterized by greater amounts of summer precipitation and less winter precipitation than modern. Ample forest fuel and warm summer temperatures allowed for large fires to occur. The middle Holocene was a transitional period between vegetation conditions and fire disturbance. The late Holocene climate is characterized as cool and wet reflecting an increase in snow cover, which reduced opportunities for fire despite increased availability of fuels. Similarities between modern forest fuel availability and those of the early Holocene suggest that warmer summers projected for the 21st century may yield substantial increases in the recurrence and ecological impacts of fire when compared to the fire regime of the last millennium.  相似文献   

17.
High‐resolution pollen, macrofossil and charcoal data, combined with accelerator mass spectrometry 14C dating and multivariate analysis, were used to reconstruct Holocene vegetation and fire dynamics at Urio Quattrocchi, a small lake in the supra‐mediterranean belt in the Nebrodi Mountains of Sicily (Italy). The data suggest that after 10 000 cal a BP increasing moisture availability supported closed forests with deciduous (Quercus cerris, Fagus sylvatica and Fraxinus spp.) and evergreen (Quercus ilex) species. Species‐rich closed forest persisted until 6850 cal a BP, when Neolithic activities caused a forest decline and affected plant diversity. Secondary forest with abundant Ilex aquifolium recovered between 6650 and 6000 cal a BP, indicating moist conditions. From 5000 cal a BP, agriculture and pastoralism led to the currently fragmented landscape with sparse deciduous forests (Quercus cerris). The study suggests that evergreen broadleaved species were more important at elevations above 1000 m a.s.l. before ca. 5000 cal a BP than subsequently, which might reflect less human impact or warmer‐than‐today climatic conditions between 10 000 and 5000 cal a BP. Despite land use since Neolithic times, deciduous supra‐mediterranean forests were never completely displaced from the Nebrodi Mountains, because of favourable moist conditions that persisted throughout the Holocene. Reconstructed vegetation dynamics document the absence of any pronounced mid‐ or late‐Holocene ‘aridification’ trend at the site, an issue which is controversially debated in Italy and the Mediterranean region. Copyright © 2012 John Wiley & Sons, Ltd.  相似文献   

18.
Extensive fires pose catastrophic threats to both human and natural ecosystems. Understanding the history of fire, particularly Holocene palaeofire activity in densely populated areas, is essential for predicting future fire risks and developing effective fire management policies. The complexity of fire activity is influenced by various factors, including climate and anthropogenic activities. In this study, we analysed microcharcoal from the top 35.36 m of a well-dated sediment core HMD1401 in Ningshao Plain, eastern China. We combined our findings with phytolith and diatom evidence to obtain a comprehensive understanding of variations in Holocene fire activity and its controls. The results showed that there was higher fire activity during the early and late Holocene and less fire activity during the mid-Holocene. More frequent fire occurred from c. 10 000–7000 cal. a BP and was primarily caused by abundant biomass and high seasonal flammability due to increased annual temperature and precipitation and warm but dry winter climate. Fire occurrences between c. 7000–2000 cal. a BP remained at a low level, except for the periods c. 5900–5600 cal. a BP and c. 5300 cal. a BP, which may have been caused by extreme climate events. The impact of fire caused by human activity was significantly enhanced during the last two millennia.  相似文献   

19.
Beetle remains from a small bog in southern Sweden contribute information concerning the forest history of the study area. The study shows that beetles are valuable indicators of woodland structures such as openness, field vegetation, presence of dead wood and disturbance factors such as climate change, fire regimes, grazing and land use. The early Holocene, ca. 8600–6450 cal. BC, was characterised by open, pine‐dominated woodlands maintained by fire and grazing disturbances. The changes in the wetland fauna, between 8600 and 7500 cal. BC, correlate well with low lake levels in southern Sweden. During the mid Holocene, ca. 6450–2400 cal. BC, the woodlands were relatively dense, with few openings in the canopy. Around 4200 cal. BC, there was a shift to a dominance of deciduous trees. Fire and grazing pressures were particularly low. Numbers of aquatic and hygrophilic beetles indicate dry conditions between ca. 5000 and 3000 cal. BC. During the late Holocene, ca. 2400 cal. BC to present, the woodlands opened up mainly through increased land use. The main disturbance factors were fire and grazing. The beetles indicate the formation of heather‐dominated heathland around 800 cal. BC. Copyright © 2008 John Wiley & Sons, Ltd. This article was published online on 23 December 2008. An error was subsequently identified. This notice is included in the online and print versions to indicate that both have been corrected (5 August 2009).  相似文献   

20.
The mid to late‐Holocene climates of most of Scotland have been reconstructed from seven peat bogs located across north–south and east–west geographical and climatological gradients. The main techniques used for palaeoclimatic reconstruction were plant macrofossil, colorimetric humification, and testate amoebae analyses, which were supported by a radiocarbon‐based chronology, aided by markers such as tephra isochrons and recent rises in pine pollen and in spheroidal carbonaceous particles (SCPs). Field stratigraphy was undertaken at each site in order to show that the changes detected within the peat profiles were replicable. Proxy climate records were reconstructed using detrended correspondence analysis (DCA) of the plant macrofossil data and a mean water table depth transfer function on the testate amoebae data. These reconstructions, coupled with the humification data, were standardised for each site and used to produce a composite record of bog surface wetness (BSW) from each site. The results show coherent wet and dry phases over the last 5000 years and suggest regional differences in climate across Scotland, specifically between northern and southern Scotland. Distinct climatic cycles are identified, all of which record a millennial‐scale periodicity which can be correlated with previously identified marine and ice core Holocene cycles. The key role of the macrofossil remains of Sphagnum imbricatum, a taxon now extinct on many sites, is discussed in relation to the identified climatic shifts. Copyright © 2005 John Wiley & Sons, Ltd.  相似文献   

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