植物始花期对气候变化响应的激素调控机理研究进展

植物物候对气候变化的响应非常敏感,是指示全球变化对生态系统影响的重要证据。其中植物花期变化影响植物繁殖与进化,具有重要意义。大量研究表明,随着全球气候变暖,北半球植物始花期普遍提前。而关于气候变暖对植物始花期影响的内在机理并没有明确解释。植物激素是植物体内对植物开花等生理活动有显著调控作用的有机物,可对环境刺激作出响应并直接参与调控植物始花期,导致始花期的提前或延后。对植物激素在植物中表达与变化的精确测定有助于了解植物始花期对气候变化响应的内在机理。论文综述了植物激素对植物开花时间的调控作用以及目前植物激素最先进的检测方法,探讨了植物激素及相关代谢产物在植物体内的含量变化对植物始花期的影响;提出通过植物激素研究植物始花期对气候变化响应的激素调控机理,为探索植物物候对气候变化响应提供新的研究思路与手段。     

Seasonal changes in the relationship between species richness and community biomass in grassland under grazing and exclosure, Horqin Sandy Land, northern China

How species diversity–productivity relationships respond to temporal dynamics and land use is still not clear in semi-arid grassland ecosystems. We analyzed seasonal changes of the relationships between vegetation cover, plant density, species richness, and aboveground biomass in grasslands under grazing and exclosure in the Horqin Sandy Land of northern China. Our results showed that in grazed and fenced grassland, vegetation cover, richness, and biomass were lower in April than in August, whereas plant density showed a reverse trend. Vegetation cover during the growing season and biomass in June and August were higher in fenced grassland than in grazed grassland, whereas plant density in April and June was lower in fenced grassland than in grazed grassland. A negative relationship between species richness and biomass was found in August in fenced grassland, and in grazed grassland the relationship between plant density and biomass changed from positive in April to negative in August. The relationship between the density of the dominant plant species and the total biomass also varied with seasonal changes and land use (grazing and exclosure). These results suggest that long-term grazing, seasonal changes, and their interaction significantly influence vegetation cover, plant density, and biomass in grasslands. Plant species competition in fenced grassland results in seasonal changes of the relationship between species richness and biomass. Long-term grazing also affects seasonal changes of the density and biomass of dominant plant species, which further affects the seasonal relationship between plant density and biomass in grasslands. Our study demonstrates the importance of temporal dynamics and land use in understanding the relationship between species richness and ecosystem function.     

A review of snow manipulation experiments in Arctic and alpine tundra ecosystems

Snow cover is one of the most important factors controlling microclimate and plant growing conditions for Arctic and alpine ecosystems. Climate change is altering snowfall regimes, which in turn influences snow cover and ultimately tundra plant communities. The interest in winter climate change and the number of experiments exploring the responses of alpine and Arctic ecosystems to changes in snow cover have been growing in recent years, but their outcomes are difficult to summarize because of the large variability in manipulation approaches, extents and measured response variables. In this review, we (1) compile the ecological publications on snow manipulation experiments, (2) classify the studies according to the climate scenarios they simulate and response variables they measure, (3) discuss the methods applied to manipulate snow cover, and (4) analyse and generalize the response in phenology, productivity and community composition by means of a meta-analysis. This meta-analysis shows that flowering phenology responded strongly to changes in the timing of snowmelt. The least responsive group of species were graminoids; however, they did show a decrease in productivity and abundance with experimentally increased snow covers. The species group with the greatest phenological response to snowmelt changes were the dwarf shrubs. Their abundance also increased in most long-term snow fence experiments, whereas species richness generally declined. We conclude that snow manipulation experiments can improve our understanding of recently observed ecosystem changes, and are an important component of climate change research.     

Reproductive phenology of Isolatocereus dumortieri (Cactaceae) in semiarid scrub in central Mexico: Effect of rain during the dry season

The giant cactus Isolatocereus dumortieri is a dominant species of the semiarid scrub of central Mexico. Its reproductive period is during the dry season, and it produces essential resources (pollen, nectar and fruits) for a great variety of animal species. We related the production of reproductive structures with water variables in the soil–plant system. The main goal of the present study was to evaluate the consequences of a change in rainfall pattern on the phenology of I. dumortieri. We watered some plants in the dry season to simulate heavy rain events, and to test the hypothesis that water availability during the reproductive season has a negative effect on fruiting and a positive effect on vegetative growth. The seasonality of rain events caused variations in soil water potential and plant osmotic potential, and we found that both variables influenced fruit production. The regression models relating the number of fruits with soil water potential, rainfall and osmotic potential were significant for all three study periods. The highest production of reproductive structures occurred in the driest year (2009), during which there was an ENSO event. Watering did not have a significant effect on osmotic potential or growth in the cacti. However, the watering × time interaction had a negative effect on the number of immature fruits. That is, at the end of the experiment, the plants that received the most water showed a decrease in the number of fruits. Plant growth during the rainy period was significantly greater than during the dry period. The results support the hypothesis that a change in rainfall pattern during the dry season has an effect on fruit production. Vegetative growth, however, occurred only during the wet season.     

Above and belowground phenology of four Mediterranean sub-shrubs. Preliminary results on root–shoot competition

Above and belowground phenology are critical aspects for plant life in areas of seasonal climate like Mediterranean regions. However, fine root growth phenology is rarely considered in most phenological studies. In this article we describe the above and belowground phenology of four species of Mediterranean sub-shrubs growing along a gradient of temperature and water availability, with special attention at the relationship between both processes. Observations were conducted monthly over a minimum of 12 months per species. Fine root growth varied significantly throughout the year, being higher in autumn than in spring and minimum in summer. In the species growing in cold areas, root growth was also reduced during winter. Shoot growth was maximum in spring for the four study species, buts its beginning was related to the site temperature, being earlier in those species growing in warmer sites. The species displaying a short vegetative period tended to separate root and shoot growth processes throughout the year. These results emphasize the importance of water availability and winter cold on fine root growth. A trade-off appears to exist between the duration of the vegetative period and the overlap between root and shoot growth processes.     

Influences of land use history and short-term nitrogen addition on community structure in temperate grasslands

Better understanding of plant community structure in relationship to land use history and nitrogen (N) will facilitate grassland conservation and projections of community succession under future N deposition. We conducted a field experiment in northern China to examine the influence of 2-year N addition on species composition and community structure in three temperate grasslands with different land use history: steppe fenced for two years (ST), steppe fenced for five years (SF), and old field grassland fenced for five years (OF). The results showed previous farming reduced species diversity and plant cover, which is mainly caused by decrease in perennial grasses (PG) and forbs (PF). Nitrogen addition increased plant cover, especially the PG cover, but had little effects on species diversity. Nitrogen enrichment also has a tendency to alter community composition by decreasing proportional cover of PF but increasing that of PG and annuals and biennials (AB), suggesting enhanced dominance of PG under increasing N deposition. Irrespective of the short-term (2 years) experimental periods, our findings highlight the dominant role of land use history in structuring plant community, and have valuable implications for grassland conservation and model projections of ecosystem succession under global scenarios of N deposition in the semi-arid grasslands.     

基于MODIS影像的土地覆被分类研究——以京津冀地区为例

在全球变化研究中,如何快速、准确获取土地覆被信息对该项研究有着至关重要的作用.随着遥感科学的不断发展和应用领域的深入,研究者可以利用遥感影像进行土地覆被分类研究,并且具有准确、快速、自动化等优点.本文利用MODIS数据具有的多光谱、多时相特点,以京津冀地区为例,选取2013 年全年16-day 的MOD13Q1/EVI时间序列数据、2013 年5 月份一期的MOD09Q1(1、2 波段)和MOD09A1(3-7 波段)产品,并运用时间序列谐波分析法对全年MOD13Q1/EVI 时间序列数据进行去云、去噪的平滑重建处理,使其数据更能反映物候周期性变化规律.选择谐波分析后的全年MOD13Q1/EVI 时间序列数据、MODIS数据的1-7 波段地表反射率和NDWI(归一化差异水体指数)、MNDWI(改进归一化差异水体指数)和NDSI(土壤亮度指数),构建了3 种特征变量组合方案的CART决策树,分别进行京津冀地区的土地覆被分类研究.结果表明:方案一(全年EVI 的23 个时相)、方案二(方案一+MOD09 的1-7 波段地表反射率)和方案三(方案二+MNDWI+NDSI+NDWI)的总体分类精度分别达到86.70%、89.98%、91.34%,Kappa系数分别为84.94%、88.66%、90.20%.研究表明,仅利用MODIS遥感影像自身多种分类特征和决策树方法对宏观土地覆被分类就可达到较高精度,显示了本文分类方法在实践中的可行性及MODIS数据在区域尺度土地覆被分类研究方面的优势与潜力.     

青藏高原维管植物物种丰富度分布的情景模拟

如何充分利用离散的观测数据,通过对维管植物物种分布丰富度及其与生境因子之间的相互作用和影响机理的定量分析,实现维管植物物种丰富度的空间分布及其情景模拟,是目前生物多样性研究前沿和核心内容之一。针对这一问题,在实现青藏高原37个国家自然保护区的维管植物物种数量收集和边界数据矢量化的基础上,分别进行维管植物物种数量与土地覆盖类型、环境因子和景观生态指数等三大类生境因子之间的相关关系的定量计算和对比分析,筛选和确定最佳相关分析方程,进而构建青藏高原维管植物物种丰富度的空间模拟分析模型。该模型中,维管植物物种丰富度与生境因子之间的复相关系数为0.94,模型验证结果表明,青藏高原的维管植物物种的平均丰富度为496.79种/100 km²,其空间分布格局整体上呈东南向西北逐渐减少趋势;另外,除柴达木盆地荒漠区域以外,维管植物物种的空间分布随海拔的升高而减少。基于CMIP5 RCP 2.6、RCP 4.5和RCP 8.5三种气候情景模拟获得的青藏高原维管植物物种丰富度未来情景结果显示,在T0-T4（2010-2100）时段内,青藏高原维管植物物种丰富度整体将呈减少趋势。RCP 8.5情景下青藏高原维管植物物种丰富度的变化幅度最大,而RCP 2.6情景下的维管植物物种丰富度的变化幅度最小。研究表明,本文构建的模型能够对青藏高原维管植物物种丰富度的空间分布格局及其未来情景进行模拟分析,模拟结果可为青藏高原生物多样性及其对气候变化响应的综合评估和情景模拟提供方法和技术支持。     

陆地生态系统模拟研究中IBIS模型开发应用的回顾和展望

全球动态植被模型(DGVM)是研究生态系统复杂过程和全球变化的强有力工具。本文基于过去20多年全球已发表的文献,对得到广泛关注和应用的DGVM之一——集成生物圈模拟器(Integrated Biosphere Simulator,IBIS)的开发、改进、发展及应用进行了总结。IBIS是一个在全球变化等多领域中有着广泛应用的模型。自1996年诞生以来,IBIS在陆地生态系统的碳、氮、水循环,植被动态、陆气耦合、水域系统耦合和气候变化影响等多个方面取得了验证和应用。本文较为系统地阐述了IBIS模型在V2.5版本后的不同发展方向,主要针对IBIS模型在水文过程(蒸散、土壤水分、地下水、径流)、植被动态(植被功能型、土地覆盖变化)、植被生理过程(植被物候、光合作用、植被生长、碳分配)、土壤生物地球化学过程(土壤碳氮循环及反馈、温室气体排放等)以及包括土地利用变化、干扰与管理等人类活动过程等方面的改进与发展进行了全面的综述;在此基础上,对模型在生态系统生产力、碳水收支、水分利用效率、温室气体排放、自然干扰(干旱、火灾、虫害)和人类活动(土地利用变化、农业经营)等方面的应用,以及模型的技术性改进方面进行了回顾;最后对模型的前景和进一步发展提出了一些见解。     

气候变化背景下1981-2010年中国玉米物候变化时空分异（英文）

Spatio-temporal changes in the differentiation characteristics of eight consecutive phenological periods and their corresponding lengths were quantitatively analyzed based on long-term phenological observation data from 114 agro-meteorological stations in four maize growing zones in China. Results showed that average air temperature and growing degree-days (GDD) during maize growing seasons showed an increasing trend from 1981 to 2010, while precipitation and sunshine duration showed a decreasing trend. Maize phenology has significantly changed under climate change: spring maize phenology was mainly advanced, especially in northwest and southwest maize zones, while summer and spring-summer maize phenology was delayed. The delay trend observed for summer maize in the northwest maize zone was more pronounced than in the Huang-Huai spring-summer maize zone. Variations in maize phenology changed the corresponding growth stages length: the vegetative growth period (days from sowing date to tasseling date) was generally shortened in spring, summer, and spring-summer maize, although to different degrees, while the reproductive growth period (days from tasseling date to mature date) showed an extension trend. The entire growth period(days from sowing date to mature date) of spring maize was extended, but the entire growth periods of summer and spring-summer maize were shortened.     

西藏纳木错植物物候及其对气候的响应

利用纳木错流域1O种代表性植物2007年和2008年的物候观测资料,分析了植物生殖物候的基本特征及气候条件对其的影响.植物最早5月中旬进入开花期,大部分植物的盛花期在6月中旬至7月下旬,开花的最适温度范围为8.5-8.9℃,花期延续到9月下旬.从6月上旬开始,大部分植物进入果期,多数植物果期与花期重叠.纳木错具有高寒地带植物物候的普遍特征:生长期较短(5个月左右)、早花现象、晚花现象.同2007年相比,2008年雨季提前,虽然气温偏低,大部分植物花期和果期普遍缩短5 d左右,但物候期提前约20 d.植物物候的年际变化对气候变化,特别是对降水量的季节分配更敏感.     

农业物候动态对地表生物物理过程及气候的反馈研究进展（英文）

Response and feedback of land surface process to climate change is one of the research priorities in the field of geoscience. The current study paid more attention to the impacts of global change on land surface process, but the feedback of land surface process to climate change has been poorly understood. It is becoming more and more meaningful under the framework of Earth system science to understand systematically the relationships between agricultural phenology dynamic and biophysical process, as well as the feedback on climate. In this paper, we summarized the research progress in this field, including the fact of agricultural phenology change, parameterization of phenology dynamic in land surface progress model, the influence of agricultural phenology dynamic on biophysical process, as well as its feedback on climate. The results showed that the agriculture phenophase, represented by the key phenological phases such as sowing, flowering and maturity, had shifted significantly due to the impacts of climate change and agronomic management. The digital expressions of land surface dynamic process, as well as the biophysical process and atmospheric process, were improved by coupling phenology dynamic in land surface model. The agricultural phenology dynamic had influenced net radiation, latent heat, sensible heat, albedo, temperature, precipitation, circulation, playing an important role in the surface energy partitioning and climate feedback. Considering the importance of agricultural phenology dynamic in land surface biophysical process and climate feedback, the following research priorities should be stressed:(1) the interactions between climate change and land surface phenology dynamic;(2) the relations between agricultural phenology dynamic and land surface reflectivity at different spectrums;(3) the contributions of crop physiology characteristic changes to land surface biophysical process;(4) the regional differences of climate feedbacks from phenology dynamic in different climate zones. This review is helpful to accelerate understanding of the role of agricultural phenology dynamic in land surface process and climate feedback.     

Influences of agricultural phenology dynamic on land surface biophysical process and climate feedback

Response and feedback of land surface process to climate change is one of the research priorities in the field of geoscience. The current study paid more attention to the impacts of global change on land surface process, but the feedback of land surface process to climate change has been poorly understood. It is becoming more and more meaningful under the framework of Earth system science to understand systematically the relationships between agricultural phenology dynamic and biophysical process, as well as the feedback on climate. In this paper, we summarized the research progress in this field, including the fact of agricultural phenology change, parameterization of phenology dynamic in land surface progress model, the influence of agricultural phenology dynamic on biophysical process, as well as its feedback on climate. The results showed that the agriculture phenophase, represented by the key phenological phases such as sowing, flowering and maturity, had shifted significantly due to the impacts of climate change and agronomic management. The digital expressions of land surface dynamic process, as well as the biophysical process and atmospheric process, were improved by coupling phenology dynamic in land surface model. The agricultural phenology dynamic had influenced net radiation, latent heat, sensible heat, albedo, temperature, precipitation, circulation, playing an important role in the surface energy partitioning and climate feedback. Considering the importance of agricultural phenology dynamic in land surface biophysical process and climate feedback, the following research priorities should be stressed: (1) the interactions between climate change and land surface phenology dynamic; (2) the relations between agricultural phenology dynamic and land surface reflectivity at different spectrums; (3) the contributions of crop physiology characteristic changes to land surface biophysical process; (4) the regional differences of climate feedbacks from phenology dynamic in different climate zones. This review is helpful to accelerate understanding of the role of agricultural phenology dynamic in land surface process and climate feedback.     

温度与降水变化的小波分析及其环境效应解释——以东北农牧交错区为例

中国东北农牧交错区属于环境变化的过渡区域和敏感区域 ,是最容易感受气候变化的地带之一。该文运用小波分析方法 ,对 5 0年代以来东北农牧交错区典型站点的温度和降水变化情景进行了分析 ,主要分析了其变化的多尺度特征和规律 ;区域和样地分析结果表明 ,降水与温度的时空变化是导致土地退化和环境变化的重要驱动力     

Winter climate change at different temporal scales in Vaccinium myrtillus, an Arctic and alpine dwarf shrub

Snow cover strongly influences plant growth in Arctic and alpine ecosystems. Snow characteristics and snowmelt timing are likely to change in a warmer climate. We studied year rings and shoot growth of the dwarf shrub bilberry ( Vaccinium myrtillus ), and species abundances of the vegetation, in response to early or late snowmelt at a study site in the Central Alps, near Davos, Switzerland. Snowmelt was manipulated on experimental plots for 3 and 30 years. Additional plots were set up along a natural snowmelt gradient, and at high and low elevation. Growth ring data showed an increasing trend in annual growth increment over the last 20 years, especially in the extraordinarily hot summer of 2003. Comparing high and low elevation sites, growth rings were wider at low elevation, but only in cold years. In years with relatively cold summers, however, xylem ring width was greater in plots with late rather than early snowmelt along the natural snowmelt gradient, possibly indicating drought stress in early snowmelt plots. Snow cover had a strong influence on species abundances along the natural snowmelt gradient, and change (not yet significant) was beginning to be seen in plots with 30 years of snow manipulation. Our results indicate that beneficial effects of early snowmelt for shrub growth may be offset in cold summers. Although early snowmelt prolongs the growing season, harsh conditions and frost events early in the growing season may become more likely, and hamper plant growth, and this could affect plant growth in all Arctic and alpine snow-dominated ecosystems.     

Display-sites selection by houbara bustard (Chlamydotis [undulata] macqueenii) in Mori, Xinjiang, People''s Republic of China

Display-site selection by houbara bustards was studied in Mori, Xinjiang, China during the breeding seasons from April to June 2000. Most of the habitats chosen for displaying were short sub-shrubby and open areas close to high shrub patches. The displaying males clearly prefer low covered areas and avoid densely covered and high vegetation sites. The vegetation density and number of plant species at display sites were significantly lower from that at randomly selected sites. The average distance to the closest shrub patch was significantly shorter at display sites than at random sites. Plant species richness, vegetative density, vegetative cover and distance to the shrub patches are possibly the most important factors that determined the display-sites selection of houbara bustard.     

气候和土地利用变化对塔里木河干流区植被覆盖变化的影响

基于遥感和地理信息系统技术,利用1998—2008年SPOT-VEGETATION归一化植被指数（NDVI）数据对塔里木河干流区1998—2007年植被覆盖的时空变化进行了监测,并从气候变化和土地利用变化双重角度分析了植被覆盖变化的原因。研究表明,塔里木河干流区植被覆盖变化经历了两个阶段:1998—2001年植被覆盖严重退化时期;2002—2007年植被覆盖度由急剧上升到缓慢下降再到持续升高时期,NDVI明显高于20世纪末期水平。塔里木河干流区植被覆盖变化存在显著的空间差异,绿洲农业灌溉区和退耕还林还草生态恢复区的植被覆盖度显著提高,天然草地植被区的植被退化严重。塔里木河干流区植被覆盖变化是气候和土地利用变化共同作用的结果。温度对植被覆盖变化的影响表现为对植被生长年内韵律的控制和秋季植被生长期的延长,年降水量的波动式上升是导致塔里木河干流区植被覆盖变化两个阶段呈现差异的主导因素。     

Perceptions and realities of land degradation in arid Otjimbingwe, Namibia

We examined the perceptions and realities of land degradation in a communal ranching area, Otjimbingwe, in arid Namibia (in south-western Africa). It is commonly perceived that large-scale degradation of Otjimbingwe has occurred due to a mixture of improper pastoral practices and pressures induced by a high human population growth rate. We sought to determine whether the inhabitants perceived land degradation to have taken place and whether their perceptions were consistent with empirical data on environmental quality. Furthermore, we wished to determine whether these pastoralists had management strategies to help them withstand the harsh environmental conditions in which they live. All respondents in our surveys perceived that the environment had become degraded. The claimed source of this degradation, a decline in annual rainfall, is inconsistent with long-term rainfall records (there was neither change nor cyclicity in rainfall over time). There is also little evidence of a decline in plant cover and soil quality in spite of the very high stocking densities. No overall pastoral strategy exists in Otjimbingwe. Options for management are extremely limited due to a variety of external and internal pressures such as a high human population growth rate, high immigration into Otjimbingwe, restricted water availability due to dams constructed upstream, and limited movement opportunities for livestock in drought periods.     

水稻发育期模型研究进展

物候是气候变化的重要指示物.随着全球变化研究的开展,已经有越来越多的研究表明,随着气象条件的变化,植物的物候期发生了明显的变化,因此,对物候的精准模拟可以帮助我们准确理解作物对全球变化的响应机制,强大的物候模型已经逐渐成为提高植物对气候变化响应的模拟精度的一个关键工具.同时作物物候的模拟也是作物模型的一个重要组成部分.水稻是最重要的粮食作物之一,水稻发育期模型研究对水稻生长模型有着重要的意义.本文对国内外水稻发育模型的发展进行了综述,并提出了目前水稻发育期模型研究中存在的问题以及发展的方向,以期后续的水稻发育期模型乃至作物模型的研究提供借鉴.     

Forest‐Cover Change from Labor‐ and Capital‐Intensive Commercial Logging in the Southern Chocó Rainforests*

This article examines two key aspects of land‐cover change in the south of the Chocó region. First, it assesses and compares the local impact on forest condition of labor‐intensive and capital‐intensive commercial logging. Second, it assesses the regional significance and permanency of these changes. Studies of land‐cover change associated with commercial logging have focused almost exclusively on capital‐intensive extraction and have assumed that after logging, degraded forests are transformed into agricultural cover. This study shows that both capital‐ and labor‐intensive logging result in similar land‐cover changes (i.e., forest degradation) if the timber sought is the same. However, labor‐intensive loggers also seek timber species not sought by capital‐intensive loggers, and this impact is statistically different from the impact of the extraction of the first group of species. Results also show that only a small fraction (20–30 percent) of the area logged is later converted to agricultural cover types. The persistence of logged forests means that up to 20 percent of the remaining forest cover could correspond to forests with significant and lasting levels of degradation. Furthermore, the different production requirements for each group of species also mean that there is a spatial differentiation in the impact of logging in the region. Logged forests are arranged into two consecutive corridors on each side of access routes (e.g., rivers). The first corridor corresponds to a narrow (approximately 1‐km) band of high‐intensity degradation. The second, broader (approximately 2‐km) forest band, with lower levels of degradation, extends inland along first‐tier corridors. A key factor determining the permanency of this land‐cover pattern is the strong control local communities have over the land in the region. This limits the spread of patterns observed in other frontier areas, especially the conversion of logged forests into agricultural cover.