Flora development in Northeastern Asia and Northern Alaska during the Cretaceous-Paleogene transitional epoch

Study of floral succession from the Cretaceous-Paleogene boundary interval in Russian Far East (Zeya-Bureya depression), Northeastern Russia (Koryak Upland), and Northern Alaska (Sagavanirktok River basin) is crucial for better understanding palaeoclimatic and palaeogeographic factors, which controlled events in vegetation evolution at that time. The succession of fossil floras in the Zeya-Bureya depression includes plant assemblages of the Santonian, Campanian, early Danian, Danian, and Danian-Selandian age. The early Danian Boguchan Flora keeps continuity in composition and dominating taxa with the Campanian Late Kundur Flora. The Koryak Flora of the Amaam Lagoon area (Northeastern Russia) is dated as late Maastrichtian based on correlation of plant-bearing beds with marine biostratigraphy, whereas the Early and Late Sagwon floras of Northern Alaska are dated back to the Danian-Selandian and early Paleocene based on palynological and macrofloristic data. The Early Sagwon Flora is most close to the late Maastrichtian Koryak Flora of the Amaam Lagoon area in composition and main dominants, while the Late Sagwon Flora is comparable with the Danian or Danian-(?) Selandian flora from the Upper Tsagayan Subformation of the Amur area. In a florogenic aspect, trans-Beringian plant migrations from northeastern Asia and southern palaeolatitudes of the Far East, which became possible due to Paleocene climate warming in Arctic, have played an important role in forming of the Paleocene floras of Northern Alaska. Floras of the Far East and high latitudes of Asia and North America show no evidence of catastrophic event at the Cretaceous-Paleogene boundary. Their development was most probably controlled by climate changes, plant evolution and migration.     

The late Cretaceous Arman flora of Magadan oblast,Northeastern Russia

The Arman flora from the volcanogenic-sedimentary beds of the Arman Formation is systematically studied using materials from the Arman River basin and the Nelkandya-Khasyn interfluve (Magadan oblast, Northeastern Russia). Seventy-three species of fossil plants belonging to 49 genera are described. They consist of liverworts, horsetails, ferns, seed ferns, cycadaleans, bennettitaleans, ginkgoaleans, czekanowskialeans, conifers, gymnosperms of uncertain systematic affinity, and angiosperms. The Arman flora shows a unique combination, with relatively ancient Early Cretaceous ferns and gymnosperms occurring alongside younger Late Cretaceous plants, primarily angiosperms. The similarity of the Arman flora to the Penzhina and Kaivayam floras of northwestern Kamchatka and the Tylpegyrgynai flora of the Pekul’nei Ridge allows the Arman flora to be dated as Turonian and Coniacian (Late Cretaceous), which is corroborated by isotopic (U-Pb and ⁴⁰Ar/³⁹Ar) age determination for the plant-bearing layers.     

Radiolarian-Based zonal scheme of the cretaceous (Albian–Santonain) of the Tethyan regions of Eurasia

A scheme of radiolarian zonal subdivision is proposed for the upper Albian–Santonian of the Tethyan regions of Eurasia. The upper Albian contains one zone: Crolanium triangulare; the Cenomanian contains three zones: Patellula spica (lower Cenomanian), Pseudoaulophacus lenticulatus (middle Cenomanian), and Triactoma parva (upper Cenomanian); the Turonian contains four zones: Acanthocircus tympanum (lower Turonian (with no upper part)), Patellula selbukhraensis (upper part of the lower Turonian), Phaseliforma turovi (middle Turonian (with no upper part)), and Actinomma (?) belbekense (upper part of the middle Turonian–upper Turonian); the Coniacian contains two zones: Alievium praegallowayi (lower part of the Coniacian) and Cyprodictyomitra longa (upper part of the Coniacian); the Santonian contains three zones: Theocampe urna (lower Santonian), Crucella robusta (middle Santonian–lower part of the upper(?) Santonian), and Afens perapediensis (upper part of the upper Santonian). The biostratigraphic subdivisions are correlated with biostrata in the schemes proposed previously for the Tethys and Pacific. A new species Patellula selbukhraensis Bragina sp. nov. is described.     

Late cretaceous Arman’ flora of Magadan oblast: Paleoclimatic interpretation

The Arman’ flora belongs to the formation of the same name in the Arman’ River basin and Nel’khandzha-Khasyn interfluve. It includes about 80 fossil plant species. The similarity of this flora to the reliably dated fossil floras of northwestern Kamchatka and the Pekul’nei Range dates it to being Turonian and Coniacian in age, which is corroborated by isotopic (U-Pb and ⁴⁰Ar/³⁹Ar) dating of the plant-bearing layers. The CLAMP method is used to estimate quantitative climatic variables, according to which the Arman’ flora existed in a temperate humid climate with warm summers and mild winters. This climate is especially similar to the conditions reconstructed from fossil plants of the Turonian Penzhina flora and the Coniacian Kaivayam and Tylpegyrgynai floras of Northeast Asia.     

Arman’ Flora of the magadan region and development of floras in the North Pacific during the Albian-Paleocene

The Arman’ Flora from volcanogenic-sedimentary deposits in the Arman’ River basin and Naraula Formation in the Nel’kandzha-Khasyn interfluve includes 82 species of fossil plants comprising liverworts, horsetails, ferns, caytonealeans, cycadaleans, ginkgoaleans, czekanowskialeans, conifers, gymnosperms incertae sedis, and angiosperms. The Arman’ Flora appears to be of Turonian-Coniacian age, as it is close to the reliably dated Penzhina and Kaivayam floras from the Northwest Kamchatka and to Tyl’pegyrgynai Flora of the Pekul’nei Ridge. The dating is consistent with isotopic dates (⁴⁰Ar/³⁹Ar and U-Pb SHRIMP) characterizing the age of plant-bearing sequences. Based on the considered position of the Arman’ Flora in the scheme of Cretaceous florogenesis, a leading role in that florogenesis was played by the gradual invasion of floras by new, angiosperm dominated, plant communities. These communities initially populated unstable habitats in the coastal lowlands of Northeast Asia and Alaska, gradually invading with time the Asiatic intracontinental areas. The peculiar combination of Early and Late Cretaceous plants characteristic of the Arman’ Flora is strong evidence that Cenophytic plant communities dominated by angiosperms colonized areas still populated in the Late Cretaceous by Mesophytic communities. Absence of Mesophytic and Cenophytic taxa mixing in the Arman’ Flora burials suggests a replacement of plant communities as whole rather than of separate plants by more advanced taxa.     

Late Cretaceous Kholokhovchan Flora of Northeastern Asia: Composition,age and fossil plant descriptions

The Kholokhovchan Flora comes from tuffaceous – terrigenous deposits of the Vetvinskaya Member (Chalbugchan Group) in the Penzhina and Oklan rivers interfluve, Northeastern Russia. The depositional environment of the plant-bearing deposits is interpreted to have been a freshwater lake. The Kholokhovchan Flora hosts 42 fossil plant species belonging to Marchantiopsida, Polypodiopsida, Ginkgoales, Leptostrobales, Bennettitales, Pinales and Magnoliopsida. It is characterised by diverse angiosperms, less diverse conifers and ferns, by the presence of relatively ancient Sphenobaiera, Phoenicopsis and Pterophyllum together with advanced Late Cretaceous Taxodium, Glyptostrobus and angiosperms, among which platanoids are quite diverse. The Kholokhovchan Flora is most similar to Penzhina and Kaivayam floras of the Anadyr-Koryak Subregion and Arman Flora of the Okhotsk-Chukotka volcanogenic belt (Northeastern Russia) and should be dated as Turonian–Coniacian. The Kholokhovchan Flora, that populated volcanic plateaus and intermontane valleys, are characterised by a mixture of ancient “Mesophytic” plants with typical Late Cretaceous “Cenophytic” taxa. This peculiar composition probably reflects a gradual penetration of new angiosperm-dominated plant assemblages into older floras: during the Late Cretaceous, “Cenophytic” assemblages migrated along river valleys and other disturbed habitats into the interior of Asia, eventually occupying volcanogenic uplands, and in places replacing the “Mesophytic” fern-gymnospermous communities that existed there. Two new angiosperm species, as well as four the most characteristic conifers of the Kholokhovchan Flora, are described: Cupressaceae gen. et sp. indet. cf. Widdringtonites sp., Taxodium cf. olrikii, Taxodium sp., Glyptostrobus sp., Ettingshausenia vetviensis sp. nov. and Parvileguminophyllum penzhinense sp. nov.     

Aptian to Coniacian (Early–Late Cretaceous) palynostratigraphy of the Gustav Group, James Ross Basin, Antarctica

The Gustav Group of the James Ross Basin, Antarctic Peninsula, forms part of a major Southern Hemisphere Cretaceous reference section. Palynological data, chiefly from dinoflagellate cysts, integrated with macrofaunal evidence and strontium isotope stratigraphy, indicate that the Gustav Group, which is approximately 2.6 km thick, is Aptian–Coniacian in age. Aptian–Coniacian palynofloras in the James Ross Basin closely resemble coeval associations from Australia and New Zealand, and Australian palynological zonation schemes are applicable to the Gustav Group. The lowermost units, the coeval Pedersen and Lagrelius Point formations, have both yielded early Aptian dinoflagellate cysts. Because the overlying Kotick Point Formation is of early to mid Albian age, the Aptian/Albian boundary is placed, questionably, at the Lagrelius Point Formation–Kotick Point Formation boundary on James Ross Island, and this transition may be unconformable. Although the Kotick Point Formation is largely early Albian on dinoflagellate cyst evidence, the uppermost part of the formation appears to be of mid Albian age. This differentiation of the early and mid Albian has refined the age of the formation, previously considered to be Aptian–Albian, based on macrofaunal evidence. The Whisky Bay Formation is of late Albian to latest Turonian age on dinoflagellate cyst evidence and this supports the macrofaunal ages. Late Albian palynofloras have been recorded from the Gin Cove, lower Tumbledown Cliffs, Bibby Point and the lower–middle Lewis Hill members. However, the Cenomanian age of the upper Tumbledown Cliffs and Rum Cove members, based on molluscan evidence, is not supported by the dinoflagellate cyst floras and further work is required on this succession. The uppermost part of the Whisky Bay Formation in north-west James Ross Island is of mid to late Turonian age and this is confirmed by strontium isotope stratigraphy. The uppermost unit, the Hidden Lake Formation, is Coniacian in age on both palaeontological and strontium isotope evidence. The uppermost part of the formation appears to be early Santonian based on dinoflagellate cysts, but strontium isotope stratigraphy constrains this as being no younger than late Coniacian. This refined palynostratigraphy greatly improves the potential of the James Ross Basin as a major Cretaceous Southern Hemisphere reference section.     

Comparative paleofloristics of the Albian-early Paleocene in the Anadyr-Koryak and North Alaska Subregions,Part 1: The Anadyr-Koryak Subregion

A high-resolution phytostratigraphic scheme based on plant macrofossils is suggested for the first time for the Albian and Upper Cretaceous of the Anadyr-Koryak Subregion of the North Pacific. Seven phytostratigraphic horizons of the subregion are distinguished based on successive stages in flora evolution substantiated by comprehensive data on described floral assemblages.     

Cretaceous stratigraphy of the Ionian Zone,Hellenides, western Greece

The Cretaceous sedimentary successions of the Ionian Zone, Hellenides, western Greece, are composed of pelagic limestones intercalated with cherty layers. The micritic and biomicritic beds with abundant chert nodules and cherty horizons, which were deposited during late Tithonian to early Santonian times, belong to the Vigla Limestone Formation, while the sediments deposited during the late Santonian to Maastrichtian, formed clastic limestone beds in which chert nodules also occur sparsely.In the Cretaceous beds calpionellids, planktonic and benthonic foraminifera characteristics of the Tethyan realm, and radiolaria have been recorded. The calpionellids, together with radiolaria, colonized the entire basin during the Berriasian to early Valanginian, the latter becoming dominant during the Hauterivian to early Albian as a result of anoxia. Planktonic foraminifera first appeared in the basin during the late Albian and persisted until the Maastrichtian. The numbers decreased, however, during the Cenomanian-early Turonian interval, when radiolaria increased owing to anoxic conditions, and during the Campanian-Maastrichtian interval because the basin became shallow. During this interval larger benthonic foraminifera colonized the basin. Zonal markers have been recognized in calpionellid and planktonic foraminiferal assemblages on the basis of which two calpionellid zones are distinguished, viz. the Calpionella alpina and Calpionellopsis Zones (Berriasian-early Valanginian) along with seven planktonic foraminiferal zones, viz. the Rotalipora ticinensis, Rotalipora appenninica (late Albian), Rotalipora brotzeni (early Cenomanian), Helvetoglobotruncana helvetica (early to middle Turonian), Marginotruncana sigali(late Turonian to early Coniacian), Dicarinella concavata (late Coniacian to early Santonian) and Dicarinella asymetrica (late early-late Santonian) Zones.The anoxic conditions that prevailed in the Ionian basin during the Barremian-early Albian, Cenomanian-early Turonian and Coniacian-Santonian intervals probably arose as a result of (a) the accumulation of large amounts of organic matter because the palaeotopography of the basin periodically hindered the circulation of water from the ocean and (b) the oxygen content of the intruding oceanic waters was low.     

Comparative paleofloristics of the Albian-Early Paleocene in the Anadyr-Koryak and North Alaska Subregions,part 2: The North Alaska Subregion

The Albian-Paleocene phytostratigraphic scheme suggested in this work for the North Alaska Subregion of the North Pacific differs from the previous ones and includes three phytostratigraphic horizons of subregional extent and four plant-bearing beds. The scheme is based on distinguished successive stages in flora evolution, being substantiated by comprehensive data on the described floral (plant macrofossils) assemblages. As is established, sedimentation environments gradually changed during the Albian-Paleogene from the west of the subregion to the east so that the marine sedimentation and accumulation of sediments in general terminated in eastern areas later than western ones.     

Comparative paleofloristics of the Albian-early Paleocene in the Anadyr-Koryak and North Alaska subregions,Part 3: Comparison of floras and floristic changes across the Cretaceous-Paleogene boundary

Floras characterizing comparable evolutionary stages in the Anadyr-Koryak and North Alaska subregions of the North Pacific show some essential distinctions despite their similarity in general. Factors responsible for appearance of their distinctive features were most likely the paleoclimatic difference between the subregions and the constrained or even interrupted trans-Beringia migration of plants in particular intervals of geological history. Floras of both subregions survived the Cretaceous-Paleogene boundary crisis without essential consequences for their evolution, and amplitude of floristic changes across the boundary was not greater than by evolutionary changes in the crisis-free Late Cretaceous. Evolution of the North Pacific floras near this boundary has been likely controlled by the long-term paleogeographic and paleoclimatic changes and by plants evolution and migration.     

Evolution of radiolarians in the late Albian–Campanian

The dynamics of radiolarian evolution in the late Albian–Campanian is analyzed, and several stages are recognized. The first stage (late Albian–middle Cenomanian), related to the MCE regional anoxic event, showed low evolutionary tempos and hence lacked structural change in radiolarian communities. The second stage (late Cenomanian–early Turonian), corresponding to ОАЕ 2, which was a global anoxic event, is characterized by a decrease in the number of genera, while many genera showed increased diversification of species composition. At this stage, a considerable number of genera became extinct and appeared, suggesting an increased rate of the radiolarian evolution. The third stage (middle Turonian–early Coniacian), including the beginning of ОАЕ 3, is characterized by a stabilized number of genera. The fourth stage (late Coniacian–Santonian) completely encompasses ОАЕ 3 except for its very beginning. At this stage, the radiolarian communities underwent a significant structural change, while their rate of evolution increased considerably. Nevertheless, during the ОАЕ 3 stage, a distinct trend toward a decrease in generic diversity continued from the late Cenomanian to the middle Turonian. The fifth stage (Campanian) is characterized by quite significant changes in the assemblage composition, while the trend toward a gradual decrease in the number of genera steadily continued. At this stage, which coincided with a considerable cooling, twice as many genera became extinct as during ОАЕ 2. The analysis of the dynamics of radiolarian evolution showed that the anoxic MCE, ОАЕ 2, and ОАЕ 3 events did not result in degradation of radiolarian assemblages. This suggests that this group has significant stratigraphic potential. In general, the evolution of radiolarians in the Late Cretaceous was gradual. By the end of the Campanian, nearly half of the generic diversity was composed of genera which appeared at the beginning of the Cretaceous and earlier.     

Evidence of palaeo-wildfire from the upper Lower Cretaceous (Serra do Tucano Formation,Aptian–Albian) of Roraima (North Brazil)

Wood fossil charcoal is identified from the upper Lower Cretaceous (Serra do Tucano Formation, Aptian–Albian) of Roraima (North Brazil). The presence of charcoal demonstrates the occurrence of Early Cretaceous palaeo-wildfires for the first time in this region and only the third time for the entirety of South America. A gymnospermous taxonomic affinity can be established for the charred woods and a relationship with conifers is likely, thus providing additional evidence for the taxonomic composition of Early Cretaceous floras in this region.     

Turonian cephalopods (ammonoids and a nautiloid) from the Wadi Daya Formation of the Talerzha Basin (South Riffian Ridges Domain,northern Morocco)

The Wadi Daya Formation, or the Calcaires crayeux of the older literature, attains a thickness of 10–40 m in the Talerhza Basin of the South Riffian Ridges. Previously, this unit was first dated as “Vraconian” (i.e., late upper Albian), but then reinterpreted as Cenomanian-Turonian and Cenomanian-Coniacian on the basis of foraminiferal and ostracod assemblages, respectively. Here, we record for the first time in the South Riffian Ridges, some typically Turonian ammonoids and a nautiloid species, namely Romaniceras (Yubariceras) cf. ornatissimum (Stoliczka), Spathites (Jeanrogericeras) cf. reveliereanus (Courtiller), Neoptychites cephalotus (Courtiller), Pachydesmoceras linderi (de Grossouvre), Lewesiceras peramplum (Mantell) and Angulithes galea (Fritsch, in Fritsch & Schlönbach). These species are herein described and illustrated. In view of these data, the underlying Marnes et marno-calcaires jaunes Formation, formerly dated as “Vraconian”, could in fact be of a middle to late Cenomanian date, in accordance with the age assignment based on planktonic foraminifera. Deposition of the overlying Marnes jaunes Formation, previously dated as Cenomanian-“Senonian”, probably started during the latest Turonian or earliest Coniacian.     

Nannostratigraphy and investigation of sedimentation conditions of the lower boundary and the upper boundary of the Aitamir Formation in the east and west Kopet Dagh,northeast of Iran

By attention to the stratigraphic value of calcareous nannoplanktons for the age determination of sedimentary beds, for the first time Late Cretaceous calcareous nannofossil taxa, their distributions and relative abundances were recorded from the lower and the upper boundary of Aitamir Formation located in northeast Iran. In the present study, biostratigraphy and paleoecological conditions were reconstructed. The Aitamir Formation comprises glauconitic sandstones and olive-green shales. In this work, samples were prepared with smear slides, and nannofossils of these boundaries are listed and figured. They were photographed under a light microscope. Based on nannoplanktons and as a result of biostratigraphic studies, the age of the lower boundary of the Aitamir Formation in the east Kopet Dagh is Early/Middle Turonian, the age of the lower boundary in the west Kopet Dagh is Late Turonian/Early Coniacian, the age of the upper boundary of the Aitamir Formation in the east Kopet Dagh is Late Santonian, and the age of the upper boundary of this Formation in the west Kopet Dagh is Early Campanian. Based on paleoecological interpretation, the Aitamir Formation was deposited in a shallow marine environment, at relatively low latitude. A deepening trend of the sedimentary basin is recognized passing from Aitamir Formation to the overlying Abderaz Formation while in the lower boundary from Sanganeh to Aitamir Formation depth decreased.     

Organic-walled dinoflagellate cyst records from a prospective Turonian – Coniacian (Upper Cretaceous) GSSP,Słupia Nadbrzeżna,Poland

A river section at Słupia Nadbrzeżna, central Poland, has been proposed as a candidate Turonian – Coniacian (Cretaceous) GSSP, in combination with the Salzgitter-Salder quarry section of Lower Saxony, Germany. Results of a high-resolution (25 cm) palynological study of the boundary interval in the Słupia Nadbrzeżna section are presented. Terrestrial palynomorphs are rare; marine organic-walled dinoflagellate cysts dominate the palynological assemblage. The dinoflagellate cyst assemblage has a low species richness (5–11 per sample; total of 18 species recorded) and diversity (Shannon index H = 0.8–1.4), dominated by four taxa: Circulodinium distinctum subsp. distinctum; Oligosphaeridium complex; Spiniferites ramosus subsp. ramosus; Surculosphaeridium longifurcatum. Declining proportions of O. complex and S. ramosus subsp. ramosus characterise the uppermost Turonian, with an increased dominance of S. longifurcatum in the lower Coniacian. The Turonian – Coniacian boundary interval includes an acme of C. distinctum subsp. distinctum in the upper Mytiloides scupini Zone, a dinoflagellate cyst abundance maximum in the Cremnoceramus walterdorfensis walterdorfensis Zone, and the highest occurrence of Senoniasphaera turonica in the basal Coniacian lower Cremnoceramus deformis erectus Zone. Most previously reported Turonian – Coniacian boundary dinoflagellate cyst marker species are absent; a shallow-water oligotrophic epicontinental depositional setting, remote from terrestrial influence, likely limited species diversity and excluded many taxa of biostratigraphic value.     

The Albian to Turonian carbon isotope record from the Shilaif Basin (United Arab Emirates) and its regional and intercontinental correlation

The sedimentary record of the Arabian Shelf offers a unique opportunity to study the Cretaceous (Albian–Turonian) greenhouse climate from a palaeoequatorial perspective. In particular, hemipelagic to pelagic carbonate successions from the extensive Shilaif intra‐shelf basin have the potential to produce an excellent record of carbon cycle perturbations during this interval. This study presents a 269 m thick chemostratigraphic (carbonate δ¹³C and δ¹⁸O) record from the Middle Albian to Early Turonian of central Abu Dhabi (United Arab Emirates), representing over 14 Myr of uninterrupted carbonate sedimentation. The Mauddud to Shilaif formations represent outer ramp to basinal intra‐shelf carbonates with variations from laminated organic‐rich to clean bioturbated intervals. Isotopic evidence of the latest Albian Anoxic Event (Oceanic Anoxic Event 1d), Middle Cenomanian Event I and the Cenomanian–Turonian Anoxic Event (Oceanic Anoxic Event 2) are confirmed and biostratigraphically calibrated by means of calcareous nannofossils. The carbon isotope record allows correlation with other regional records and well‐calibrated records across the Tethyan Ocean and represents a significant improvement of the chronostratigraphic framework of the United Arab Emirates (Shilaif) and Oman (Natih) intra‐shelf basins. The study further confirms that low carbon isotope values corresponding to the two source rock intervals in the Shilaif Formation clearly precede the isotopic expressions of Oceanic Anoxic Event 1d and Oceanic Anoxic Event 2.     

Sequence architecture and carbonate platform configuration (Late Cenomanian–Santonian), Sinai, Egypt

Abstract Relative sea‐level changes on the mixed carbonate–siliciclastic platform of Sinai are manifested in shifts of distinct facies belts (deep‐water facies, high‐energy subtidal, shallow subtidal, lagoon, shallow shoreface siliciclastics, supratidal) and are interpreted in terms of sequence stratigraphy. Eight sedimentary sequences are recognized for the Upper Cenomanian to Santonian. Their correlation along a north–south transect reveals distinct changes in lithofacies and progradation/retrogradation patterns within the individual systems tracts. The number and stratigraphy of the sequence boundaries of Sinai correlate well with those from adjacent areas. Patterns of increased subsidence are documented for the Central Sinai Basin since the Late Cenomanian by increased thickness of the stratal packages (post‐CeSin 7 HST, post‐TuSin 1 LST and HST, post‐TuSin 2 LST) and are balanced by varying accumulation rates. Based on new sedimentological and biostratigraphic data, large‐scale palaeogeographic maps and cross‐sections show the: (1) temporal and spatial evolution of the Central Sinai Basin, e.g. its latest Cenomanian initial formation, Lower Turonian deep‐water facies, Middle Turonian to Coniacian synsedimentary subsidence; (2) drowning of the Cenomanian platform coinciding with the latest Cenomanian to Early Turonian relative sea‐level rise; (3) re‐establishment of the platform in Middle–Late Turonian times; and (4) a Coniacian basin and swell morphology.     

Cenomanian–Turonian carbonate platform deposits in west central Jordan

We studied upper Albian to Turonian shallow-marine shelf deposits (Ajlun Group) of west central Jordan along a NNE-SSW running transect. The carbonate-dominated succession includes few siliciclastic intercalations, claystones and shales, and can be subdivided into five formations. The Naur, Fuheis and Hummar Formations of upper Albian to upper Cenomanian age represent shallow subtidal to supratidal platform environments. The uppermost Cenomanian to middle Turonian Shueib Formation includes deeper water deposits of the inner/mid-shelf and locally TOC-rich black shales. Shallow-marine platform environments once again dominate the Wadi As Sir Formation (middle-upper Turonian). A new multibiostratigraphic framework is based on ammonites (mainly of the middle Cenomanian rhotomagense Zone to the middle Turonian woollgari Zone) and calcareous nannofossils (biozones CC 9–CC 11), supplemented by benthic and planktonic foraminifers and ostracods. It forms the base of a sequence stratigraphic subdivision, containing eight sedimentary sequences (S1–S8), which are separated by four Cenomanian sequence boundaries (CeJo1–CeJo4) and three Turonian sequence boundaries (TuJo1–TuJo3). This scheme allows the correlation of the platform succession from distal to proximal shelf areas in contrast to previous correlations using lithologic units. Furthermore, comparisons between the platform successions and sequence patterns of west central Jordan and those from neighbouring areas allow to differentiate local, regional, and global controlling factors of platform development within the study area.     

Biogeography of the Saharan Gretaceous and Paleocene epicontinental transgressions

The first trans-Saharan epicontinental transgression began in the Late Cenomanian and peaked in the Early Turonian. The sea entered through rifts (Nigeria) and subsiding areas (north Africa) and, at maximum extent, spilled out onto stable craton. The central Sahara was land in late Early Turonian time; the sea seems to have remained in the central part of the Benue rift, to sally forth again during the minor Coniacian transgressive pulse. The Coniacian transgression went no further than eastern and central Niger Republic. For a short time during the Cenomanian-Turonian link-up between the Tethys and the South Atlantic, some organisms (mainly ammonites and pelecypods) passed across the Sahara. The Coniacian marine deposits of eastern Niger contain several elements typical of the Mungo River Formation of Cameroun, and coastal Nigeria. These faunistic indications imply that the topographical obstacle formed by the Zambuk ridge of northeastern Nigeria was swamped at maximum transgression. A third, very extensive epicontinental transgression began in latest Campanian time and reached its acme in the Early Maastrichtian. In the central Sahara, at least, there was a retreat of the sea in later Maastrichtian times. A final transgression peaked during the Paleocene, after which the sea withdrew permanently from the northwest African hinterland. The same marine ostracod associations occur in the Paleocene of coastal Nigeria, the Sokoto embayment, Mali and Libya, thus providing strong evidence of a marine connexion, albeit brief, between the South Atlantic and the Tethys, despite the lack of outcrop evidence in the crucial Niger valley region.