辽西首次发现保存毛发和软件的中生代哺乳动物化石

中国辽西地区素以产出丰富多彩的中生代热河生物群闻名于世。特别是近几年来,该区大量长羽毛的恐龙、原始鸟类、原始哺乳类和被子植物化石的发现更加引起了国际科学界的极大关注。1997年秋,作者在辽宁北票四合屯地区发现了一块完整的保存有毛发和软体印痕的对齿兽类哺乳动物化石。身体保存有手发和软体印痕的中生代哺乳动物化石在世界上也是罕见的。对对于研究对齿兽形态结构、生物习性、系统关系等方面均具有重要意义。     

A Possible Long-tailed Bird with a Pygostyle from the Late Mesozoic Yixian Formation, Western Liaoning, China

AbstractAn incomplete caudal vertebral series (IVPP Vl1309) from the Yixian Formation of late MesozoicJianshangou area of Beipiao, western Liaoning Province, may represent a new bird. The tail is composed of at least 12 free caudal vertebrae and the most distal 5 caudal vertebrae co-ossified into a pygostyle. The pygostyle is plate-like and slightly curved dorsally. The anterior free caudals are amphiplatyan. The anterior caudal surfaces of the last three free caudals are concave, but their posterior articular surfaces are convex. The pygostyle is regarded as the first appeared flight apparatus during the evolutionary process from Archaeopteryx to neornithes. The pygostyle appeared in most early fossil birds and almost all the modem birds. Although their morphologies are different, they are basically formed by at least four last caudal vertebrae. The specimen V11309 is regarded as a bird rather than a non-avian theropod dinosaur based on the following characters: short caudal vertebrae, numerous pits present on the surfaces of the centra and, a foramen present between the basal part of the fused neural spines, which is similar to that of Struthio camelus. The discoveries of pygostyles from the therizinosauroids and oviraptorosaurs may provide strong evidence for supporting the origin of birds from small theropod dinosaurs. The structure of the pygostyle in specimen Vl1309 is different from those of Beipiaosaurus (Therizinosauroid) and Nomingia (oviraptorosaur). The most parsimonious interpretation is that these pygostyle-like structures are independently acquired by Beipiaosaurus and Nomingia during their evolutionary process.     

A New Troodontid (Theropoda: Troodontidae) from the Lower Cretaceous Yixian Formation of Western Liaoning, China

A specimen collected from the Lower Cretaceous Yixian Formation of western Liaoning, China, represents a new genus and species of troodontid theropod. The new taxon is named and described on the basis of the holotype and the only known specimen, which comprises an articulated skeleton with the presacral vertebral, shoulder girdle and forelimbs missing as preserved. Diagnostic features of the new species include nasals that are sinusoid in lateral view, absence of a passage connecting the antorbital and maxillary fenestrae, relatively large teeth, plate-like chevrons forming a band along most of the length of the tail, and a long neck between the femoral head and shaft. The temporal constraints of the three paravian groups (Troodontidae, Dromaeosauridae and Aves) combined with the character distributions among the earliest known troodontids indicate a rapid evolution at the base of the Troodontidae.     

A New Boreopterid Pterodactyloid Pterosaur from the Early Cretaceous Yixian Formation of Liaoning Province, Northeastern China

<正>A new boreopterid pterosaur:Zhenyuanopterus longirostris gen.et sp.nov.from the Yixian Formation of western Liaoning is erected,based on the complete skeleton with a skull and lower jaws preserved.It is characterized by:a large boreopterid pterosaur with a high number of teeth,where the anterior teeth are much larger than posterior ones;the length of the dorsal + sacral vertebrae is nearly half the length of the skull;ratio of the length of the humerus to metacarpal IV is approximately 91%and the,humerus,femur and third wing phalanx are all equal in length and the feet are specially small.It represents the largest boroepterid pterosaur discovered from western Liaoning and its surrounding areas so far     

Pygostyle-like Structure from Beipiaosaurus (Theropoda, Therizinosauroidea) from the Lower Cretaceous Yixian Formation of Liaoning, China

Pygostyle was previously considered as a unique structure of ornithothoracine birds, used to maneuver tail feathers. A similar structure from an oviraptorosaurian dinosaur was considered functionally associated with the rectrices as in birds. We report a pygostyle-like structure from a therizinosauroid dinosaur. The presence of filamentous integuments, but absence of rectrices, on the tail of this therizinosauroid, combined with other lines of evidence, suggests that the initial function of the pygostyle was not related to the rectrices.     

A New Enantiornithes (Aves) from the Early Cretaceous of China

A new bird from the early Cretaceous Yixian Formation of Liaoning, China is described. This new species, Grabauornis lingyuanensis, shares several synapomorphies with the Enantiornithes. The specimen is relatively well preserved. The skeletal morphology of Grabauornis bears close resemblance to that of other Chinese members of this clade. The brachial index （the ratio between the lengths of humerus and ulna） is 0.95, which is close to the average for enantiornithine birds. It indicates that Grabauornis was a rather good flyer, and the presence of an alula in the wing further testifies to this.     

Largest Toothed Pterosaur Skull from the Early Cretaceous Yixian Formation of Western Liaoning,China, with Comments on the Family Boreopteridae

A new pterosaur Moganopterus zhuiana gen. et sp. nov. is erected based on a complete skull with lower jaws and anterior cervical vertebrae. It is characterized by much elongated upper and lower jaws with at least 62 total, long, curved teeth with sharp tips, a well developed parietal crest extending posterodorsally, forming an angle of 15 degrees with the ventral margin of the skull, the ratio of length to width of cervical vertebrae greater than 5:1. The skull length is 750 mm, and it is the largest toothed pterosaur found so far in the world. Based on this new pterosaur, the Boreopteridae can be divided into two subgroups: Boreopterinae sub-fam. nov. and Moganopterinae sub-fam. nov., which is also confirmed by the phylogenetic analysis.     

辽西义县组蝾螈类化石新发现

在辽西葫芦岛市附近的早白垩世义县组地层中发现1处新的蝾螈化石点。新台门一带出露的湖相沉积地层主要是一套灰绿色安山岩与紫色泥质粉沙岩的互层。该层中除了产蝾螈化石外,还有诸如Ephemeropsis,Eosestheria,Liaoningocladus等其他动植物化石。过去发现的Liaoxitriton化石点是在同一地区比新化石层稍高的层位中发现的。该类蝾螈化石曾报道产于早白垩世的九佛堂组,但是新近的野外调查表明,该化石点所在层位也属义县组。     

A New Ornithocheirid from the Early Cretaceous of Liaoning Province, China

Based on a nearly complete skeleton with skull from the Early Cretaceous of Liaoning Province, a new ornithocheirid pterosaur: Boreopterus cuiae gen. et sp. nov. is erected. Boreopterus cuiae is different from other pterosaurs preserved with skulls known from the western Liaoning Province and its neighboring areas. Tiffs new oterosaur has more and larger teeth than those in other ornithocheirids. Its anterior nine pairs of teeth are larger than other teeth. The fourth pair of upper and lower teeth are slightly larger than the third pair. Overall, Boreopterus cuiae shows much small range of tooth size variation than Anhanguerapiscator and Coloborhynchus robustus. The new taxon shares with other ornithocherids in having a relatively large size of the third and fourth pairs of teeth.     

中国辽宁中华侏罗兽的发现代表早期哺乳动物演化的一个新里程碑

真兽类哺乳动物是现代最为丰富多彩的哺乳动物类群,其起源和早期演化是国际科学界高度关注的科学问题之一.中国辽宁中侏罗世髫髻山组产出的中华侏罗兽是目前世界上发现的最古老的真兽类哺乳动物,代表了早期哺乳动物演化的一个新里程碑.研究表明,中华侏罗兽的发现将以前国际上的白垩纪化石记录提前了3500万年,填补了早期哺乳动物演化的化石记录间隔,帮助校正了现代基于DNA方法研究的哺乳动物演化历史,并与基于 DNA 方法测定的真兽类哺乳动物起源时间的研究结果相吻合,为哺乳动物演化历史建立了以化石为标准点的新标尺.中华侏罗兽的精确年龄有助于确定真兽类哺乳动物与其它哺乳动物,如后兽亚纲(它们的后代包括有袋类,如袋鼠)和单孔类(如鸭嘴兽),分异发展的时间.     

辽西热河生物群一新的巨龙型类恐龙

初步研究了辽西地区热河生物群的一具近完整的蜥脚类恐龙骨架,其代表了一新的巨龙型类恐龙——中国辽宁巨龙(Liaoningotitan sinensis gen.et sp.nov)。其主要特征为:上颌骨腹缘下凸,上颌齿列短且位于前部;颧骨前端近平齐于眶前窗的前缘;翼骨的方骨支基部收缩;上颌齿呈叠瓦状排列;上颌齿冠窄勺形,切面呈D字形,缺失唇侧沟和小锯齿;下颌齿9枚,明显小于上颌齿,排列稀松;下颌齿冠不对称,切面呈椭圆形,舌侧的沟和嵴发育,齿冠基部向舌侧膨大;肱骨近端扩展,约为长度的54.9%;肠骨前髋臼突前端较尖。初步的支序分析显示,辽宁巨龙属于巨龙型类恐龙,是比盘足龙和梁龙更为进步的类群。     

A New Specimen of Liaoceratops yanzigouensis (Dinosauria:Neoceratopsia) from the Early Cretaceous of Liaoning Province, China

A new specimen of basal neoceratopsian dinosaur Liaoceratops yanzigouensis is described. The specimen comes from the Lujiatun Bed of the Lower Cretaceous Yixian Formation in Beipiao City of western Liaoning Province, and is represented by a very well preserved three-dimensional partial skull and mandible. It is also the smallest among the three specimens of L. yanzigouensis, and several features in the new specimen, such as the short preorbital length and the round rostroventral orbital rim, can be ontogenetically-related. The superb exposure of the palatal complex may be caused by the removing of its brain by a small predator in the contemporary Jehol Biota.     

A New Giant Compsognathid Dinosaur with Long Filamentous Integuments from Lower Cretaceous of Northeastern China

1 Introduction Compsognathids are a small group of theropods commonly placed at the basal position within Coelurosauria (Sereno, 1999; Rauhut, 2003; Holtz and Osmólska, 2004; Norell and Xu, 2005). Recently new findings make the number of valid compsognathid genera increase rapidly. This group contains the relatively complete Compsoganthus (Ostrom, 1978; Bidar et al., 1972) and Juravenator (G?hlich and Chiappe, 2006) from the Upper Jurassic of West Europe, and Sinosauropteryx (Ji and J…     

辽宁西部翼龙类化石的首次发现

辽宁北票义组中的翼龙个体中等大小，尾短，具腹肋，前肢粗壮，桡、尺骨长于翼掌骨，骨肢跖骨极细长，第Ｖ趾退化但未消失等特征，表明它应为翼手龙亚目早期类型中一个新的属种－－杨氏东方翼龙。这是辽西翼龙类化石的首次发现，为进一步确定热河动物群的性质，地质时代，以及研究古地理古环境等提供了新资料。     

A New Angiosperm Genus from the Lower Cretaceous Yixian Formation, Western Liaoning, China

The abundant fossils of early angiosperms found in the Yixian Formation (125 Ma, Early Cretaceous) have demonstrated unexpectedly high diversity and various reproductive strategies, and thus have shed new light on early angiosperm evolution. Here, using light microscope and scanning electron microscope, we document several fructifications of Baicarpus gen. nov. from Liaoning province, China. Each fructification of the new taxon includes several apocarpous carpels/fruits attached in an urn-formed gynobase. Each fruit/carpel has a persistent straight style and an ovary enclosing a single seed. These fossil plants demonstrate a new floral organization and thus enhance our understanding of the diversity of angiosperms in the Yixian Formation. Together with previously reported ones, this fossil angiosperm suggests an origin time for angiosperms earlier than widely accepted.     

Postcranial Anatomy of the Mesozoic Dalinghosaurus (Squamata): Evidence from a New Specimen of Western Liaoning

The postcranial skeleton of a new specimen of the long-tailed lizard Dalinghosaurus longidigitus was excavated from the Yixian Formation in Lingyuan, western Liaoning. The new specimen provides more anatomical information about this species, especially about the anterior dorsal vertebrae, shoulder girdle and forelimbs. This lizard can be included within the clade Scleroglossa by its 27 or more presacrals, moderately long pubis, and gently notched distal end of tibia. But the detailed systematic position for this taxon remains undetermined. The features of the much longer hind limbs and pes compared with forelimbs and manus, metatarsal Ⅳ longer than Ⅲ, pedal phalanges robust, and penultimate phalanx not longer than other phalanges etc. suggest that this lizard was a running and ground swelling animal.     

辽宁凌源义县组恐龙皮肤印痕化石

辽宁西部凌源大王杖子义县组恐龙皮肤印痕化石的发现，是热河生物群中恐龙皮肤鳞片的又一次记录。这些鳞片的形态以不规则多边形为主、互不叠覆，同时鳞片的大小、排列方式等随其着生部位的不同而明显变化。从与该皮肤鳞片保存在一起的数枚上颌齿及部分前后肢骨来判断，这一恐龙化石应归鸟臀类，且极有可能为鹦鹉嘴龙类(psittacosaurids)。     

A New Troodontid Dinosaur from the Lower Cretaceous Yixian Formation of Liaoning Province, China

A new troodontid dinosaur,Daliansaurus liaoningensis gen.et sp.nov.,is erected based on a nearly complete specimen from the Lower Cretaceous Yixian Formation of Beipiao City,Liaoning Province,China.This well preserved skeleton provides important new details of the anatomy for Liaoning troodontids,and gives new insight into their phylogenetic relationships and evolution.Daliansaurus is distinguished from other troodontids by an enlarged ungual on pedal digit IV,which is approximately the same size as the sickle-shaped second ungual,and is differentiated from other Liaoning troodontids by a number of characters of the skull,manus,pelvis,and hindlimb.A phylogenetic analysis recovers Daliansaurus within a subclade of Liaoning troodontids that also includes Sinovenator,Sinusonasus,and Mei.We erect a name for this group—Sinovenatorinae—and argue that it reflects a localized radiation of small-bodied troodontids in the Early Cretaceous of eastern Asia,similar to previously recognized radiations of Liaoning dromaeosaurids and avialans.As more Liaoning theropods are discovered,it is becoming apparent that small,feathered paravians were particularly diverse during the Early Cretaceous,and future work is needed to clarify how this diversity arose,which species coexisted,and how these numerous species partitioned niches.     

Discovery of a New Pterosaur in Western Liaoning,China

The pterosaur from the Yixian Formation in Beipiao, Liaoning Province, ischaracterized by the medium-sized individual, short tail, presence of the gastralia, strongforelimbs, radius and ulna longer than wing-metacarpal, extremely narrow and elongatemetatarsals, degenerated and small phalanx V of hindlimb and so on. It must be a new genusand species, Eosipterus yangi gen. et sp. nov., in the early groups of the SuborderPterodactyloidea. This is the first record of pterosaurs found in Liaoning Province. The fossilprovides new evidence for further determining the nature and geological time of the JeholFauna and for studying the palaeogeography and palaeoenvironment.     

A Dichocarpum-like Angiosperm from the Early Cretaceous of China

The Yixian Formation (the Early Cretaceous) of China has yielded a diverse group of early angiosperms, revealing the charm of angiosperms from the formation is far from finishing. Here we report a fossil angiosperm, Nothodichocarpum gen. et sp. nov, with physically connected female parts, male parts, branches, and leaves with pinnate reticulate venation from the Yixian Formation of Liaoning, China. In addition to the previous reports, this report underscores the diversity of angiosperms in the Early Cretaceous Yixian Formation and earlier origin of angiosperms. The morphological variations in the reproductive parts in different stages allow us to infer the development of this early angiosperm. This fossil plant demonstrates a unique character assemblage that sheds new light on the origin and evolution of carpels.