科尔沁沙地优势固沙灌木的生物量预测模型

灌木生物量预测模型对于快速估算固沙灌木林生产力具有重要作用。以科尔沁沙地3种常见的固沙灌木小叶锦鸡儿（Caragana microphylla）、差不嘎蒿（Artemisia halondendron）、黄柳（Salix gordejevii）为对象,基于对灌木地上、根系和整株生物量及高度、冠幅等的测定,建立3种灌木地上、根系和总生物量的估算模型,通过决定系数（R²）、估计值的标准误（SEE）和回归检验显著水平（P<0.05）筛选最优的生物量预测模型。结果表明：小叶锦鸡儿和黄柳生物量的最佳估算变量为冠幅体积,而差不嘎蒿的最佳预测变量为冠幅面积。幂函数回归模型具有最大的R²和较小的SEE,说明相对生长方程是估算小叶锦鸡儿、黄柳、差不嘎蒿灌木生物量较理想的模型。通过实测值检验,3种灌木幂函数模型预测生物量的预估精度在93%以上,具有较好的预测精度。     

Role of nurse shrubs in restoration of an arid rangeland: Effects of microclimate on grass establishment

A perennial forage grass, Agropyron desertorum was sown under the canopies of four shrub species or in open areas to test for facilitation during seedling establishment in an arid rangeland in Karnakh, Northeast Iran. Height, area and volume of shrubs were measured. Microclimate conditions and seedling establishment were assessed three times within two consecutive growing seasons. Near surface light intensity and air temperature were lower under shrubs, which led to initially higher soil moisture and grass establishment under the canopy of some shrub species. The leguminous shrub (Astragalus gossypinus) showed facilitation during moderate stress (summer 2009), but shifted to a negative effect during the severe drought (summer 2010). Competition, possibly for light, reduced the establishment of Agropyron seedlings under the cushion-like shrub (Acantholimon prostegium). Salsola arbusculiformis and Artemisia kopetdaghensis respectively showed a neutral and a facilitation effect in the first season, but a combined effect of allelopathy and drought led to the high mortality of Agropyron seedlings under their canopy. In conclusion in this arid rangeland, shrubs may facilitate establishment of understory plants under moderate drought stress, and for non-resource factors (light and temperature), but these positive effects are suppressed due to competition under severe drought conditions.     

Short-term effects of shrub control on two different plant communities in Argentina

The aim of this work was to study the effect of controlling jarilla (Larreaspp.), nonforage shrubs for livestock, on two plant communities with different forage production in the arid mid-west of Argentina. Total vegetation cover, forage species cover and carrying capacity were determined during three growing seasons, for cleared and uncleared plots. At the end of the study, significant differences were found in theLarrea cuneifoliacommunity, where average forage species cover increased by 156% over the control, and greater carrying capacity (129%) was also obtained from the cleared treatment. No significant differences were found in theLarrea divaricatacommunity. Data lead us to recommend shrub control as an improvement practice only in areas with low forage species cover and poor carrying capacity.     

Stem radial growth indicate the options of species,topography and stand management for artificial forests in the western Loess Plateau,China

An understanding of the differences in artificial forest between tree species,slope aspects,and management options in arid environments is fundamentally important for efficient management of these artificial systems;however,few studies have quantified the spatial and temporal differences in stem radial growth of trees in the arid western Loess Plateau of China.Using dendrochronology,we assessed the growth of three woody species(the native shrub Reaumuria soongorica,the exotic shrub Tamarix ramosissima and tree Platycladus orientalis)by measuring the annual stem radial increment.We also describe the long-term growth trends and responses to climatic factors on slopes with different aspects during periods with and without irrigation.We found that precipitation during the main growing season was significantly positively correlated with ring growth for all three species and both slope aspects.In addition,supplemental water(e.g.,irrigation,rainwater harvesting)greatly relieved drought stress and promoted radial growth.Our results suggest that as the main afforestation species in the Loess Plateau used for soil and water conservation,P.orientalis is more suitable than T.ramosissima under rain-fed conditions.However,a landscape that combined a tree(P.orientalis)with a shrub(R.soongorica)and grassland appears likely to represent the best means of ecological restoration in the arid western Loess Plateau.     

Desert grassland canopy arthropod species richness: temporal patterns and effects of intense, short-duration livestock grazing

Arthropods living in the canopies of two woody shrub species (a sub-shrub (Gutierrezia sarothrae) and a large shrub (Prosopis glandulosa)) and perennial grasses plus associated herbaceous species, were sampled on 18, 0.5 hectare plots in a Chihuahuan Desert grassland for five consecutive years. Mesquite shrubs were removed from nine plots, six plots were grazed by yearling cattle in August and six plots were grazed in February for the last 3 years of the 5 year study. Arthropod species richness ranged between 154 and 353 on grasses, from 120 to 266 on G. sarothrae, and from 69 to 116 on P. glandulosa. There was a significant relationship between the number of families of insects on grass and G. sarothrae and growing season rainfall but species richness was not a function of growing season rainfall on any of the plants. Several of the arthropod families that were the most species rich in this grassland were found on all of the plants sampled, i.e. Salticid spiders, Bruchid and Curculionid beetles, Cicadellid and Psyllid homopterans, and ants (Formicidae). There were more species rich families that were shared by grasses and the sub-shrub G. sarothrae than with mesquite. The absence of a relationship between growing season rainfall and species richness was attributed to variation in life history characteristics of arthropods and to the non-linear responses of annual and perennial desert grassland plants to rainfall.There were no significant differences in insect family or species richness on any of the plant types as a result of removal of mesquite (P. glandulosa) from selected plots. Intense, short duration (24 h) grazing by livestock during in late summer resulted in reduced species richness in the grass-herb vegetation layer but had no effect on insect species richness on snakeweed or mesquite shrubs. Livestock grazing in winter had no effect on insect species richness on any of the vegetation sampled.     

Gender-related differences of shrubs in stands of Atriplex canescens with different histories of grazing by cattle

Atriplex canescens is a relatively common dioecious shrub in western North America. It is considered a valuable forage resource for both wild and domestic herbivores. Sex ratios and shrub dimensions were recorded in stands of tetraploid A. canescens that had been either protected from cattle grazing or summer- or winter-grazed by cattle for at least 20 years. Stem diameter and crown shape were used as surrogates for shrub age which could not be estimated by counting growth rings. Shrub sex ratios in exclosures were significantly more female biased than the empirically derived ratio for tetraploid A. canescens (55 female:35 male: 10 monecious). Conversely, shrub sex ratios in grazed pastures were not significantly different from the empirical ratio. Proportion of female shrubs in exclosures was significantly higher than in grazed pastures. Proportion of male shrubs, on the other hand, was similar in exclosures and grazed pastures. Winter-grazed shrub stands were apparently younger than both summer-grazed and protected shrubs. Protected shrubs appeared to be the oldest. Grazed female shrubs were apparently younger than grazed males, however, shrub ages of protected male and female shrubs were apparently not different. Cattle-grazing may have affected female shrubs more negatively at this site, causing gender-based differential mortality, and/or sex-shifting. Such processes could account for the differences in sex ratios, and for the apparent gender-related differences in shrub age that were observed.     

祁连山高山灌丛生物量及其分配特征

分布于青海云杉林上线的祁连山高山灌丛是祁连山水源涵养林效益最佳林型,研究高山灌丛生物量是生态系统生产力的重要体现,也是群落结构和功能的主要测度之一。以祁连山排露沟流域3 300～3 700 m高山灌丛为研究对象,采用标准地、样方收获法以及壕沟挖掘方法,对不同海拔高度10个固定样地灌丛生物量,器官生物量进行调查,分析了不同海拔梯度灌丛的叶、枝、须根、细根、粗根烘干重数据及其生物量分配特征。结果表明：祁连山高山灌丛总生物量为12 869.39±3 306.16 kg/hm²(平均值±标准差,n=10),其中器官分配以枝生物量所占比例最高,达32.21%,叶、须根、细根、粗根的比例分别为15.70%、14.06%、11.13%和26.90%。不同海拔梯度灌丛生物量器官分配比例差异较大,地上生物量平均为6 097.17kg/hm²,地下生物量平均为6 772.22kg/hm²,不同海拔根茎比在0.56～1.93之间变化,海拔3 500 m处根茎比达最大。祁连山高山灌丛生物量与海拔呈现显著的负相关（R² = 0.898 7,p＜0.01）,随着海拔的升高,灌丛总生物量,地上以及地下生物量均呈现下降的趋势。研究结果可为内陆河流域生态环境的保护及其高山灌丛对全球气候变化的响应研究提供重要理论依据和应用资料。     

Plant Area Index and microclimate underneath shrub species from a Chilean semiarid community

In drylands, environmental conditions under shrub canopy differ from those found in open sites. We should expect that microclimate conditions under shrubs with distinct canopy architecture should also be different. Plant Area Index (PAI) of the three most abundant shrubs species (Porlieria chilensis, Adesmia bedwellii and Proustia cuneifolia) in Bosque Fray Jorge National Park, north-central Chile was measured using a Plant Canopy Analyzer. During two years (2004-2005), we recorded the Relative Humidity and Air Temperature underneath and away from the canopy of the shrubs.The three shrub species showed significant differences in PAI. Microclimate at 30 cm and 2 m above the soil in the open conditions were drier and warmer than underneath shrub canopies. Vegetation patches generate moderate microclimate conditions.Canopy structure can buffer climatic variability, contributing to high herbaceous productivity as well as shrub recruitment. Reflecting shrub architecture and observed PAI values, the lowest microclimate variations were observed under the canopies of P. chilensis, followed by P. cuneifolia and finally A. bedwellii. We bring a novel approach quantifying the Plant Area Index instead of the Plant cover and using a low cost method that integrates the distribution of leaves and may be derived from remote sensing products.     

地表径流对荒漠灌丛生境土壤水分空间特征的影响

土壤水分是干旱区多尺度生态水文过程的关键影响因素和驱动因子,其时空格局是生态、水文、气象、地形等自然过程研究的重要参数。笔者研究了降水径流事件后3种不同灌丛个体尺度土壤水分空间异质性特征,结果表明,珍珠灌丛个体尺度土壤水分空间分布特征表现为灌丛边缘>灌丛内部>灌丛间裸地,驼绒藜灌丛表现为灌丛内部>灌丛边缘>灌丛间裸地,而狭叶锦鸡儿灌丛不同微生境土壤水分差异不显著。珍珠和驼绒藜灌丛同一微生境土壤水分存在坡位梯度,珍珠灌丛3个微生境土壤水分均表现为上坡位大于下坡位,而驼绒藜灌丛边缘表现为上坡位小于下坡位,其他两个微生境无明显规律;狭叶锦鸡儿灌丛土壤水分无明显的坡位梯度。3种灌丛不同微生境土壤含水量随土层深度增加的变化不明显。这说明在降水径流事件中,不同斑块的反应差异引起了地表径流的形成以及随之发生的资源再分配,从而导致了景观内土壤水分的空间异质性。     

The effects of the identity of shrub species on the distribution and diversity of ground arthropods in a sandy desert ecosystem of northwestern China

Shrub is an important factor on structuring ground arthropod communities in desert ecosystems. In this study, in order to determine how shrubs and their species influence ground arthropod distribution patterns in a sandy desert scrubland dominated by two different shrub species, Calligonum mongolicum and Nitraria sphaerocarpa, the ground arthropods were sampled with pitfall traps during spring, summer and autumn. At the community level, total arthropod abundance was shown to be significantly higher under shrubs than in intershrub bare areas in spring; similar patterns occurred in terms of the richness of arthropod groups in the spring and over three seasons, suggesting season-specific shrub presence effects on arthropod activity. In addition, more arthropods were found under N. sphaerocarpa shrubs than under C. rnongolicum shrubs in autumn, suggesting season-specific effects of shrub species of arthropod activity, whereas more arthropods taxa were captured under C. mongoIicum than N. sphaerocarpa. At the trophic group level, the abundances of predator and herbivore arthropods were significantly greater under shrubs than in intershrub bare habitats, whereas herbivore arthropods were more abundant under N. sphaerocarpa than C. rnongolicum, and an opposite rule was detected for predator arthropods At the family level, the mean abundances of Carabidae, Curculionidae, Gnaphosidae and Lycosidae were significantly higher in the shrub microhabitats than in the intershrub bare habitat, there was no significant difference between habitats on the mean abundances of Formicidae and Tenebrionidae. The study results suggested that shrub presence and shrub species variation are important determinants of ground arthropod assemblages in this desert ecosystem, but the responses of ar- thropods differed among trophic and taxonomic groups.     

黄土高原半干旱区天然锦鸡儿灌丛对土壤水分的影响

本文选择黄土高原半干旱区定西地区的一种地带性植被,天然甘蒙锦鸡儿灌丛,将其0～99m深的土壤水分含量与人工柠条锦鸡儿灌丛、人工杏树林、天然草地、放牧荒坡和农地的土壤水分含量进行了比较。结果发现,天然锦鸡儿灌丛在1m以下土壤各层的水分含量均高于人工柠条灌丛和人工杏树林,而与放牧荒坡和农地的土壤湿度接近,略低于农地。天然锦鸡儿灌丛4m以上土层的土壤湿度还明显高于天然草地;天然锦鸡儿灌丛形成的难效—无效水层深度在2m土层以上,而人工柠条灌丛形成的难效无效水层则深达56m,人工杏树林42m,天然草地、放牧荒坡分别为36m和33m,农地1m。     

Effects of shrub removal and nitrogen addition on soil moisture in sagebrush steppe

I monitored soil moisture profiles in sagebrush steppe in SE Idaho, USA, for 6 years, a period that included four consecutive years of drought. Recharge of soil moisture was primarily the result of winter and early spring precipitation, and soil moisture declined during the spring and summer growing season. Recharge was substantially lower during drought years, and a single year of average precipitation did not restore soil moisture to pre-drought levels. Relative to control plots, there was more soil moisture on shrub removal plots and less soil moisture on nitrogen addition plots, particularly at depths of 100–180 cm. These differences suggest that in this ecosystem shrubs extract more moisture from deeper in the soil profile than perennial grasses, and that increased nitrogen availability can significantly affect soil water balance.     

Effect of plant species on shrub fertile island at an oasis–desert ecotone in the South Junggar Basin, China

Shrub fertile islands are a common feature in arid ecosystems. To examine the effect of plant species on the spatial patterns of soil chemical and physical properties surrounding individual shrubs, two deciduous shrub species with different morphologies (Tamarix spp. and Haloxylon ammodendron Bge.) were studied at an oasis–desert ecotone in South Junggar Basin. Soil samples were collected under the shrub crown (canopy), at the vertically projected limit of shrub crown margin (periphery), and in the space between shrub crowns (interspace) at two depths, 0–10 and 10–20 cm, to analyze their physical and chemical properties. The results show that the fertile islands of Tamarix spp. are enriched with more soil nutrients (significantly higher, P<0.05; soil organic matter (SOM); total nitrogen (TN) and available nitrogen (AN); to a deeper depth (>20 cm) and in a larger area (beyond the canopied area) compared to that of H. ammodendron (significantly higher, P<0.05, soil nutrients detected only for AN; <20 cm in depth; smaller than the canopied area). Soil texture patterns surrounding the shrubs of the two species are even more different, with more coarse particles under the Tamarix spp. canopies compared to the interspace between shrubs but fewer under the H. ammodendron canopies compared to the interspaces. These variations are attributed to the difference in morphology of the two studied species: the Y-shaped crowns of H. ammodendron are less capable of capturing and maintaining litter under them than the hemispheroidal crowns of Tamarix spp., which leads to the less well developed fertile islands surrounding H. ammodendron shrubs.     

Thermal utility of desert vegetation for the Egyptian tortoise and its conservation implications

We examined the monthly shrub use and the thermal utility of the desert shrub, Artemisia monosperma, by the globally endangered Egyptian tortoise, Testudo kleinmanni, in Sinai, Egypt. A. monosperma is a common desert shrub that is often used by the Egyptian tortoise as a refuge. Egyptian tortoises had the highest mean carapace temperature and lowest carapace temperature range during summer months, which coincided with the aestivation period and the Egyptian tortoise's use of larger shrubs. Our results suggest that the thermal characteristics of larger shrubs are important to the microhabitat use of the Egyptian tortoise and likely long-term survival. Larger shrubs were effective as thermal refuges that ameliorated and stabilized micro-climatic variation. Smaller shrubs were not an adequate thermal refuge in the summer because the mean temperature, and its fluctuation range beneath them was too high to offer adequate refuge during aestivation. The tortoises used the smaller shrubs in the fall, winter, and spring seasons for basking, thermoregulation, and concealment and used shrubs with a cover diameter greater than one meter during the aestivation season. The future conservation of the Egyptian tortoise should focus on protecting and restoring existing habitat by maintaining large perennial shrubs for refuge and aestivation sites.     

金沙江干热河谷不同植被坡面土壤水分时空分布特征

于2016年7~12月和2017年4月的旱、雨季期间,以金沙江干热河谷苴那小流域内的银合欢（Leucaena Benth）林地、车桑子（Dodonaea angustifolia）灌丛地和扭黄茅（Heteropogon cantortus）草地为研究对象,通过网格法和土钻法采集并测定了（0~100 cm）土层的土壤含水量,应用经典统计法和地统计学方法分析该区域不同林草植被下坡面土壤水分的动态变化特征。结果表明：（1）研究区土壤含水量总体较低,雨季显著大于旱季,旱、雨季均表现为灌丛地>草地>林地,呈中度至强度变异（0.07~0.28之间）。（2）不同林草植被下旱、雨季土壤水分具有相似的空间自相关性,自相关系数均由正向负转变,但由正向负转变的滞后距离有所不同,且雨季大于旱季,呈中等或强等空间自相关性。（3）不同林草植被下的土壤水分空间结构不同,林地、灌丛地和草地旱雨季最佳拟合模型均为球状模型;相同林草植被下各土层旱、雨季土壤水分的空间分布特征相似,但旱季的分布格局差异更显著,不同林草植被下深层土壤水分分布比表层土壤水分的分布更为复杂,土壤水分呈明显的斑块或条带状分布,含水量高值区和低值区位置不固定。总之不同林草植被类型会改变局部地段土壤水分空间分布,降雨会加强这种差异的趋势,但土壤水分仍具一定空间连续性。     

乔木和灌木枝水分传导脆弱性沿降水量递增的分化

植物枝条失去50%水分传导时对应的水势(ψ50),称为枝的水分传导脆弱性。该指标是反映植物干旱适应能力的关键指标,也决定着植物沿降水梯度的分布。乔木和灌木枝水分传导脆弱性沿降水量递增的变化是否一致,有待揭示。基于已发表的文献,筛选出236种乔木、137种灌木,建立ψ50与分布地年均降水量及分布地干旱指数关系,来确认两大类植物的ψ50沿降水递增的变化的异同。结果表明:乔木和灌木的ψ50均随降水梯度、干旱指数的增加而增加,一元线性回归方程显著;ψ50与降水梯度、干旱指数回归方程的回归系数在乔木和灌木间差异不显著,但同一降水量下,乔木比灌木具有较低的ψ50值。我们认为灌木和乔木枝水分传导脆弱性对降水量和干旱指数递增具有相似的适应性。     

POST-FIRE DEVELOPMENT OF CANOPY STRUCTURE IN A MEDITERRANEAN SHRUB,QUERCUS COCCIFERAL.

The canopy structure, including density, the apportionment of biomass in basal stem and branch diameter classes and foliage, and leaf characteristics, was examined for a dense Quercus coccifera garrigue in southern France at 3,9, and 33 years since the last fire. Indications of changing canopy structure were considered in relation to succession, regeneration strategies, water relations, and flammability. Other shrub species and herbs are most abundant at 3 and 33 years following fire, but at 9 years Q. coccifera is especially dominant. At 9 years the density of stems in the middle range of diameter classes is greatest, overall biomass reaches its peak, and leaf area index is highest. The canopy closes quickly, but by 33 years evidence for self-thinning, slowed but continuing basal sprouting, and declining leaf area and biomass is found; senescence may be beginning, but it is not an important factor in the first 33 years of canopy development. The continuity of the below-ground biomass in this and similar vegetation probably plays an important role in determining the optimum tactics for ramets through cyclic successions. Renewed competition in later years in a more open canopy of Q. coccifera, as indicated by the data, probably would be limited by recurrent fire. [Key words: biomass, canopy, density, mediterranean, physiognomy.]     

Influence of shrubs on soil characteristics and their function in Sahelian agro-ecosystems in semi-arid Niger

Soil beneath shrubs form ‘fertile islands’ in fallow sites and millet fields in semi-arid Niger. To gain more information about this phenomena different shrub species in fallow sites following a gradient from 350–650 mm precipitation were examined. For each shrub two different areas were distinguished: an area under the canopy of the shrubs and an area in the nearby open land. Soil samples were taken from a depth of 0–10 cm and analysed for C_org, N_total, P_Bray, pH(H₂O), exchangeable cations, effective cation exchange capacity (ECEC) and soil texture. Significantly higher concentrations between 38–51% for C, N, P and 22% on ECEC for K⁺were found in the soil under the shrubs. The pH showed only slight but significant differences, whereas Al³⁺and H⁺rates on ECEC under the shrubs were increased by 44–55%. For Guiera senegalensis, the most common shrub of the studied area, enrichment ratios of most soil properties increased relatively more with increasing aridity. In general, enrichment ratios decreased with the age of the fallows, whereas concentrations showed no clear evolution. The chemical composition of the shrub litter seems to influence the degree of soil enrichment. The main step of fertile island formation takes place during the cultivation period by trapping wind-blown sediment. This work shows that shrubs are of vital importance for the accumulation of nutrients and maintenance of soil fertility within agro-ecosystems of Niger.     

古尔班通古特沙漠灌木冠幅预测模型

古尔班通古特沙漠是中国第二大沙漠,也是中国固定和半固定沙丘主要分布区,固沙灌木种较多。冠幅不仅是反映固沙灌木可视化的重要参数,也是反映沙漠植被生长情况的重要变量。以3种沙丘（固定沙丘、半固定沙丘和流动沙丘）上主要固沙灌木为研究对象,利用12种基础模型、BP(Backpropagation Neural Network)神经网络和支持向量机（Support Vector Machine,SVM）机器学习算法建立了基于固沙灌木株高和冠长率的冠幅预测模型,同时将两种机器学习算法拟合结果与基础模型进行比较,最终选出了适合研究区的冠幅预测模型。结果表明：（1）不同沙丘类型和不同灌木种类的最优冠幅预测模型不同,且固定沙丘和半固定沙丘模型优于流动沙丘。3种沙丘类型最优拟合为M2(Quadratic Model)模型;（2）白梭梭（Haloxylon persicum）在半固定沙丘和流动沙丘上拟合的最优模型分别为M2、M7(Gompertz),沙拐枣（Calligonum mongolicum）最优模型为M2,蛇麻黄（Ephedra distachya）和油蒿（Artemisia ordosica）在...     

山岳旅游地生态服务价值时空分异及其与环境因子关系——以武夷山风景名胜区为例

以山岳型旅游地武夷山风景名胜区为例,估算了风景区景观生态服务价值,分析1986年、1997年、2009年风景区景观生态服务价值时空变化特征,并运用相关分析和逐步回归分析方法对生态服务价值与环境因子的关系进行了探讨,以期揭示风景区景观生态服务价值与环境因子间的作用规律。结果表明:1.武夷山风景名胜区景观类型单位面积生态服务价值最高为水体(9 3707元/(hm2.a)),最低为裸地(354元/(hm2.a)),建设用地价值(12 000元/(hm2.a))仅高于裸地、农田、灌草等景观类型,杉木林、马尾松林、竹林、阔叶林等森林景观服务价值相差不大;2.风景区内景观生态服务价值在1986—1997年间服务价值有较大损失,在1997—2009年间得到一定程度弥补,但1986-2009年间景观生态服务价值变化呈亏损趋势,植被景观类型向建设用地的转变导致景观生态服务价值的降低;3.风景区内景观类型生态服务价值与环境因子(坡向除外)间呈现中等相关性,其中,郁闭度与生态服务价值相关系数最高(0.7**),环境因子中蓄积量与郁闭度相关系数最高(0.86**);4.植被景观生态服务价拟合结果(R2=0.7524,p<0.0001)值优于非植被景观(R2=0.5370,p<0.0001),拟合效果能较好地揭示景观类型生态服务价值与环境因子间的数量关系。