Miocene pinnipeds of the otariid subfamily Allodesminae in the North Pacific Ocean: Systematics and relationships

Abstract Fossil pinnipeds in the extinct otariid subfamily Allodesminae are large, relatively highly evolved marine carnivores that became abundant and diverse in Middle Miocene time and were restricted to the North Pacific Ocean. Their record extends from early Middle Miocene through Late Miocene, with records from California, Oregon, Washington, Baja California and Japan. Allodesmines are characterized by extreme sexual dimorphism, a large orbit, retracted orbital margin of the zygomatic arch, a deeply mortised jugal-squamosal junction, wide palate, bulbous cheek tooth crowns, nearly flat tympanic bulla with wrinkled ventral surface, a large tympanohyal fossa, large ear ossicles and deep mandible. Eleven allodesmine species are known (eight of which are named), in at least four genera, and most belong to the typical genus Allodesmus Kellogg, 1922. The earliest and most generalized allodesmine known is from the early Middle Miocene (ca 16 Ma) Astoria Formation in coastal Oregon. The last known records are from Late Miocene rocks (ca 10 Ma) in California and Washington. New taxa proposed here are: the genus Brachyallodesmus Barnes and Hirota, to contain Allodesmus packardi Barnes, 1972; the genus Megagomphos Hirota and Barnes, to contain Allodesmus sinanoensis (Nagao, 1941); the species Allodesmus sadoensis Hirota, (Middle Miocene, Japan); the species Allodesmus megallos Hirota (Middle Miocene, Japan); and the species Allodesmus gracilis Barnes (Middle Miocene, California). Additionally, the genus Atopotarus Downs, 1956, and the species Allodesmus kelloggi Mitchell, 1966, are resurrected. Allodesmines were apparently a rapidly evolving group, and most appear to have been adapted to roles later filled by otariine, dusignathine and imagotariine otariids, and the Phocidae (true seals). They became extinct in Late Miocene time and left no living descendants. Although some of their characters evolved convergently with various living species of the pinniped family Phocidae, Allodesminae are an otariid group and not part of the evolutionary history of Phocidae.     

Miocene fossil pinnipeds of the genera Prototaria and Neotherium (Carnivora; Otariidae; Imagotariinae) in the North Pacific Ocean: Evolution, relationships and distribution

Abstract Fossil otariid pinnipeds of the extinct genera Prototaria Takeyama and Ozawa, 1984, and Neotherium Kellogg, 1931, known from Middle Miocene deposits bordering the North Pacific Ocean, are small, primitive pinnipeds in the subfamily Imagotariinae. They have a small supraorbital process of the frontal or have lost it entirely, a three-rooted first molar, small paroccipital process, and ear morphology indicating that they belong in the subfamily Imagotariinae. Their unique derived characters include extreme intertemporal constriction and highly modified cheek teeth, the premolars having become molarized by the addition of protocones and lingual cingula. Prototaria Takeyama and Ozawa, 1984, the most primitive known imagotariine genus, contains two species, P. primigena Takeyama and Ozawa, 1984, and P. planicephala Kohno, 1994, both of early Middle Miocene age from Japan. Prototaria has a few derived characters, including a large antorbital process, narrow intertemporal region, and large orbit, but its primitive characters apparently were inherited from enaliarctine ancestors. The long enigmatic Neotherium mirum Kellogg, 1931, of Middle Miocene age from California, USA, is related to Prototaria, but differs by having an elongate skull, very slender zygomatic arch, ventrally exposed median lacerate foramen, and smaller but more molarized premolars. A more primitive new genus and species, Proneotherium repenningi Barnes, related to N. mirum, is from the early Middle Miocene Astoria Formation, coastal Oregon, USA. It shares some derived characters with Prototaria, and shares many other important derived characters with N. mirum. Imagotariines probably arose from some species of Early Miocene enaliarctines, became diverse in Middle and Late Miocene time, and are only known from the North Pacific realm. Although they might include the ancestors of true walruses of the subfamily Odobeninae, no known imagotariines appear to have been adapted for mollusk feeding as are the highly evolved modern walruses. Instead, imagotariines appear to have retained a primitive piscivorous diet, as did the fur seals and sea lions of the subfamily Otariinae.     

A new fossil kentriodontid dolphin (Cetacea; Kentriodontidae) from the Middle Miocene Takinoue Formation, Hokkaido, Japan

Abstract Kentriodon hobetsu , a new species, is described from a partial skull from the Middle Miocene Takinoue Formation, in Hokkaido, Japan. The holotype skull includes the base of the rostrum and a fairly well preserved braincase. The new species is distinguished clearly from previously described species of Kentriodon Kellogg, 1927, by the shape of the anterior margin of the pterygoid sinus fossa, the large premaxilla, rounded facial margin, and lesser development of rugosities and processes. This new record of Kentriodon , the first named kentriodontid dolphin from the western North Pacific, extends the geographic range of the genus significantly.     

Authigenic gypsum found in gas hydrate-associated sediments from Hydrate Ridge, the eastern North Pacific

Hydrate Ridge is located at the second accretion-ary ridge along the Cascadia margin of Oregon in the eastern North Pacific (fig. 1). The Bottom Simulating Reflector (BSR) underlies the entire Hydrate Ridge[1]. The Ocean Drilling Program (ODP) in 1992 at Site 892 and the TECFLUX99 and 2000 showed that the gas hydrate occurs just beneath the thin sediment- covered surface and at the horizon of around 64 meter below seafloor (mbsf) on Hydrate Ridge[25]. The col-lision of the Juan de …     

North Pacific sea ice cover, a predictor for the Western North Pacific typhoon frequency?

The relationship between the sea ice cover in the North Pacific and the typhoon frequency has been studied in this paper. It follows that the index for the sea ice cover in the North Pacific (ISA) both in December-January-February (DJF) and in March-April-May (MAM) is negatively correlated with annual typhoon number over the western North Pacific (TNWNP) during 1965―2004, with correlation coeffi-cients of -0.42 and -0.49 respectively (above 99% significant level). Large sea ice cover in the North Pacific tends to decrease TNWNP. Positive ISA (MAM) is associated with the tropical circulation and SST anomalies in the North Pacific, which may lead to unfavorable dynamic and thermal conditions for typhoon genesis over WNP from June to October (JJASO). The variability of the atmospheric circula-tion over the North Pacific, associated with the ISA anomaly in MAM is connected to the tropical at-mospheric circulation variability in MAM via the teleconnection wave train. Besides, as the tropical circulation has strong seasonal persistency from the MAM to JJASO, thus, the ISA in MAM-related variability of the tropical atmospheric circulation as well as the SST can affect the typhoon activity over the western North Pacific.     

Relationships between the North Pacific Oscillation and the typhoon/hurricane frequencies

Relationships between the North Pacific Oscillation (NPO) and the typhoon as well as hurricane fre-quencies are documented. The correlation between NPO index in June-July-August-September and the annual typhoon number in the western North Pacific is 0.37 for the period of 1949―1998. The NPO is correlated with the annual hurricane number in the tropical Atlantic at -0.28 for the same period. The variability of NPO is found to be concurrent with the changes of the magnitude of vertical zonal wind shear, sea-level pressure patterns, as well as the sea surface temperature, which are physically asso- ciated with the typhoons and hurricanes genesis. The NPO associated atmospheric circulation vari- ability is analyzed to explain how NPO is linked with variability of the tropical atmospheric circulation in the western Pacific and the tropical Atlantic, via the atmospheric teleconnection.     

A new type of primitive baleen whale (Cetacea; Mysticeti) from Kyushu, Japan

Abstract An important and relatively complete fossil mysticete cetacean specimen was collected from the Oligocene age Ashiya Group in Kyushu, Japan. The specimen includes a complete skull, with the anterior part of the vertebral column, several ribs, and the left pectoral limb. It provides significant information about the early evolution of cetotheres and transitional steps from the Archaeoceti to Mysticeti. Several features of the specimen, such as the anterior position of the nasal opening and large mandibular foramen, are primitive, but others are derived, such as the wide maxilla, broad supraorbital process of the frontal, occipital shield overhanging the temporal fossae, and the flat articulation of the distal end of the humerus. This animal, therefore, has a mixture of primitive and derived characters. It represents a stage of evolution in mysticetes that has been previously unrecognized.     

南太平洋和北太平洋年代际振荡与华北盛夏降水的关系及可能物理机制

利用降水、大气环流和海表温度等多种再分析资料和偏相关方法,研究了1951—2007年南太平洋年代际振荡（SPDO）和北太平洋年代际振荡（即PDO,本文称为NPDO）分别与华北盛夏（7—8月）降水在年代际时间尺度上的关系及其可能物理机制.结果表明：在去除SPDO和NPDO的相关性之前,它们与华北盛夏降水的关系均偏弱;但在去除两者相关性之后,SPDO（NPDO）与华北盛夏降水存在显著正（负）相关关系.去除两者相关性之后,当SPDO处于正位相时,热带西北太平洋海温异常显著偏暖,这将在对流层中下层从热带西太平洋—东亚沿岸激发出"气旋-反气旋-气旋"的负位相东亚—太平洋型遥相关（EAP）波列,该波列导致东亚夏季风异常增强,有利于低纬地区水汽输送至华北地区,从而使得华北盛夏降水异常偏多,反之,当SPDO处于负位相时,华北盛夏降水异常将偏少;对NPDO来说,当其处于正位相时,不仅热带西北太平洋异常显著偏冷,而且印度洋大部分海温异常显著偏暖,在两者共同作用下,对流层中下层从热带西太平洋—东亚沿岸出现"反气旋-气旋-反气旋"的正位相EAP波列,这将引起东亚夏季风异常减弱,不利于低纬地区水汽输送至华北地区,华北盛夏降水异常因此减少,反之,当NPDO处于负位相时,华北盛夏降水异常将偏多.     

Classification and distribution of Oligocene Aetiocetidae (Mammalia; Cetacea; Mysticeti) from western North America and Japan

Abstract Fossil whales in the very rare, primitive, extinct cetacean family Aetiocetidae are small, relict, toothed mysticetes that persisted into Late Oligocene time after more highly derived baleen-bearing mysticetes had already evolved. No known aetiocetid could be ancestral to baleen-bearing mysticetes, but aetiocetid morphology is in many ways intermediate between archaeocetes and baleen-bearing mysticetes, demonstrating the probable transitional steps passed through in the evolution of baleen-bearing mysticetes. Their discovery indicates that mysticetes evolved from Archaeocetes, and supports theories of the monophyly of Cetacea. Late Oligocene aetiocetids have been found on both sides of the North Pacific Ocean: on Vancouver Island, British Columbia, Canada; in Oregon and Washington, USA; in Baja California Sur, México; and the islands of Kyushu and Hokkaido, Japan. The most primitive North American aetiocetid, Chonecetus sookensis Russell, 1968, is from the early Late Oligocene Hesquiat Formation on Vancouver Island, British Columbia, Canada. A more derived, Late Oligocene species, Chonecetus goedertorum Barnes and Furusawa, new species, from the Late Oligocene Pysht Formation, Olympic Peninsula, Washington, has the primitive placental mammalian tooth count of 11/11. The type genus of the family, Aetiocetus Emlong, 1966, has as its type species, A. cotylalveus Emlong, 1966, known only from the Late Oligocene Yaquina Formation on the coast of Oregon. It has 11 upper teeth on each side of the rostrum. A more derived species, Aetiocetus weltoni Barnes and Kimura, new species, from a higher stratigraphic level in the Yaquina Formation, has a more posteriorly positioned cranial vertex and a tooth count of 11/12. We describe four new species of aetiocetids in three genera from the Late Oligocene Morawan Formation near Ashoro, Hokkaido, Japan. The most primitive, Ashorocetus eguchii Barnes and Kimura, new genus and species, has a primitive stage of cranial telescoping, and is closely related to Chonecetus Russell, 1968. Another, Morawanocetus yabukii Kimura and Barnes, new genus and species, in some ways intermediate between Chonecetus and Aetiocetus, has a suite of unique derived characters, including a much foreshortened brain case. The third, Aetiocetus tomitai Kimura and Barnes, new species, is the most primitive species of Aetiocetus yet discovered. The fourth, Aetiocetus polydentatus Sawamura, new species, the most derived species of Aetiocetus known, has a highly telescoped cranium, homodonty, polydonty and a dental count of 13–14/14–15. The fossil record now indicates considerable diversity in the family, with several different contemporaneous lineages in three new subfamilies: Chonecetinae, Morawanocetinae and Aetiocetinae. Aetiocetids are not known outside the North Pacific. Many Recent mysticetes are essentially cosmopolitan, and aetiocetids might have also been relatively widely dispersed. We suspect that with time their remains will be found around other ocean basins also. If so, then they may be potentially useful in trans-oceanic geological correlations.     

Water exchange between the subpolar and subtropical North Pacific in an OGCM

The water exchange between the subpolar and subtropical gyres of the North Pacific is demonstrated by the simulation of chlorofluorocarbon (CFC) using an ocean general circulation model. The simulated CFC concentration in the North Pacific is in good agreement with observations. The water exchange is clearly illustrated by the tongues of CFC concentration. The subpolar waters with high CFC are transported southward into the eastern subtropical gyre, whereas the subtropical waters with low CFC are transported northward into the western subpolar gyre. The simulated exchange transport along 42°N in the layer of σθ< 26.8 indicates that the northward mass transport is about 15 Sv (1 Sv = 106 m3·s-1) west of 165°E, and about 5 Sv between 175°W and 150°W. The southward mass transport is about 5 Sv between 165°E and 175°W, and about 2 Sv east of 150°W.     

Summary of taxa and distribution of Sirenia in the North Pacific Ocean

Abstract North Pacific fossil sirenians comprise representatives of three subfamilies of the Dugongidae: Halitheriinae ( Metaxytherium arctodites , Middle Miocene, North America), Hy-drodamalinae ( Dusisiren spp., Early-Late Miocene, and Hydrodamalis spp., Late Miocene-Pleistocene, North America and Japan), and Dugonginae ( Dioplotherium allisoni , Early-Middle Miocene, North America). Indeterminate dugongid remains are also known from the Late Oligocene of Japan, and the discovery of additional taxa in the western Pacific, especially in Paleogene rocks, can be anticipated. The known North Pacific Neogene taxa apparently dispersed into the Pacific from the Caribbean. Metaxytherium gave rise in the Pacific to Dusisiren ; a series of chronospecies of the latter genus eventually culminated in Hydrodamalis , which was exterminated by humans circa AD 1768. Dioplotherium left no known descendants in the Pacific. The Recent Dugong probably entered the Pacific from the Indian Ocean. The presence in the North Pacific Miocene of at least three sympatric dugongid lineages, together with desmostylians, is evidence for a diversity of marine plants that was reduced by subsequent climatic cooling.     

北太平洋风暴轴的变异特征及其与中纬度海气耦合关系分析

本文使用欧洲ECMWF（ERA40）再分析资料,通过经验正交函数（Empirical Orthogonal Function,EOF）分解探讨了冬季北太平洋风暴轴的变异特征,使用回归分析得到了与风暴轴空间异常型相关的冬季大气平均流异常、表层海温（Sea Surface Temperature,SST）异常的空间耦合型.研究结果表明,冬季北太平洋风暴轴主要有两种空间异常型,第一种是风暴轴中东部明显北抬（南压）,使得整个风暴轴向东北（东南）倾斜,与此同时,在中纬度北太平洋海区,冬季暖（冷）异常的洋面上是异常高压（低压）,海气系统在垂直向表现为一种暖脊（冷槽）配置,在对流层中高层是太平洋-北美（Pacific-North American,PNA）型负（正）位相.第二种是风暴轴整体性加强（减弱）并偏北（南）,此时,黑潮区海温异常偏暖（冷）,低层阿留申低压和高空的西风急流略偏北（南）,对流层中高层表现为西太平洋（Western Pacific,WP）型负（正）位相.风暴轴EOF分解的时间系数与阿留申低压指数、PNA指数、WP指数,以及与尼诺3区（NINO3）指数、黑潮海温指数间显著的相关性再次证实了在北太平洋中纬度地区存在着SST异常、风暴轴异常和大气平均流异常三者间的空间耦合型.     

A new species of Middle Miocene sperm whale of the genus Scaldicetus (Cetacea; Physeteridae) from Shiga-mura, Japan

Abstract A nearly complete skeleton of a fossil sperm whale from the Middle Miocene age Bessho Formation at Shiga-mura, Nagano Prefecture, the most complete fossil physeterid skeleton found in Japan, is here named Scaldicetus shigensis, new species. Its abruptly tapered rostrum, deep supracranial basin, and high occipital crest are typical of physeterids. Large teeth with crenulate enamel on conical crowns, present in both the palate and mandible, are consistent with the genus Scaldicetus du Bus, 1867, a genus originally based on fossils discovered in the Antwerp Basin, Belgium. Scaldicetus shigensis is relatively primitive, having a relatively long, slender rostrum, large zygomatic arches, large tympanic bullae, prominent occipital condyles, low tooth count and a sloping occipital shield that is deeply emarginated laterally by large temporal fossae. The probable mandibular tooth count is 12, only one more than the primitive eutherian mammalian dentition. Despite its abundant primitive characters, Scaldicetus shigensis has very asymmetrical external nares, comparable to the living sperm whale, Physeter catodon, and is in this way more derived than most of the contemporaneous fossil physeterids. At a time when other sperm whales were more highly evolved, Scaldicetus shigensis retained a primitive occipital shield, large tympanic bullae, and low tooth count, as in primitive Oligocene odontocetes such as Agorophiidae. It was a relict form in Middle Miocene time, and provides an indication of a primitive stage of sperm whale evolution heretofore undocumented. Fossils from elsewhere previously referred to as Scaldicetus, largely on the basis of tooth characters, range in age from Early Miocene to Plio-Pleistocene time. Cranial material that would clarify relationships is lacking for most of these, and this time range seems too long for a cetacean genus. Because of this simplistic approach to identifications, Scaldicetus is probably a grade taxon, not a natural biological grouping. However, the tooth morphology of Scaldicetus shigensis is correct for the genus Scaldicetus, so we provisionally assign it to this genus. Redefinition of the genus Scaldicetus and revision of the Physeteridae are beyond the scope of the present study.     

Relationship between the Asian-Pacific oscillation and the tropical cyclone frequency in the western North Pacific

The relationship between the Asian-Pacific oscillation (APO) and the tropical cyclone frequency over the western North Pacific (WNP) in summer is preliminarily investigated through an analysis of ob- served data. The result has shown clearly that APO is significantly and positively correlated to the tropical cyclone frequency in the WNP. If APO is above (below) the normal in summer, more (less) tropical cyclones will tend to appear in the WNP. The present study also addresses the large-scale at- mospheric general circulation changes underlying the linkage between APO and the WNP tropical cy- clone frequency. It follows that a positive phase of summer APO is concurrent with weakened as well as northward and eastward located western Pacific subtropical high (WPSH), low-level convergence and high-level divergence, and reduced vertical zonal wind shear in the WNP, providing favorable envi- ronment for the tropical cyclone genesis, and thus more tropical cyclones will come into being, and vice versa.     

New predictors and a new prediction model for the typhoon frequency over western North Pacific

In this paper, the impacts of the atmospheric circulation during boreal winter-spring on the western North Pacific (WNP) typhoon frequency (WNPTF) are studied. Several new factors in winter-spring in- fluencing the typhoon frequency were identified, including the sea ice cover in the North Pacific and the North Pacific oscillation. Based on these results, the multi-linear regression was applied to establishing a new forecast model for the typhoon frequency by using the datasets of 1965―1999. The forecast model shows a high correlation coefficient (0.79) between the model simulated and the actual typhoon frequencies in the period of 1965―1999. The forecast model also exhibits reasonable hindcasts for the typhoon frequencies for the years 2000―2006. Therefore, this work demonstrates that the new pre- dictors are significant for the prediction of the interannual variability of the WNPTF, which could be potentially used in the operational seasonal forecast of the typhoon frequency in the WNP to get a more physically based operational prediction model and higher forecast skill.     

Potential radionuclide transport pathways from seafloor dumpsites: Kamchatka region of the North Pacific Ocean

Encapsulated nuclear waste materials, dumped by Russia, are present at two deepwater seafloor locations in the offshore north-west Pacific Ocean, south-east of the Kamchatka Peninsula. This paper assesses potential pathways by which these wastes might, if released from their containers, disperse away from the dumpsites and through the surrounding ocean. A review of large-scale ocean circulation theory and of field and model results suggests that mean abyssal currents are north-eastward to eastward from the dumpsite locations and would advect leaking materials toward the north-eastern Pacific. Results of advective and diffusive horizontal plume transport models are consistent with this sense of flow. Trajectory speeds are, however, subject to considerable uncertainty. Our results suggest that as little as 5 years or as long as 100 years might be required for material to be transported from the dump sites to the north-east Pacific. Dilution by 4 or 5 orders of magnitude is predicted during this transit. Vertical mixing or upwelling are necessary in order to transport contaminants upward from north-east Pacific abyssal waters to the near-surface layers before they can potentially impact productive coastal regions, such as those off Alaska. Information concerning such upwelling mechanisms is inadequate for estimation of rates or to identify geographical areas that might be at risk.     

The Sumatra tsunami of 26 December 2004 as observed in the North Pacific and North Atlantic oceans

The M _w=9.3 megathrust earthquake of December 26, 2004 off the coast of Sumatra in the Indian Ocean generated a catastrophic tsunami that caused widespread damage in coastal areas and left more than 226,000 people dead or missing. The Sumatra tsunami was accurately recorded by a large number of tide gauges throughout the world's oceans. This paper examines the amplitudes, frequencies and wave train structure of tsunami waves recorded by tide gauges located more than 20,000 km from the source area along the Pacific and Atlantic coasts of North America.     

开边界海盆尺度环流模式模拟北太平洋CFCs分布

利用一个开边界北太平洋海盆尺度环流模式模拟了北太平洋CFCs的吸收和分布.模式计算了CFC_11海气界面通量及其在海洋中的输运和再分布,考察了开边界对模式模拟结果的影响以及开边界模式中不同沿等密度面扩散系数的影响.打开南边界可以很好地改善赤道以南海区的CFC_11模拟.增加沿等密度面扩散系数使得137°E和165°E断面模拟的高CFC_11水向南、向下输送的强度和影响范围都有明显增大,能很好地改善西北太平洋的模拟结果.西北太平洋是CFC_11的一个重要的汇区,此外在赤道东太平洋上升流区还存在一个小的汇区.     

北太平洋海表面高度的年际变化及其机制

利用15年（1993～2007年）月平均的海表面高度（SSH）异常资料,分析了北太平洋海表面高度的年际变化的时空结构,并研究了热通量和风应力两个因子对其的强迫作用.结果表明,北太平洋年际时间尺度SSH变化的大值区在黑潮延伸区和西太平洋暖池区.EOF分解第一模态的空间结构沿纬向呈带状分布,第二模态为沿经向呈带状分布.热通量强迫作用在中纬度的东北太平洋可以解释SSH年际变化40%以上.风应力对SSH的作用包括正压和斜压两个方面.正压Sverdrup平衡模型模拟的SSH年际变化较弱,仅能解释高纬度副极地环流西部的20%~40%.由大尺度风应力强迫的第一阶斜压Rossby波模型可以解释热带地区的20%~60%,中纬度中部的20%~40%,以及阿拉斯加环流东部和副极地环流西部的20%~60%.风应力强迫的一阶斜压Rossby波模型对SSH的强迫机理又可分为局地风应力强迫和西传Rossby波作用.其中,风应力的局地强迫作用（Ekman抽吸）在东北太平洋、白令海以及热带中部有显著的预报技巧,可以解释SSH年际变异的40%以上.Rossby波的传播作用在中纬度海域的副热带环流中西部和夏威夷岛以东起着重要作用,可解释20%~60%.