The diets of ungulates from the hominid fossil-bearing site of Elandsfontein, Western Cape, South Africa

The dietary regimes of 15 ungulate species from the middle Pleistocene levels of the hominid-bearing locality of Elandsfontein, South Africa, are investigated using the mesowear technique. Previous studies, using taxonomic analogy, classified twelve of the studied species as grazers (Redunca arundinum, Hippotragus gigas, Hippotragus leucophaeus, Antidorcas recki, Homoiceras antiquus, Damaliscus aff. lunatus, Connochaetes gnou laticornutus, Rabaticerus arambourgi, Damaliscus niro, Damaliscus sp. nov., an unnamed “spiral horn” antelope and Equus capensis), one as a mixed feeder (Taurotragus oryx) and two as browsers (Tragelaphus strepsiceros and Raphicerus melanotis). Although results from mesowear analysis sustain previous dietary classifications in the majority of cases, five species were reclassified. Three species previously classified as grazers, were reclassified as mixed feeders (H. gigas, D. aff. lunatus and R. arambourgi), one previously classified as a grazer, was reclassified as a browser (the “spiral horn” antelope), and one previously classified as a mixed feeder, was reclassified as a browser (T. oryx). While current results broadly support previous reconstructions of the Elandsfontein middle Pleistocene environment as one which included a substantial C₃ grassy component, the reclassifications suggest that trees, broad-leaved bush and fynbos were probably more prominent than what was previously thought.     

Pollen records of vegetation and inferred climate change in southwestern Madhya Pradesh during the last ca. 3800 years

Pollen analysis of 1.5m deep sediment core from Sapna Lake, Betul District (M.P.) has demonstrated that between 3,800 and 2,700 yr BP, open Acacia-dominant scrub forests interspersed with stretches of herbaceous vegetation comprising grasses, sedges, Asteraceae, Cheno/Am, etc., occurred in the region under a regime of dry climate. A few trees of Madhuca indica, Holoptelea, Shorea robusta, Lagerstroemia parviflora, etc. were also sparingly distributed therein. The vicinity of the lake was under cereal-based agricultural practice as portrayed by the presence of Cerealia and other culture pollen taxa viz., Cheno/Am, Caryophyllaceae, Artemisia and Cannabis sativa. Around 2,700 to 1,260 yr BP the Acacia-dominant scrub forests were succeeded by the open mixed deciduous forests as evidenced from the improvement in frequencies of Madhuca indica, Sapotaceae, Holoptelea, Shorea robusta as well as sporadic invasion of Terminalia, Mitragyna, Flacourtia, Grewia, Lannea coromandelica, etc. This enhancement in the forest floristic reflects the initiation of a warm and humid climate possibly in response to increasing monsoon precipitation. Owing to the prevailing favourable climatic condition there was acceleration in the agricultural practice as indicated by the rising trend of Cerealia and other culture pollen taxa. Since 1,260 yr BP onwards the depletion in the prominent forest constituents such as Madhuca indica, Sapotaceae, Shorea robusta, Holoptelea and other associated trees implies that the forests turned more sparse and less diversified due to the inception of a warm and less humid climate, attributable to reduced monsoon precipitation. However, the agricultural practice continued with almost same intensity as before, because the Cerealia and other culture pollen taxa remain unchanged.     

Methyl ketones in high altitude Ecuadorian Andosols confirm excellent conservation of plant-specific n-alkane patterns

To reconstruct past shifts in the upper forest line (UFL) in the Northern Ecuadorian Andes we are studying the applicability of plant-specific patterns of lipids preserved in soils as proxies for past vegetation along an altitudinal transect. Longer chain length n-alkanes, (C₁₉–C₃₅) were previously found to occur in plant-specific patterns in the dominant vegetation in the area as well as in preliminary soil samples, and may serve as such a proxy. In the present study, we assessed the preservation of n-alkane patterns with depth in soils from five excavations along an altitudinal transect 3500–3860 m above sea level (m.a.s.l) in the area. We used the carbon preference index (CPI) as well as chain length distributions of n-alkanes and their most likely degradation products, n-methyl (Me) ketones, n-alcohols and n-fatty acids. Clear n-alkane patterns were found in all the soils and at all depths, while a clear relationship with the observed patterns of n-Me ketones identified them as the primary degradation product of the former. Very low average n-Me ketone/n-alkane ratio values were found, ranging from 0.03 to 0.15 at the top of the mineral soil, to 0.05–0.20 at the interface with an underlying palaeosol several thousand years old. The concurrent high CPI values indicate very limited degradation of n-alkanes with depth. Except for C₃₃, the shifts in n-Me ketone/n-alkane values were similar for all chain lengths investigated, signifying an absence of preferential degradation of individual n-alkanes. With one exception, all the soils showed a similar increase in n-Me ketone/n-alkane values with depth, indicating that the degradation rates were not influenced by altitude. This means that, even if the total concentration of n-alkanes decreases over time, the characteristic pattern remains intact, conserving their potential as a biomarker for past vegetation reconstruction in the area, as well as for investigation of degradation processes of soil organic carbon.     

Early Barremian orbitolinid record from the Moghan area,NW Iran: Northern margin of the Neotethys

Orbitolinid foraminifers are reported for the first time from lower Barremian shallow marine carbonate deposits of the Moghan area, NW Iran. According to geologic map of the Razi, these rocks were previously assigned to general age of the Early Cretaceous. The early Barremian age is documented based on stratigraphic range of the marker Valserina turbinata (Foury). Other orbitolinids such as Eopalorbitolina charollaisi Schroeder, Eopalorbitolina pertenuis (Foury), Paleodictyoconus cf. cuvillieri (Foury), Montseciella cf. alguerensis Cherchi and Schroeder, Paracoskinolina cf. maynci (Chevalier), Orbitolinopsis cf. buccifer Arnaud-Vanneau and Thieuloy, and Dictyoconus? pachymarginalis Schroeder also coexist. Obtained biostratigraphic data suggest that the range of Dictyoconus? pachymarginalis, hitherto known from the Aptian, has to be extended into the early Barremian. Most of the orbitolinid taxa are well known in Europe as northern Tethyan margin endemic forms. Therefore, the study area can be considered as part of the northern Tethys margin during the Barremian. This study also extends the palaeobiogeographic distribution of northern Tethyan orbitolinids eastwards as far as northwest Iran.     

Turonian cephalopods (ammonoids and a nautiloid) from the Wadi Daya Formation of the Talerzha Basin (South Riffian Ridges Domain,northern Morocco)

The Wadi Daya Formation, or the Calcaires crayeux of the older literature, attains a thickness of 10–40 m in the Talerhza Basin of the South Riffian Ridges. Previously, this unit was first dated as “Vraconian” (i.e., late upper Albian), but then reinterpreted as Cenomanian-Turonian and Cenomanian-Coniacian on the basis of foraminiferal and ostracod assemblages, respectively. Here, we record for the first time in the South Riffian Ridges, some typically Turonian ammonoids and a nautiloid species, namely Romaniceras (Yubariceras) cf. ornatissimum (Stoliczka), Spathites (Jeanrogericeras) cf. reveliereanus (Courtiller), Neoptychites cephalotus (Courtiller), Pachydesmoceras linderi (de Grossouvre), Lewesiceras peramplum (Mantell) and Angulithes galea (Fritsch, in Fritsch & Schlönbach). These species are herein described and illustrated. In view of these data, the underlying Marnes et marno-calcaires jaunes Formation, formerly dated as “Vraconian”, could in fact be of a middle to late Cenomanian date, in accordance with the age assignment based on planktonic foraminifera. Deposition of the overlying Marnes jaunes Formation, previously dated as Cenomanian-“Senonian”, probably started during the latest Turonian or earliest Coniacian.     

Germanium crystal chemistry in hematite and goethite from the Apex Mine,Utah, and some new data on germanium in aqueous solution and in stottite

At the Apex Mine in southwest Utah, fine-grained hematite contains as much as 1.0 wt. percent Ge, and fine-grained goethite contains as much as 0.7 wt. percent Ge. The mode of Ge incorporation in these minerals was investigated by high-resolution K-edge fluorescence spectroscopy using synchrotron radiation. Analysis of extended fine structure (EXAFS) K-edge data for Ge and Fe shows that Ge substitutes for Fe in the octahedral metal sites of the studied hematite and goethite, with average Ge-ligand bond lengths of 1.88 Å. The solid solution in hematite probably occurs through the coupled substitution 2Fe(III) = Ge(IV) + Fe(II), similar to the coupled substitution 2Fe(III) = Ti(IV) + Fe(II) that occurs in the solid solution series hematite-ilmenite. The solid solution in goethite probably occurs by the loss of an H atom from an OH group, through the coupled substitution Fe(III) + H(I) = Ge(IV). In related experiments, EXAFS data indicate that in a neutral aqueous solution containing 790 ppm Ge, the Ge occurs predominately as Ge(OH)₄, with tetrahedral Ge-OH bond lengths of 1.74 Å. In stottite, FeGe(OH)₆, Fe(II) and Ge(IV) occur in octahedral sites with average Fe-OH and Ge-OH bond lengths of about 2.20 Å and 1.88 Å.     

New palaeontological evidence suggests an early Middle Pleistocene age for the lower levels of Sun Hole Cave,Cheddar, Somerset,UK

Sun Hole, a small fissure cave on the north side of Cheddar Gorge, Somerset, is best known as the site where late Upper Palaeolithic artefacts are associated with human and other mammal remains that immediately pre-date the Lateglacial interstadial. These remains from the upper levels (Unit I) overlie sediments (Unit II) that are thought to have accumulated during a full glacial period, below which are sediments (Unit III) attributed to an interglacial stage. The extinct land snail Retinella (Lyrodiscus) sp., which in Britain was only known from the Hoxnian Stage, had previously been recovered from Unit III. The interglacial at the base of the sequence was therefore assigned to the Hoxnian Stage (Marine Isotope Stage (MIS) 11, 424–374 kyr). Here we provide additional analyses of samples from Unit III, which as well as containing further Retinella (Lyrodiscus), also yielded a specimen of Monachoides incarnatus, its first record from the British Pleistocene. An associated assemblage of small mammals included a few that are rare in the British Pleistocene, such as birch mouse (Sicista cf. betulina) and garden dormouse (Eliomys quercinus). The vole Lasiopodomys gregaloides and the shrew Sorex (Drepanosorex) savini do not occur as late as the Hoxnian and therefore suggest an older age in the early Middle Pleistocene. The record from Sun Hole therefore has parallels with the well-known sequence from the upper Calcareous Member in the nearby cavern system at Westbury-sub-Mendip.     

The influence of morphology in determining the decomposition of two salt marsh macrophytes

Annual decomposition rates of Spartina alterniflora height forms and Juncus roemerianus were determined in situ in three North Carolina salt marshes using the litter bag method. The decomposition of Spartina was significantly influenced by size, i.e., height form, with the taller plants which had greater amounts of stem tissue, being more resistant to decay. Instantaneous decay rates for short and medium Spartina were not significantly different at any site, but they were both significantly greater than that of the tall form at two of the three study sites. Juncus decomposed more slowly than Spartina during the first 8 months, but had decomposed as completely as all three height forms of Spartina at two of the study sites by the end of the 13-month study period. Constant submergence appeared to inhibit decomposition since there was twice as much undecomposed plant material remaining in bags placed in tidal creeks as in those on the marsh surface.     

The distribution of calcareous nannofossils and foraminifers in the Callovian, Oxfordian, and Volgian deposits in the southwest of Moscow

The distribution of calcareous nannofossils and foraminifers occurring in the Callovian-Oxfordian deposits in the southwest of Moscow is studied. Nannoplankton-bearing beds and foraminiferal zones are distinguished. The Retecapsa incompta Beds correspond in range to the Ophthalmidium sagittum-Epistomina volgensis and Ophthalmidium strumosum-Lenticulina brestica foraminiferal zones as well as the lower part of Epistomina uhligi-Lenticulina russiensis Zone. The Watznaueria manivitae, Crepidolithus perforata, and Watznaueria fossacincta (lowermost part) beds span interval of the Epistomina uhligi-Lenticulina russiensis Zone. The Watznaueria fossacincta Beds are concurrent to the Lenticulina ponderosa-Flabellamina lidiae Zone of the foraminiferal scale.     

The earliest Attagenus species (Coleoptera: Dermestidae: Attageninae) from Upper Cretaceous Burmese amber

A new species belonging to the extant dermestid genus Attagenus, Attagenus burmiticus sp. nov., is described and illustrated based on a well-preserved specimen from the Upper Cretaceous Burmese amber. The discovery suggests that Attagenus is an ancient group, originating as early as in the mid-Cretaceous. Along with another species of Attagenus known from the Upper Cretaceous New Jersey amber, it implies that Attagenus were widespread in the Mesoozic.     

Upper Cretaceous amber from Vendée,north-western France: Age dating and geological,chemical, and palaeontological characteristics

The Upper Cretaceous lignite deposits of La Garnache, Vendée (north-western France), consist of two lignitic clay series, Garnache 1 and Garnache 2, separated by a fault. The first series cropped out to the south of the fault during road works until 2002 but is now covered by an embankment. It has provided numerous pieces of amber containing arthropod and micro-organism inclusions. The second lignitic series, exposed to the north of the fault, is rich in fossil wood but devoid of amber. Palynological analysis of Garnache 1 revealed several Normapolles species belonging to the genera Atlantopollis, Complexiopollis, Osculapollis, Plicapollis and Trudopollis, but larger forms typical of Senonian deposits are absent. By contrast, Garnache 2 proved to be dominated taxonomically and numerically by spores (Appendicisporites, Camarozonosporites, Gleicheniidites, Patellasporites, Stereisporites), associated with a few gymnosperm (Cerebropollenites, Phyllocladidites, Classopollis) and angiosperm (Liliacidites, Retitricolpites and a single specimen of the Normapolles Complexiopollis) taxa. Garnache 1 is, therefore, younger than Garnache 2, the latter being clearly Cenomanian in age whereas Garnache 1 is more likely to be Turonian. Lignitic clay of Garnache 1 contains numerous translucent, orange to red, pieces of amber. Vendean amber is rich in aquatic arthropods, such as tanaids and epicarideans (Crustacea), as well as marine or brackish siliceous micro-organisms such as diatoms and sponge spicules. These aquatic inclusions indicate that resin-producing trees grew along and close to the seashore. The amber-bearing clay was deposited in a calm, estuarine or lagoonal, muddy environment.     

Distribution of the dentary groove of theropod dinosaurs: Implications for theropod phylogeny and the validity of the genus Nanotyrannus Bakker et al., 1988

This study examines the phylogenetic distribution of a morphologic character, described as a groove containing pores, on the lateral surface of the dentary bone in theropod dinosaurs. The nature of this groove is a feature unique to theropods. Of the 92 theropod taxa examined for the presence and absence of this feature, 48 possessed and 44 lacked this feature. Distribution of this character was compared to published phylogenetic analyses of theropods, in order to evaluate the utility of the dentary groove as a diagnostic feature. 80% of pre-Tyrannoraptoran theropods possessed the dentary groove, with only 6 reversals in basal theropod clades. Theropods with beaks or edentulous jaws all lacked a dentary groove. Tyrannosauroidea is marked by mosaic distribution of this character. Among tyrannosauroids, the dentary groove occurs only in Dryptosaurus and the Albertosaurinae (Albertosaurus + Gorgosaurus). Nanotyrannus lancensis, sometimes described as representing juvenile Tyrannosaurus rex, also possesses this groove, unlike the remainder of the Tyrannosaurinae. Nanotyrannus lancensis was included in a phylogenetic analysis of Tyrannosauroidea and was recovered within Albertosaurinae. We recommend that Nanotyrannus stand as a valid taxon nested within the Albertosaurinae, based on the presence of this groove, as well as other features of the skull.     

Saurischian track assemblages from the Lower Cretaceous Shenhuangshan Formation in the Yuanma Basin,Southern China

Cretaceous dinosaur tracks from Hunan Province are historically significant as the basis for three ichnotaxa: Xiangxipus chenxiensis, Xiangxipus youngi, and Hunanpus jiuquwanensis all representing theropodan tracks, described from a single site in 1982. Although the type locality has since been destroyed, the type specimen and replicas remained available for restudy in 2006, when paratype Hunanpus tracks and sauropod footprints were described from a second, nearby locality. Material from both localities is here re-described in detail. It is proposed that while Xiangxipus chenxiensis can be regarded as a distinct ichnospecies, probably representing an ornithomimid trackmaker, Xiangxipus youngi cannot be accommodated in the same ichnogenus. We consider it similar to the ichnogenus Wupus from the Lower Cretaceous of Sichuan Province, and therefore of possible avian affinity. We also find no justification for regarding Hunanpus jiuquwanensis as generically distinct from the widespread ichnogenus Grallator, and therefore rename it Grallator jiuquwanensis comb. nov. The Hunan track record is generally consistent with the sparse record of theropod skeletal remains in the province, but adds evidence of sauropods that was previously lacking.     

Leymeriellidae (Cretaceous ammonites) from the lower Albian of Esfahan and Khur (Central Iran)

Leymeriellid ammonite faunas are described from northeast of Esfahan and the Khur area (Central Iran). The faunas comprise Leymeriella (L.) tardefurcata, L. (L.) germanica and L. (L.) acuticostata? as well as L. (Neoleymeriella) regularis, L. (N.) diabola and L. (Neoleymeriella) pseudoregularis?. The leymeriellids of Iran are closely related to faunas from northwest Europe, the Vocontian Basin and Transcaspia (Mangyschlak) and thus allow for biostratigraphic correlations, indicating the earliest Albian Leymeriella tardefurcata Zone with its superimposed L. acuticostata and L. regularis subzones. We suggest that the sudden appearance of common representatives of the genus Leymeriella in the northern and Central Iranian basins (Leymeriella acme) can be used as a useful proxy marker to define the base of the Albian Stage in Iran.     

New data on the palaeobiogeography of Loftusia genus (Foraminiferida). An in situ presence of the genus in eastern Greece (Boeotia)

An Upper Maastrichtian horizon rich in Loftusia is for the first time described in situ in Greece in the province of Boeotia. It is found in a continuous undisturbed carbonate sequence of the eastern Greece platform (Subpelagonian zone) of Maastrichtian–Paleocene age followed by flysch sedimentation. Loftusia is found in a facies reflecting an outer shelf environment, associated with debris of rudists, Orbitoides spp., Siderolites calcitrapoides, Omphalocyclus macroporus, Hellenocyclina beotica, Sulcoperculina sp., and echinoderms. Similar fauna, but without Loftusia, is found in the surrounding levels of Late Maastrichtian age as well. This recovery of the genus in the western part of its distribution area, where it is not as abundant as in the eastern part, is considered significant for the palaeobiogeography of the genus in Tethys Ocean during Late Cretaceous. To cite this article: A. Zambetakis-Lekkas, A. Kemeridou, C. R. Geoscience 338 (2006).     

The rudist bivalve Pironaea milovanovici Kühn, 1935, a multiple-fold Hippuritidae,from south-eastern Spain. Taxonomic implications. Pore and canal system constructional morphology

The study of the external and internal shell characters of Pironaea milovanovici Kühn, as well as of their ontogenetic and intra-specific variability, observed on abundant, well preserved, and matrix-free specimens from Tavernes de la Valldigna, Valencia province, Spain, revealed that other Pironaea species (e.g. P. bacevicensis Milovanović, P. petkovici Milovanović, Sladić-Trifunović, and Grubić, P. postdalmatinica Pamouktchiev, P. signumpraeverta Pamouktchiev, and Pironaea sp. ex gr. polystyla in Lupu), based on one or a very small number of specimens, are junior synonyms of P. milovanovici Kühn. The pore and canal system of the left valve is described in detail, based on differently weathered valves and differently oriented sections; it is constrained by the presence of secondary pillars in both valves. Being the pore and canal system and its relation with the secondary pillars relatively simple and well understood in P. milovanovici, by comparison, an interpretation of these structures in more complex multiple-fold Hippuritidae such as those of the Americas is hypothesized.     

http://www.sciencedirect.com/science/article/pii/S167498711300090X

The Khami group within the petroliferous Zagros Basin of Iran includes the Surmeh, Hith, Fahliyan, Gadvan and Dariyan formations. The Dariyan Formation, previously known as Orbitolina limestone is very signif-icant as a reservoir rock in the Zagros Basin. In this study, the age of the Dariyan Formation is examined based on biostratigraphic and paleontological investigations. 236 m of Dariyan Formation sediments in the sections at Kuh-e Rahmat （north） and Nour-Abad （west） of Iran were selected to study its biostratigraphy. According to the lithostratigraphic investigations, the Dariyan Formation in these sections is subdivided into three： lower, middle and upper, which include thin-bedded to medium-bedded and massive lime-stone. Micropaleontologic study of the Dariyan Formation has led to the recognition of two biozones in the Nour-Abad area and one biozone in the Kuh-e Rahmat stratigraphic column. Diagnostic foraminifera in these sections are as follows： Globigerinel oides algerianus Cushman and Ten Dam, Globigerinelloides fer-reolensis Moullade, Globigerinelloides maridalensis （Bolli）, Globigerinelloides paragottisi Verga and Permoli Silva, Hedbergel a aptiana Bartenstein, Hedbergel a excelsa Longoria, Hedbergel a luterbacheri Longoria, Hedbergella occulta Longoria, Hedbergella ruka （Banner et al.）, Hedbergella similis Longoria, Hedbergella trocoidea （Gandolfi）, Muricohedbergella delrioensis （Carsey）, Pseudoplanomalina cheniourensis Sigal, Con-icorbitolina conica d＇Archiac, Iraqia sp., Mesorbitolina parva Douglass, Mesorbitolina pervia Douglass, Meso-rbitolina ovalis Gorog and Arnaud-Vanneau, Mesorbitolina subconcava Leymerie, Mesorbitolina texana （Roemer）, Mesorbitolina ovalis-pervia Cherchi and Schroeder, Debarina hahounerensis Fourcade et al., Marssonel a trochus （d＇Orbigny）, Nezzazata sp., Orbitolina sp., Textularia sp.     

Climatic controls of the middle marsh zone in the Bay of Fundy

The ecological importance of Plantago maritima within a salt marsh on the Bay of Fundy is documented through measurements of cover, density, and biomass. During late August 1993, peak standing crops of Plantago were as high as 532 g m^?2, and composed as much as 96% of the biomass of a stand of vegetation. Plantago is a dominant component of the marsh vegetation at an elevation just above the Spartina alterniflora-dominated low marsh, and is found as a dominant when growing in association with a number of plant species characteristic of the high marsh. We hypothesize that the existence of this community is dependent upon regular ice-shearing of Spartina patens, which would otherwise competitively exclude Plantago. This hypothesis is supported by the elevational limits of Plantago dominance and the geographical limitation of Plantago communities to portions of the northwestern Atlantic subjected to winter temperatures which average below 0°C.     

Pachycephalosaurs (Dinosauria: Ornithischia) from the Upper Cretaceous (upper Campanian) of New Mexico: A reassessment of Stegoceras novomexicanum

Pachycephalosaurs, a group of ornithischian dinosaurs with distinctive cranial ornamentation and skull domes, underwent dramatic changes in cranial morphology during ontogeny. This has caused debate about whether some specimens belong to juveniles or adults, which impacts studies of pachycephalosaur phylogeny and evolution. One such debate concerns a small skull roof specimen from the Campanian (Upper Cretaceous) of New Mexico, NMMNH P-33898, which was originally described as an indeterminate juvenile but later regarded as a mature adult and erected as the holotype of a new small-bodied species, Stegoceras novomexicanum. We restudied NMMNH P-33898 using computed tomography scanning, morphometric and phylogenetic analyses, and comparisons to growth series of other pachycephalosaurs (Stegoceras validum, Pachycephalosaurus wyomingensis). We conclude that two purported paratype specimens of Stegoceras novomexicanum cannot be referred to the same taxon as the holotype, that the holotype and controversial paratypes all belong to immature specimens and not aberrant small-bodied adults, but that current evidence cannot clearly determine whether NMMNH P-33898 is a juvenile belonging to its own diagnostic species (S. novomexicanum) or is a juvenile of Stegoceras validum, Sphaerotholus goodwini, or another known taxon. We review the pachycephalosaur record of New Mexico and demonstrate that pachycephalosaurs were important components of dinosaur faunas in the southern part of Western North America during the ∼15 million years before the end-Cretaceous extinction, just as they were in roughly contemporaneous northern localities.