Sub-fossil Chironomidae from East Africa. 2. Chironominae (Chironomini and Tanytarsini)

This paper and a companion article present illustrated guides to the identification of sub-fossil chironomid larvae (Insecta: Diptera: Chironomidae) preserved in the sediments of low- and mid-elevation lakes in East Africa. They are based on analysis of surface-sediment death assemblages from 61 lakes located in the humid to semi-arid environments in equatorial East Africa (Uganda, Kenya, Tanzania), supplemented with similar surface-sediment samples from 12 lakes in the Horn of Africa (Ethiopia), and sub-recent core samples from six lakes in Kenya and two in Uganda. We analyzed about 11,000 specimens and identified 98.4% of these to species, species group, genus, or tribe level, depending on current -taxonomic knowledge of the various considered genera and the taxonomic resolution of preserved diagnostic features. We distinguished 90 different sub-fossil morphotypes, of which 16 are Tanypodinae, 19 are Orthocladiinae, and 55 are Chironominae. In this paper we focus on the subfamily Chironominae (tribes Chironomini and Tanytarsini). The diagnostic characters distinguishing these morphotypes from each other resemble differences at the species level in the better-known Holarctic fauna, hence we consider most of our morphotypes equivalent to morphological species or groups of closely related species. Given that core samples yielded only seven morphotypes not also found in the surface-sediment samples, the current inventory of 90 taxa likely represents the large majority of distinct sub-fossil chironomid larval types to be found in East African lakes, excluding the few very large Rift lakes, cold-water lakes above treeline, and special standing-water environments such as fens and bogs. Consistent use of a single set of morphological characters to identify both fossil and living chironomid larvae would ensure exchangeability of information between modem and paleoenvironmental studies on aquatic invertebrate communities in African lakes, and increase the relevance of paleoenvironmental reconstructions to water-quality evaluations aimed at sustainable management of scarce, fluctuating surface-water resources in tropical East Africa.     

Subfossil Chironomidae from Lake Tanganyika, East Africa 1. Tanypodinae and Orthocladiinae

Analysis of subfossil remains of larval Chironomidae in 38 surface-sediment samples from between 53 and 189 m depth in Lake Tanganyika (East Africa) yielded 77 different morphotypes, including 7 Tanypodinae, 19 Orthocladiinae, and 51 Chironominae. Character-state differences between these morphotypes resemble differences at the species level in the better-known Holarctic fauna, hence we consider most of our Lake Tanganyika morphotypes equivalent to morphological species. Individual morphotypes were identified to species, genus, or tribe level depending on current alpha-taxonomic knowledge on the larvae of the group concerned, and the taxonomic resolution of preserved diagnostic features. This paper presents taxon diagnoses and an illustrated key to the Tanypodinae and Orthocladiinae in this Lake Tanganyika collection, with the aim to promote consistency of identification in studies using African Chironomidae as biological indicators of natural and anthropogenic environmental change in lacustrine ecosystems.     

Subfossil Chironomidae from Lake Tanganyika, East Africa. 2. Chironominae (Chironomini and Tanytarsini)

Analysis of subfossil remains of larval Chironomidae in 38 surface-sediment samples from between 53 and 189 meter depth in Lake Tanganyika (East Africa) yielded 77 morphotypes, among which 7 Tanypodinae, 19 Orthocladiinae, and 51 Chironominae. Character-state differences between these morphotypes resemble differences at the species level in the better-known Holarctic fauna, hence we consider most of our Lake Tanganyika morphotypes equivalent to morphological species. Individual morphotypes were identified to species, genus, or tribe level depending on current alpha-taxonomic knowledge on the larvae of the group concerned, and the taxonomic resolution of preserved diagnostic features. This paper presents taxon diagnoses and an illustrated key to the Chironominae (Chironomini and Tanytarsini) in this collection. As the living chironomid fauna of Lake Tanganyika has never been comprehensively studied, we also briefly discuss faunistic aspects, and the ecology of the recovered species and genera in relation to benthic habitat diversity in Lake Tanganyika.     

Sieve mesh size and quantitative chironomid paleoclimatology

The minimum sieve mesh size for fossil chironomid analysis is usually set at 100 μm, to ensure adequate recovery of small species. Yet taking into account the labor intensity of sorting and identifying fossil chironomid remains, the large numbers of samples requiring processing in paleoclimate studies with high temporal resolution, and the increasing need to engage non-specialist analysts in this work, it seems appealing to sieve samples through a larger mesh size that would mainly retain easier-to-see, easier-to-extract and easier-to-identify fossil specimens. In this study we evaluated the influence of sieve mesh size on chironomid-based quantitative paleoenvironmental reconstructions in African lakes. We developed two chironomid-salinity calibration data sets based on either the >100 μm or >150 μm size fractions of surface-sediment fossil assemblages, and compared the performance of salinity-inference models derived from them. We find that, despite ∼35% additional fossil loss, restriction to >150 μm data did not appreciably affect individual taxon optima or tolerances, and resulted in only a modest reduction of mean taxon richness per lake. Parameters of statistical model performance were as good, or better than those of models based on the >100 μm data, albeit only after excluding two atypically dilute (<100 μS/cm) lakes from the calibration. Application of inference models based on >150 μm or >100 μm data to a 200-year fossil record from Lake Abiyata, Ethiopia, produced very similar trends, amplitudes and uncertainty ranges of inferred past salinity change. Restriction to >150 μm data reduced the mean fossil yield of core samples by ∼20% on average, i.e. fossil loss was markedly lower than in surface-sediment samples. Using the larger mesh size reduced sample processing time by up to 50%, partly by removing a significant proportion of visually obstructive organic debris, and partly by allowing a greater fraction of chironomid fossils to be identified directly in the sorting tray. The fraction of 1st instar group taxa in surface-sediment samples was reduced from 13% to 3%, increasing the mean taxonomic resolution of fossil assemblages, and thus their ecological specificity.     

Spatial variability of chironomid death assemblages in the surface sediments of a fluctuating tropical lake (Lake Naivasha,Kenya)

Studies addressing within-lake variability of fossil chironomid assemblages are very few, and all deal with hydrologically stable temperate lakes where the question of spatial integration mostly relates to the mixing of faunal assemblages associated with shallow, warm-water habitat and those associated with deeper, cold-water habitat. Here we study within-lake variability of surface-sediment chironomid assemblages in the fairly large (∼100–170 km² since 1983) and shallow (Z _max = 5–8 m) fluctuating tropical lake basin of Lake Naivasha, Kenya, and compare the patterns observed with those in two smaller adjacent basins, one similarly shallow (Lake Oloidien, 5.1–5.7 km2, 5–8 m), the other deep and stratified (Crescent Island Crater, 1.9 km², 14–17 m). Chironomid assemblages were analysed in core-top samples and surface sediments along inshore to offshore transects, and how well individual samples represented the total (basin-wide mean) subfossil assemblage was considered both in terms of taxon richness and taxon percent composition. Within-lake variability of subfossil chironomid concentrations (with generally higher absolute values in nearshore samples) could be explained by effects of sediment winnowing and focusing, whereas between-lake variability reflected their relative susceptibility to wind-driven sediment disturbance or bottom anoxia. In all study lakes, but most significantly in lakes Naivasha and Oloidien, species distribution in the subfossil chironomid assemblages showed a strong nearshore to offshore gradient, which in these shallow lakes, reflects the dominant control of substrate and food quality on species distribution in the living community. Particularly in the larger basins, nearshore samples better represented the total lake assemblage than offshore samples, because the former always contained a component of mud-dwelling species whereas the latter often lacked a component of macrophyte-dwelling species. Our results show that although sedimentation dynamics in the shallow, wind-stressed Lake Naivasha is dominated by frequent resuspension and random sediment redistribution, the near- to offshore gradient in chironomid habitat remains imprinted on subfossil assemblages. We conclude that also in shallow fluctuating lakes, given sufficient size, incomplete pre-burial spatial integration of habitat-specific chironomid assemblages can be exploited for within-lake calibration of environmental gradients.     

Paleotemperature reconstruction in tropical Africa using fossil Chironomidae (Insecta: Diptera)

Fossil assemblages of chironomid larvae (non-biting midges) preserved in lake sediments are well-established paleothermometers in north-temperate and boreal regions, but their potential for temperature reconstruction in tropical regions has never before been assessed. In this study, we surveyed sub-fossil chironomid assemblages in the surface sediments of 65 lakes and permanent pools in southwestern Uganda (including the Rwenzori Mountains) and central and southern Kenya (including Mount Kenya) to document the modern distribution of African chironomid communities along the regional temperature gradient covered by lakes situated between 489 and 4,575 m above sea level (a.s.l). We then combined these faunal data with linked Surface-Water Temperature (SWTemp: range 2.1–28.1°C) and Mean Annual Air Temperature (MATemp: range 1.1–24.9°C) data to develop inference models for quantitative paleotemperature reconstruction. Here we compare and discuss the performance of models based on different numerical techniques [weighted-averaging (WA), weighted-averaging partial-least-squares (WA-PLS) and a weighted modern analogue technique (WMAT)], and on subsets of lakes with varying gradient lengths of temperature and other environmental variables. All inference models calibrated against MATemp have a high coefficient of determination ( r_\textjack² r_{\text{jack}}^{2}  = 0.81–0.97), low maximum bias (0.84–2.59°C), and low root-mean-squared error of prediction (RMSEP = 0.61–1.50°C). The statistical power of SWTemp models is generally weaker ( r_\textjack² r_{\text{jack}}^{2}  = 0.77–0.95; maximum bias 1.55–3.73°C; RMSEP = 1.39–1.98°C), likely because the surface-water temperature data are spot measurements failing to catch significant daily and seasonal variation. Models based on calibration over the full temperature gradient suffer slightly from the limited number of study sites at intermediate elevation (2,000–3,000 m), and from the presence of morphologically indistinguishable but ecologically distinct taxa. Calibration confined to high-elevation sites (>3,000 m) has poorer error statistics, but is less susceptible to biogeographical and taxonomic complexities. Our results compare favourably with chironomid-based temperature inferences in temperate regions, indicating that chironomid-based temperature reconstruction in tropical Africa can be achieved.     

Kenya's regional ambitions or China's Belt‐and‐Road? News media representations of the Mombasa‐Nairobi Standard Gauge Railway

Kenya's Standard Gauge Railway (SGR), and its construction and financing by China, exemplifies the prominence accorded to mega‐infrastructural projects in contemporary economic development, as well as the dependence this has engendered on external loans. Alongside, multiple geographies intertwine in the SGR which is both a component of China's global ‘Belt and Road Initiative’ (BRI), as well as an East African regional link between coastal Kenya and neighbouring landlocked countries. In most popular and academic analyses, the role of China is emphasized over the SGR's positioning within East Africa. Does this then lead to an undermining of the meanings that are attached to the SGR within Kenya? This article seeks to add Kenyan perspectives to the debate over China's infrastructural loans by examining the range of news media discourses that have emerged around the SGR. Utilizing the theoretical lens of South‐South cooperation and African agency, this study analyzes how material infrastructure becomes meaningful within the context of a political superstructure. For Kenya, this political superstructure is shaped not only by Kenya‐China relations, but also by infrastructural competitions within East Africa, as well as within Kenya itself.     

东非国家小城镇形成与发展的潜力——基于最大熵模型分析

21世纪以来,小城镇已成为东非快速城市化的重点区域。论文对坦桑尼亚、乌干达2个东非典型国家的城市进行了等级划分与数量统计,确定以10万人以下的城镇、小城市为研究对象。选取7个自然环境变量与6个社会经济变量,结合城市数据,构建最大熵模型。利用模型得到的平均生境适宜性指数(HSI)判别小城镇形成适宜度、划分潜力区,并与人口密度进行对比分析。研究发现：(1)距区域性主要交通干线距离、距全国性主要交通干线距离、归一化植被指数(NDVI)与距大城市距离对两国城镇、小城市的贡献率均较高;不同国家不同规模城市有其特有的高贡献率变量。(2)随城市规模增大,交通干线贡献率下降,自然环境变量贡献率上升。(3)坦桑尼亚北部HSI值整体高于南部,HSI高值区城镇呈团块状,小城市呈条带状。乌干达全境HSI较低,HSI高值区城镇以坎帕拉为起点呈放射网状,小城市为碎片状。两国HSI高值区基本位于路网、河流湖泊附近。(4)坦桑尼亚人口密度普遍小于乌干达,潜力区总面积则大于乌干达。潜力区与人口密度关系主要分为3种：比周边人口密度大,为历史悠久、经济实力强、靠近大中城市地区;与周边人口密度相近,为快速发展中的地区;比周边...     

Chrysophyte cysts as paleolimnological indicators of environmental change due to cottage development and acidic deposition in the Muskoka-Haliburton region, Ontario, Canada

Chrysophyte cysts preserved in recent and pre-industrial lake sediment samples from 54 Muskoka-Haliburton (Ontario) lakes were used in a paleolimnological study to determine the impact of acidic precipitation and cottage development on water quality. A total of 246 cyst morphotypes were identified. Ecological preferences of cyst morphotypes were determined using multivariate statistical analysis, cluster analysis, and species-environment correlations. Recent cyst assemblages were related to water chemistry and lake morphometric variables using Redundancy Analysis (RDA). The distribution of morphotypes was related to a gradient of acid neutralising capacity (ANC), expressed through the association of variables related to buffering (i.e. longitude, watershed area, and ionic concentration) with the first axis (₁ = 0.29). Cyst assemblages were also defined, to a lesser extent (₂ = 0.06), by a trophic status gradient, created through the combination of total nitrogen (TN), total phosphorus (TP), volume-weighted cottage density, and lake depth variables. The identification of lakewater pH and trophic status as important determinants of cyst assemblage structure allowed for the reconstruction of acidification and eutrophication related water chemistry changes using fossil cyst assemblages. The reconstruction of pre-industrial (pre-1850) water quality conditions with fossil cyst assemblages indicated that pH significantly decreased in 24.1% of the study lakes and increased in 16.7% of the lakes. Increases in pH in more alkaline drainage basins are attributed to alkalinity generation processes induced by acidic precipitation as has been shown in other studies. Total phosphorus (TP) concentrations significantly declined in 12.9% of the lakes and increased in 16.6% of lakes. Increases in [TP] were linked to cottage development. Decreases in trophic status may be due to landuse changes, the result of the acidification occurring in the area, or warmer and drier climates. A comparison of chrysophyte cyst and diatom water quality inferences show similar trends in pH changes. There is a good agreement between diatom and chrysophyte bioindicators with respect to [TP] changes in oligotrophic lakes (< 10 g/L); however, diatom inferences suggest that lakes with current [TP] values greater than 10 g/L have decreased in trophic status over time, while chrysophyte reconstructions suggest that these same lakes have become more productive systems.     

Reconstructing fluctuations of a shallow East African lake during the past 1800 yrs from sediment stratigraphy in a submerged crater basin

The sedimentology of an 8.22-m long core of late-Holocene deposits in the submerged Crescent Island Crater basin of Lake Naivasha, Kenya, is used to reconstruct decade-scale fluctuations in lake-surface elevation during the past 1800 yrs. Lake-depth inference for the past 1000 yrs is semi-quantitative, based on (1) relationships between lake level and bottom dynamics predicted by wave theory, and (2) historical validation of the effects of lake-level fluctuation and hydrologic closure on sediment composition in Crescent Island Crater and nearby Lake Oloidien. In these shallow fluctuating lakes, organic-carbon variation in a lithological sequence from clayey mud to algal gyttja is positively correlated with lake depth at the time of deposition, because the focusing of oxidized littoral sediments which dilute autochthonous organic matter before burial is reduced during highstands. The lake-level reconstruction for Lake Naivasha agrees with other adequately dated lake-level records from equatorial East Africa in its implication of dry climatic conditions during the Mediaeval Warm Period and generally wet conditions during the Little Ice Age. Crescent Island Crater survived widespread aridity in the early-19th century as a fresh weedy pond, while the main basin of Lake Naivasha and many other shallow East African lakes fell dry and truncated their sediment archive of Little Ice Age climatic variability.     

非洲世界遗产的分布特征

以世界文化和自然遗产为主要类型, 通过图表分析和计算密度, 分全洲、地区和国家3 个尺度, 研究非洲世界遗产的分布特征。从全洲看, 非洲大陆沿海和海岛地区、中等河流流域与非洲东部局部区域世界遗产分布相对集中。不同地区中, 东部、北部和西部非洲比较稠密, 其中东部非洲密度最高。各国中, 塞内加尔、突尼斯、冈比亚、佛得角、毛里求斯和塞舌尔密度最高, 其次为科特迪瓦、摩洛哥、多哥、肯尼亚、乌干达、马拉维和津巴布韦等国。世界文化遗产在东部、北部和西部非洲同样密集, 密度较高的有埃及、摩洛哥、突尼斯、贝宁、加纳、塞内加尔、多哥、冈比亚、佛得角、埃塞俄比亚、肯尼亚、马拉维、毛里求斯和津巴布韦等国。世界自然遗产以东部非洲密度最高, 主要分布在突尼斯、尼日尔、几内亚、科特迪瓦、塞内加尔、喀麦隆、中非、民主刚果、坦桑尼亚、肯尼亚、塞舌尔、乌干达、马达加斯加、南非、津巴布韦和马拉维等国。     

The sensitivity of East African rift lakes to climate fluctuations

Sequences of paleo-shorelines and the deposits of rift lakes are used to reconstruct past climate changes in East Africa. These recorders of hydrological changes in the Rift Valley indicate extreme lake-level variations on the order of tens to hundreds of meters during the last 20,000 years. Lake-balance and climate modeling results, on the other hand, suggest relatively moderate changes in the precipitation-evaporation balance during that time interval. What could cause such a disparity? We investigated the physical characteristics and hydrology of lake basins to resolve this difference. Nine closed-basin lakes, Ziway-Shalla, Awassa, Turkana, Suguta, Baringo-Bogoria, Nakuru-Elmenteita, Naivasha, Magadi-Natron, Manyara, and open-basin Lake Victoria in the eastern branch of the East African Rift System (EARS) were used for this study. We created a classification scheme of lake response to climate based on empirical measures of topography (hypsometric integral) and climate (aridity index). With reference to early Holocene lake levels, we found that lakes in the crest of the Ethiopian and Kenyan domes were most sensitive to recording regional climatic shifts. Their hypsometric values fall between 0.23–0.29, in a graben-shaped basin, and their aridity index is above unity (humid). Of the ten lakes, three lakes in the EARS are sensitive lakes: Naivasha (HI = 0.23, AI = 1.20) in the Kenya Rift, Awassa (HI = 0.23, AI = 1.03) and Ziway-Shalla (HI = 0.23, AI = 1.33) in the Main Ethiopian Rift (Main Ethiopian Rift). Two lakes have the graben shape, but lower aridity indices, and thus Lakes Suguta (HI = 0.29, AI = 0.43) and Nakuru-Elmenteita (HI = 0.30, AI = 0.85) are most sensitive to local climate changes. Though relatively shallow and slightly alkaline today, they fluctuated by four to ten times the modern water depth during the last 20,000 years. Five of the study lakes are pan-shaped and experienced lower magnitudes of lake level change during the same time period. Understanding the sensitivity of these lakes is critical in establishing the timing or synchronicity of regional-scale events or trends and predicting future hydrological variations in the wake of global climate changes.     

Diatom-based models for inferring past water chemistry in western Ugandan crater lakes

Diatom surface sediment samples and corresponding water chemistry were collected from 56 lakes across a natural conductivity gradient in western Uganda (reflecting a regional climatic gradient of effective moisture) to explore factors controlling diatom distribution. Here we develop a regional training set from these crater lakes to test the hypothesis that this approach, by providing more appropriate and closer analogues, can improve the accuracy of palaeo-conductivity reconstructions, and so environmental inferences in these lake systems compared to larger training sets. We compare this output to models based on larger, but geographically and limnologically diverse training sets, using the European Diatom Database Initiative (EDDI) database. The relationships between water chemistry and diatom distributions were explored using canonical correspondence analysis (CCA) and partial CCA. Variance partitioning indicated that conductivity accounted for a significant and independent portion of this variation. A transfer function was developed for conductivity (r _jack ² ?=?0.74). Prediction errors, estimated using jack-knifing, are low for the conductivity model (0.256 log₁₀ units). The resulting model was applied to a sedimentary sequence from Lake Kasenda, western Uganda. Comparison of conductivity reconstructions using the Ugandan crater lake training set and the East Africa training set (EDDI) highlighted a number of differences in the optima of key diatom taxa, which lead to differences in reconstructed values and could lead to misinterpretation of the fossil record. This study highlights issues of how far transfer functions based on continental-scale lake datasets such as the EDDI pan-African models should be used and the benefits that may be obtained from regional training sets.     

Hydrologic control on the oxygen-isotope relation between sediment cellulose and lake water, western Taimyr Peninsula, Russia: Implications for the use of surface-sediment calibrations in paleolimnology

Systematic variability occurs between the oxygen isotopic composition of lake water sampled in mid-summer 1993 and cellulose extracted from surficial sediments of a suite of lakes spanning the forest-tundra transition near Noril'sk, Russia. Some tundra and all forest-tundra lakes show greater deviation from expected cellulose-water isotopic separation than forest lakes, apparently because of greater sensitivity to ¹⁸O-depleted snowmelt contributions. Cellulose derived from aquatic plants naturally integrates fluctuations in lake water ¹⁸O, providing a signal that is inherently more representative of average thaw season lake water ¹⁸O than the measure of instantaneous ¹⁸O obtained from an individual sample of lake water. Thus, indiscriminate use of empirical cellulose-water relations derived from calibration samples could lead to erroneous assessment of paleohydrology from the oxygen-isotope stratigraphy of sediment cores from arctic lakes. However, deviation from the expected cellulose-water fractionation is a source of lake-specific hydrologic information useful for qualifying paleoenvironmental interpretations and possibly constraining non-isotopic methods that rely on surface-sediment calibrations.     

Midges of the genus Pseudodiamesa Goetghebuer (Diptera, Chironomidae): current knowledge and palaeoecological perspective

Larvae of the genus Pseudodiamesa Goetghebuer, which includes 12 described valid species and is divided by Makarchenko and Makarchenko (1999) into two species groups, Pseudodiamesa branickii group and Pseudodiamesa nivosa group, are widespread in lakes and running waters of cold, high-latitude and high-altitude areas of the Northern Hemisphere. Larval remains of the genus are often found in subfossil assemblages from these cold regions, but intra-genus morphotypes usually are not distinguished by palaeoecologists. Current knowledge of the ecology of the species indicates that the Pseudodiamesa nivosa group is more cold-adapted than the Pseudodiamesa branickii group and, therefore, these two intra-genus morphotypes cannot be amalgamated into a larger taxonomic unit without losing substantial information. Here we present the morphological characters of head capsules of Pseudodiamesa larvae attributed to the different species-group morphotypes, which are clearly visible in subfossil specimens. The information summarized in this paper will help provide more reliable chironomid-based palaeoclimatic inferences from lake sediment records covering colder phases through the late Quaternary in the Northern Hemisphere.     

Solar and anthropogenic imprints on Lake Masoko (southern Tanzania) during the last 500 years

The Masoko crater-lake in southern Tanzania provides a continuous record of environmental changes covering the last 500 years. Multi-proxy studies were performed on a 52 cm sediment core retrieved from the deepest part of the lake. Magnetic, organic carbon, geochemical proxies and pollen assemblages indicate a dry climate during the ‘Little Ice Age’ (AD 1550–1850), confirming that the LIA in eastern Africa resulted in marked and synchronous hydrological changes. However, the direction of response varies between different African lakes (low versus high lake-levels), indicating strong regional contrasts that prevent the clear identification of climate trends over eastern Africa at this time. Inferred changes in Masoko lake-levels closely resemble the record of solar activity cycles, indicating a possible control of solar activity on the climate in this area. This observation supports previous results from East African lakes, and extends this relationship southward. Finally, anthropogenic impact is observed in the Masoko sediments during the last 60 years, suggesting that human disturbance significantly affected this remote basin during colonial and post-colonial times.     

A cladoceran-based paleolimnological assessment of the impact of forest harvesting on four lakes from the central interior of British Columbia, Canada

To investigate the impact of forest harvesting on lake ecosystems, six lakes (four impact, two reference) from central British Columbia, Canada were assessed using cladoceran remains preserved in the lake sediment cores. Two temporal resolutions were analyzed: a decadal scale for the past 70 years, and a high-resolution scale (2 year intervals for ten years before and after harvesting). Three lakes which experienced forestry activity in their watersheds in the early 1960s, and one lake which experienced forestry activityin the mid 1970s, showed subtle but statistically significant changes in cladoceran species composition following forest harvesting (analysis of similarity tests) at the decadal scale, whereas only two of these lakes showed significant changes at the higher resolution. These changes may be due to increased nutrient levels that might be associated with forest harvesting. The two reference lakes, which had not experienced any known large-scale anthropogenic watershed disturbances in the past century, showed no significant shifts in the cladoceran species assemblages at either temporal scale.     

Detecting ancient wild rice (Zizania spp. L.) using phytoliths: a taphonomic study of modern wild rice in Minnesota (USA) lake sediments

Wild rice (Zizania spp. L.) is a North American native grain with spiritual and dietary significance to many native people. Wild rice is also an important aquatic plant that provides critical habitat to wetland and aquatic wildlife. Past distribution of wild rice in North America is poorly understood, largely because of the limited taxonomic resolution of Poaceae pollen. A novel technique for detecting diagnostic Zizania silica phytoliths allows unambiguous identification of this taxon in lake sediments. We need to better understand modern depositional patterns of phytoliths in lake sediments, however, before attempting detailed paleoreconstructions. We analyzed distributions of diagnostic Zizania and other Poaceae phytoliths in modern sediments from three lakes with variable percent cover of wild rice and a non-wild rice control lake in central Minnesota. Absolute counts of phytoliths per gram sediment were achieved using an exotic diatom marker. Non-Zizania short-cell phytoliths, i.e. phytoliths from wetland grasses Phragmites australis and Muhlenbergia glomerata, dominate the assemblages in all lakes. Most Poaceae short-cell phytoliths appear to be derived locally, with little evidence for regional inheritance from eolian or alluvial processes. Because of anatomical differences in decay of plant debris and other taphonomic issues, Zizania inflorescence rondel phytoliths were most abundant, with morphotypes from other parts rarely encountered. Even in sediments under the densest wild rice stands, Zizania phytoliths contributed a maximum of 9 % to total Poaceae phytolith abundance. Lake morphology also affects the depositional pattern of phytoliths in modern sediments, so coring locations should be considered carefully. At least 500 phytoliths should be counted to detect a sufficient number of wild rice phytolith morphotypes. Diagnostic Zizania phytoliths are a reliable tool for wild rice detection in paleolake sediments.     

东非油气资源开发的历史透视与现状解析

东非油气资源开发始于一战前,二战后出现第一个勘探高峰期。随着冷战结束,东非油气资源开发的力度不断加大,国际能源公司纷纷入驻。在中小型能源公司的积极开拓下,于2006年以后相继取得重大勘探成果。东非油气资源已呈现出分布广、储量大的特点,并吸引一批国际大型能源公司的投资,加快了能源产业的发展,推动了能源立法的建设。随着东非在全球能源版图中地位的凸显,地区外大国在该地区的博弈也日趋激烈。东非各国则获得发展的新机遇,并将推进地区一体化进程。然而,若东非各国不能妥善处理政府能力问题、腐败问题、经济结构问题、利益分配问题、恐怖主义问题等,将有可能陷入"资源诅咒"的陷阱中。