Change in Extent of Meadows and Shrub Fields in the Central Western Cascade Range,Oregon∗

We examined change in areal extent of mountain meadows and fields of deciduous shrubs and conifer saplings in the central western Cascade Range of Oregon, based on analysis of aerial photographs taken in 1946 and 2000. These nonforest vegetation patches are distinctive habitats in dominantly forested landscapes, such as the Cascades, and change in extent of these habitats is of interest to scientists and land managers. We mapped and dated even-aged forest stands of probable postfire origin around the nonforest patches, using tree-ring analysis and interpretation of aerial photographs. We used these and archival data to interpret possible influences of past wildfire and sheep grazing on the extent of nonforest patches. The total area of nonforest vegetation patches decreased from 5.5 percent of the study area in 1946 to 2.5 percent in 2000. Significantly more cases of contracted patches were observed for mesic and xeric meadows which have adjacent forest established after wildfire in the approximately 150 years preceding 1946. A higher proportion of mesic (47 percent, n = 47) than xeric (17 percent, n= 115) meadows contracted between 1946 and 2000. Broad-leaved shrub fields were unchanged, probably because of topo-edaphic controls, dense cover of shrubs, and snow effects; but all fields of conifer saplings underwent succession to forest. We observed no strong influences of sheep grazing on the extent of meadows.     

羊啃食动力学模拟与内蒙古西部气候变化适应

将定性分析、民主参与式评估和定量数值模拟方法结合起来,对阿拉善气候变化适应问题进行了简要评估并重点分析了土地退化、放牧和草原管理策略之间的关系。阿拉善主要的适应政策是禁牧和生态移民。禁牧后荒漠草原植被得到恢复,但移民增加了绿洲压力,过度开垦又造成周边土地沙化;过度抽取地下水致使水位下降并伴随咸水入侵和地表盐碱化。禁牧的另一个副作用是造成草原鼠害加重及火灾风险增加,生物多样性也呈现先升后降趋势。一些灌木因缺乏啃食出现退化迹象等,说明一定程度的啃食有利于维持草原生态平衡。因此,动物是草原生态系统不可或缺的成员之一。我们用一个包含动物啃食等的森林生态动力模式模拟了啃食动力学,结果表明,有限的放牧有利于草场更新并能充分利用草原资源;过度放牧导致草场退化并伴随羊群死亡。因此,需要将圈养和限制性放牧结合起来,制定可持续性适应政策。     

A thousand years of environmental change and human impact in the alpine zone at Mt Kosciusko,New South Wales

This study reconstructs erosion, productivity, fire and vegetation records at Club Lake, in the alpine zone of Kosciusko National Park (the highest mountain region in Australia), and uses them to compare the prehistoric and historic periods. While disturbance in the prehistoric period was found to be minimal and mainly activated by fire, the impact of land uses after European arrival initiated a change in the erosion and fire regime and brought new grazing animals and exotic plant species. These triggered temporal changes in eutrophication and the nature of erosion, and significant vegetation changes. There was a reduction in the stability and persistence of species representation, especially in herbfield vegetation, and little recovery is evident despite the cessation of summer grazing. It is apparent that the area is very sensitive to disturbance by human impact and large fires.     

Effects of fire, grazing, and the presence of shrubs on Chihuahuan desert grasslands

Responses of herbaceous and suffrutescent species to fire, grazing, and presence of Prosopis glandulosa were examined in a Chihuahuan desert grassland in south-central New Mexico. Treatments were assigned randomly to eight 12×8 m plots within each of two blocks. Following fires in June 1995, unfenced plots were exposed to livestock grazing over 4 years. Plots were established that either included or excluded P. glandulosa. Perennial grass cover, primarilyBouteloua eriopoda , decreased by 13% in burned plots but increased 5% in unburned areas. Conversely, perennial forb cover was 4% greater after fire. Perennial grass frequency decreased 30% more and perennial forb frequency increased 10% more following burning. Further, increases in evenness after fire resulted in a 225% increase in species diversity. Grazing also resulted in a decrease in perennial grass cover while frequency decreased 22% more in grazed than ungrazed plots. Only frequency and not cover of perennial forbs and annual grasses increased more following grazing. Presence of P. glandulosa had no differential effect on responses of non-shrub species. Fires were conducted during near drought conditions while grazing occurred during years of precipitation equivalent to the long-term average. Precipitation immediately following fire may be critical for recovery of B. eriopoda -dominated desert grasslands; relationships between fire and post-fire precipitation patterns require future investigation.     

Thick moss layers and high cover of grasses: Potential threats to herb diversity in hay meadows in Norway

In an old, species-rich hay meadow in Varalds?y in Hardanger, the only known remaining managed site of its kind in West Norway, the moss layer is increasing in thickness in some places. Thick moss is associated with a high cover of monocotyledons, while a low cover of monocotyledons is associated with high numbers of herbs, including vulnerable species, which mainly occur in herb-rich areas. The increase in moss is probably caused by a reduction in both traditional means of moss removal and in sheep trampling and it may represent a potential threat to plant species diversity in traditionally managed hay meadows in the region. In order to increase the area of open, species-rich hay meadows, and thus the potential population sizes for indicator species of traditional management, increased grazing intensity (back to traditional levels) and removal of mosses during haymaking is recommended. The study demonstrates that, for conservation purposes, it is not easy to replicate past conditions unless the details of traditional management practices are known and understood, and these are now passing from general knowledge.     

Effects of fire on rangeland vegetation in south-western Mendoza plains (Argentina): composition,frequency, biomass,productivity and carrying capacity

The effect of both a non-prescribed summer fire and grazing at high stocking rate following fire on plant community composition, the frequency of occurrence of bare soil, grasses and shrubs, species diversity and biomass of herbaceous forage were evaluated for three growing seasons after fire. Changes in community composition occurred as a consequence of both fire and grazing. Communities were dominated by unpalatable shrubs and grasses in unburned sites as a result of a long history of overgrazing. Fire contributed to a conversion of those shrublands to communities with a more favourable balance between woody and herbaceous species. The frequency of palatable grasses and herbaceous forage biomass increased by a factor ofc. 3 at the expense of woody vegetation. Grazing after fire had significant positive (i.e. decrease in undesirable grasses) and negative (i.e. increase in bare soil) effects.     

Structural and nutritional heterogeneity of riparian vegetation in Patagonia (Argentina) in relation to seasonal grazing by sheep

Riparian meadows of southern Patagonia are temporally and spatially heterogeneous habitats. They are thought to play a key role in regulating the dynamics of arid grazed steppes of the region. We conducted a 2-year study with sheep to determine the influence of two grazing conditions (deferment and spring grazing) on structural and nutritional parameters of three vegetation types apparently associated with a soil moisture gradient in a riparian meadow in southern Santa Cruz (Argentina). Spring deferment allowed forage accumulation in very moist and intermediate vegetation types by the beginning of the summer and had no detrimental impact on forage quality. Deferment had no effect on biomass accumulation of the drier vegetation type at our study site. Structural attributes of the vegetation appeared to affect spatial grazing patterns of sheep. Soil-related patchiness, rather than sheep grazing, appeared to control vegetation structure and nutritional value. The proportion of more mesic plant communities in riparian habitats of Patagonia could determine the outcome of plant–animal interactions in these systems. Practical recommendations of grazing capacity in paddocks, or more theoretical considerations of ecosystem dynamics of the Patagonian steppes need to explicitly consider the internal patchiness of riparian habitats.     

Seedbank responses to spring and fall prescribed fire in mountain big sagebrush ecosystems of differing ecological condition at Lava Beds National Monument,California

Land use change in the American West has altered sagebrush dominated ecosystems, facilitating exotic grass invasion, increased woody species, altered fire regimes, and native species losses. Restoring fire regimes may be integral to the restoration of sagebrush ecosystems. However, little is known about how the timing of prescribed fire differentially impacts the soil seedbank. Further, sites of different ecological condition may variably respond to fire. To address these uncertainties, we quantified germination of invasive cheatgrass and plant functional groups (native dicots, exotic dicots, native bunchgrasses, and native shrubs) following spring and fall prescribed burns in three mountain big sagebrush communities at Lava Beds National Monument, California. At cheatgrass and native-dominated sites, there were fewer cheatgrass seedlings (91% and 86%, respectively) immediately following spring fire than in unburned controls, but this reduction did not persist one year later. Fall burns did not significantly impact cheatgrass or exotic dicot germination. At a native dominated site, native dicots increased one year following spring (1620%) and fall burns (67%), suggesting adaptation to a natural fire regime. This research shows that the prefire condition of an ecosystem greatly influences the postfire germination response, and that fire-adapted ecosystems can benefit from the natural disturbance regime.     

Structural and nutritional heterogeneity of riparian vegetation in Patagonia (Argentina) in relation to seasonal grazing by sheep

Riparian meadows of southern Patagonia are temporally and spatially heterogeneous habitats. They are thought to play a key role in regulating the dynamics of arid grazed steppes of the region. We conducted a 2-year study with sheep to determine the influence of two grazing conditions (deferment and spring grazing) on structural and nutritional parameters of three vegetation types apparently associated with a soil moisture gradient in a riparian meadow in southern Santa Cruz (Argentina). Spring deferment allowed forage accumulation in very moist and intermediate vegetation types by the beginning of the summer and had no detrimental impact on forage quality. Deferment had no effect on biomass accumulation of the drier vegetation type at our study site. Structural attributes of the vegetation appeared to affect spatial grazing patterns of sheep. Soil-related patchiness, rather than sheep grazing, appeared to control vegetation structure and nutritional value. The proportion of more mesic plant communities in riparian habitats of Patagonia could determine the outcome of plant–animal interactions in these systems. Practical recommendations of grazing capacity in paddocks, or more theoretical considerations of ecosystem dynamics of the Patagonian steppes need to explicitly consider the internal patchiness of riparian habitats.     

干扰对生物土壤结皮及其理化性质的影响

生物土壤结皮作为荒漠地区特殊环境的产物,具有较强的抗风蚀、水蚀功能,也是干旱荒漠地区植被演替的重要基础。随着人类活动的加剧,生物土壤结皮也受到不同类型和不同程度的干扰,主要包括放牧、火烧、车辆碾压等形式。干扰对生物土壤结皮的影响主要表现在生物土壤结皮结构及盖度的变化、土壤理化性质的改变、土壤微生物数量及活性的变化等几个方面。放牧对土壤物理性质的影响还没有一致的结论,除了干扰的程度还与土壤含水量有关系。火烧虽然改善了土壤结构,但是破坏了地表植被的盖度,而且恢复比较困难,也加速了外界侵蚀的力度,对土壤化学性质也有显著影响。机械碾压的破坏力最强,且没有任何的积极意义。     

Paleoecology and its application to fire and vegetation management in Kootenay National Park, British Columbia

High-resolution analysis of macroscopic charcoal and pollen ratios were used to reconstruct a 10,000 yr history of fire and vegetation change around Dog Lake, now in the Montane Spruce biogeoclimatic zone of southeastern British Columbia. Lake sediment charcoal records suggest that fire was more frequent in the early Holocene from 10,000 to 8200 calendar yrs BP, when climate was warmer and drier than today and forest fuels were limited. Fire frequency increased and reached its maximum during the early to mid-Holocene from 8200 to 4000 calendar yrs BP, corresponding to the dry and warm Hypsithermal period in the Rocky Mountains. During the Hypsithermal period forests around Dog Lake were dominated by Pseudotsuga/Larix,Pinus and open meadows of Poaceae that were subject to frequent fire. From 4000 calendar yrs BP to present, fires became less frequent with the onset of cooler and wetter Neoglacial climate and an increase in wet-closed Picea and Abies forests in the valley. Changes in fire frequency are supported by dry-open/wet-closed pollen ratio data indicating that forest type and disturbance regimes vary with changing climate. The fire frequency and forest cover reconstructions from Dog Lake are a first attempt at defining a range of natural variability for Montane Spruce forests in southeastern British Columbia. Fire and vegetation management in Kootenay National Park can now use this century to millennial-scale range of variability to define the context of current forest conditions and potential changes under global warming scenarios.     

A review on the eco-geography of soil fauna in China

This review summarizes main research findings in soil fauna eco-geography in China in the past 30 years. The subject areas and main results were overviewed including biodiversity and eco-geological distribution of soil fauna communities. Studies of ecological distributions of soil fauna and dynamic ranges in space from tropical, subtropical to temperate regions, and in categories from forest, grassland, desert, wetland, farmland to urban ecosystems, the responses and indications of soil fauna to soil environments. Effects of intensive disturbance such as fire, grazing, farming, fertilization on soil fauna include sensitive (e.g. nematode) and rare groups and community indexes. The functions of soil fauna were discussed including environmental construction, environmental purification, litter decomposition and elements cycling. Interactions between soil fauna and other biota in soil ecosystems and linking between aboveground and belowground diversity and the effects of global change on soil fauna community in China were also included. Finally, the authors pointed out common interests in soil fauna eco-geographical studies, which include application of molecule biology into soil fauna taxa; function and mechanism of soil fauna community diversity; interaction between aboveground and belowground ecosystems; effects of disturbance, pollution, biological invasion, and global change on soil fauna community and function. The review is to provide a scientific basis for promoting soil fauna eco-geographical studies in China.     

Effects of grazing and climate change on sandy grassland ecosystems in Inner Mongolia

To understand the effects of grazing activities and climate change on sandy grassland ecosystems in northern China, a livestock field grazing and enclosure experiment was conducted from 1992 to 2006 in Horqin Sand Land, Inner Mongolia. The results showed that sustained heavy grazing resulted in serious degradation of the vegetation; moderate grazing can maintain vegetation stabilization; and light grazing can promote rapid restoration of degraded vegetation. The livestock productivity was the highest in the moderate grazing grassland, and sustained heavy grazing resulted in rapid decrease of the livestock productivity. Heavy grazing can cause a retrogressive succession of grassland vegetation, whereas moderate and light grazing may promote progressive succession of plant species. The effects of changing climate on succession processes were not significant in the short term; a warm-humid climate is favorable to restoration of degraded vegetation, whereas a sustained warm-drought climate may result in degradation of grassland vegetation. Heavy livestock grazing should be stopped for the sustainable use of grassland; the proper grazing intensity for sandy grassland is two to three sheep or sheep equivalents per hectare in Inner Mongolia.     

Rocky Substrate and the Lower Treeline Ecotone of Yellowstone's Northern Range

Lower treeline ecotones are influenced by several processes, including fire, grazing, and climatic variables, but rocky substrate is an under-recognized factor affecting their patterns and dynamics. On the Northern Range of Yellowstone National Park, we hypothesized that north-facing slopes and riparian zones would explain the distribution of most forested areas, but that rocky substrate would explain a substantial additional portion of the lower treeline, and that advancement of conifers into grassland has occurred along rocky substrate. We used GIS software and change detection analysis to assess the spatial distribution and change of the forest with respect to north-facing slopes, riparian areas, and rocky substrate. As in other lower treeline studies, we found that north-facing slopes and riparian zones contained the majority of forested land (63%). However, areas with rocky substrate accounted for more forested area than north-facing slopes and riparian zones combined (67.4%), including 28% of forest that was not explained by the first two variables. Change detection analysis between 1954 and 2006 revealed that conifer advancement was insignificant but the number of forested patches increased and the mean size of patches decreased. These results suggest a more fragmented lower treeline environment that may indicate forest advancement in small patches. Field surveys revealed that rocky substrate did not provide conifers with protection from browsing ungulates or fire, yet conifers preferentially established on rocky areas. Our landscape analysis suggests that weathered rock increases soil coarseness in a landscape dominated by fine glacial till, which improves conditions for seedling establishment by reducing competition with grasses and shrubs sufficiently to overcome the negative influence of dry, nutrient-poor microsites in a moisture-stressed environment.     

HUMAN IMPACTS ON THE ENVIRONMENT: UNPREDICTABILITY AND THE PRIMACY OF PLACE

Landscapes, environments, and ecosystems (i.e., places) are historically and spatially contingent and therefore unique. Generalizations and laws are based on “all other things being equal.” Owing to historical and spatial contingency, all other things are never completely equal and often do not approach that ideal closely enough to allow for reliable prediction. The problem is illustrated by examining effects of grazing and fire suppression on transitions between grass and woody vegetation communities. In three different environments in the southern United States (central Texas, southwestern Virginia, and eastern North Carolina), three different relationships exist between grazing and fire regime, and vegetation. Other examples are common in the geographic and environmental literature. This suggests that the search for a single applicable generalization or predictive model of a given human-induced environmental change that can be applied without accounting for the particular characteristics of places may be futile. Several general ways to address this issue are suggested. First is the recognition that place matters. Conditions specific to a location or region will influence what, how, or even if generalizations apply. Second is the adoption of a “demographic” approach. This is based on the idea that for places, like people, it is possible to make aggregate or probabilistic generalizations or predictions, but these cannot be applied to deterministic prediction of individuals. Third is a synoptic approach based on forecasting not via simple cause-effect relationships, but based on a typology of situations that recognizes and incorporates the unique characteristics of site and situation. Several converging threads of inquiry in geography and geosciences explicitly or implicitly address the issue of historical and spatial contingency. [Key words: landscape, environment, place, prediction.]     

Driving Mechanism of Gross Primary Production Changes and Implications for Grassland Management on the Tibetan Plateau

The contribution of climatic change and anthropogenic activities to vegetation productivity are not fully understood. In this study, we determined potential climate-driven gross primary production (GPPp) using a process-based terrestrial ecosystem model, and actual gross primary production (GPPa) using MODIS Approach in alpine grasslands on the Tibetan Plateau from 2000 to 2015. The GPPa was influenced by both climatic change and anthropogenic activities. Gross primary production caused by anthropogenic activities (GPPh) was calculated as the difference between GPPp and GPPa. Approximately 75.63% and 24.37% of the area percentages of GPPa showed increasing and decreasing trends, respectively. Climatic change and anthropogenic activities were dominant factors responsible for approximately 42.90% and 32.72% of the increasing area percentage of GPPa, respectively. In contrast, climatic change and anthropogenic activities were responsible for approximately 16.88% and 7.49% of the decreasing area percentages of GPPa, respectively. The absolute values of the change trends of GPPp and GPPh of meadows were greater than those of steppes. The GPPp change values were greater than those of GPPh at all elevations, whereas both GPPp and GPPh showed decreasing trends when elevations were greater than or equal to 5000 m, 4600 m and 4800 m in meadows, steppes and all grasslands, respectively. Climatic change had stronger effects on the GPPa changes when elevations were lower than 5000 m, 4600 m and 4800 m in meadows, steppes and all grasslands, respectively. In contrast, anthropogenic activities had stronger effects on the GPPa changes when elevations were greater than or equal to 5000 m, 4600 m and 4800 m in meadows, steppes and all grasslands, respectively. Therefore, the causes of actual gross primary production changes varied with elevations, regions and grassland types, and grassland classification management should be considered on the Tibetan Plateau.     

扎龙湿地火烧严重度分析及火灾对丹顶鹤生境的影响

利用ETM 遥感影像资料,分析了扎龙湿地火烧严重度及火灾对扎龙湿地丹顶鹤(Grus japonensis)生境的影响。采用火灾前、后的ETM 影像数据,计算差异性归一化燃烧率(dNBR)和制作扎龙火烧严重度分布图,用综合燃烧指数(CBI)分析植被死亡率对生境变化的影响程度,并分析两者的相关关系;利用两种燃烧指数与对应的高程、植被类型、坡度、功能分区和丹顶鹤巢位等因子叠加分析作为火烧区的评价指标;再根据火烧严重度分类图定量化分析火灾对丹顶鹤生境和巢址变化的影响。结果表明,CBI与dNBR之间有非线性关系(n=59,R2=0.8724);判定高程在148.3~154.6m和坡度在0°~0.185°之间,且分布有大量芦苇(Phragmites australis)沼泽的核心区低洼地易发生火灾;在火烧强度较高的情况下,迫使丹顶鹤选择次适宜的栖息环境,降低了丹顶鹤对隐蔽条件和食物资源的标准,导致丹顶鹤重新选择巢址的位置,其种群数量迅速减少。研究结果对扎龙湿地动物资源可持续发展具有一定的指导意义。     

The Coyote Brush Invasion of Southern California Grasslands and the Legacy of Mechanical Disturbance

This study investigates the recent invasion of exotic grasslands by coyote brush in La Jolla Valley, California. We test the “event dependent” hypothesis that mechanical disturbances during the past century were a key cause. To examine the relationship between past mechanical disturbances and vegetation dynamics we first conducted a review of the historical literature on practices of shrub removal and documented disturbance history using historical imagery. We next analyzed vegetation‐cover change over time using remotely sensed imagery and a vegetation map to document the history of native shrub advancement into exotic grassland by species association. Finally, we determined the topographic characteristics associated with different phases of shrub advancement. We found that mechanical disturbances historically varied by topography with upper and steeper slopes being least intensively disturbed. We found that shrub advancement rates, following release from grazing, varied by slope, elevation, and time period, and that Baccharis pilularis (coyote brush) was the main species to invade the more intensively disturbed sites at lower elevations. Our results indicate that mechanical disturbances played an important role in modifying the original vegetation cover with long‐lasting consequences, including a facilitating role for the subsequent Baccharis pilularis invasion. We concluded that the practice of grazing often included exotic mechanical disturbances that had long‐lasting impacts on native plants.     

Studying the post-fire performance of tussock grasses in Patagonia: Survival, biomass production and early competition

Patagonia grasslands are subjected to two main disturbances, fire and grazing, but little information is available about its effects on vegetation. We studied post-fire survival and resprouting ability of two dominant grass species, Stipa speciosa and Festuca pallescens, for four years; evaluated the effect of early post-fire defoliation on both species; and tested whether competition is important in post-fire recovery in San Ramón Ranch, NW Patagonia (Argentina). To simulate grazing, a clipping treatment was applied at the beginning and end of growing seasons. Survival rates were high (>60%) and, after three years, biomass of both species in the burned area was similar to the unburned area. Competition seems to play an important role in the early post-fire recovery of both species, particularly in the case of F. pallescens that increased 87% the biomass production without competition. Fire may improve forage quality by eliminating the standing dead material, but early post-fire grazing might endanger the persistence of F. pallescens. We suggest requiring a resting period before livestock introduction and controlling grazing intensity.     

科尔沁沙质放牧草地土壤动物多样性特征研究

在科尔沁沙质放牧草地,人为的小面积实验放牧情况下,土壤动物群落结构随放牧强度的提高发生变化,主要体现在动物密度降低、群落丰富度减小。放牧强度提高时,土壤动物生物量、密度季节动态和多样性指数无明显变化规律,重度放牧下以上指标并不明显低于封育区。土壤动物优势类群对放牧扰动的响应各自不同。以相似性指数比较土壤动物对不同牧压的响应,结果为中度放牧区较接近封育区,而重牧区分异最大。