生物结皮中优势蓝藻胞外多糖对几种荒漠草本植物种子萌发的影响

蓝藻是荒漠生物结皮的重要组成。选取生物结皮中的优势蓝藻(具鞘微鞘藻(Microcoleus vaginatus)、眼点伪枝藻(Scytonema ocellatum)、地木耳(Nostoc commune)),通过提取其胞外多糖,研究了不同蓝藻胞外多糖对荒漠植物种子萌发的影响。结果表明:具鞘微鞘藻、眼点伪枝藻的胞外多糖对粗枝猪毛菜(Salsola subcrassa)的种子萌发表现为低浓度抑制、高浓度促进作用,地木耳的胞外多糖仅在高浓度时对粗枝猪毛菜的种子萌发具有促进作用(p<0.05);具鞘微鞘藻胞外多糖对小花荆芥(Nepeta micrantha)种子萌发无明显的影响作用,但眼点伪枝藻和地木耳对小花荆芥种子萌发具有抑制作用。具鞘微鞘藻、眼点伪枝藻和地木耳胞外多糖对琉苞菊(Hyalea pulchella)种子萌发具有明显的抑制作用。藻种胞外多糖组分的差异和不同种子的生物学特性差异可能是造成不同藻种胞外多糖对种子萌发产生不同影响的主要原因。     

Extracellular polysaccharides from cyanobacterial soil crusts: A review of their role in dryland soil processes

Cyanobacterial soil crusts are a community of microorganisms living in the soil surface of different habitats worldwide. Through photosynthesis, cyanobacteria produce extracellular polysaccharides (EPS) increasing the soil carbon (C) pool as carbohydrates. The layer of polysaccharides also acts as a mechanical structure surrounding the filamentous cyanobacteria that together with the soil particles form stable aggregates in the topsoil thus decreasing C loss by erosion. Thus despite their apparent importance to the dryland system we have only a limited understanding of their role and possible applications in dryland soil environments. This review draws on these disparate sources of information in order to provide a summary of our understanding of the characteristics, behaviour and influence of cyanobacterial EPS in dryland soils and makes recommendations for further research.     

阿拉善高原沙漠地区植物钙质根管的矿物组成特征

在阿拉善高原地区,植物钙质根管的成因及其是否具有一定的环境指示意义,目前尚存在较大争论。分析该区域植物钙质根管的矿物组成,可以为植物钙质根管的成因提供新的证据解释。本文通过近5年的野外考察,先后采集了巴丹吉林沙漠、腾格里沙漠及乌兰布和沙漠腹地33组植物钙质根管和33个地表风积砂样品,通过X衍射的方法,分析了上述样品的矿物组成。结果显示:上述沙漠地区植物钙质根管和地表风积砂中矿物组成以硅酸盐矿物、氧化物和氢氧化物矿物、碳酸盐类矿物为主,没有明显的区域差异。其中碳酸盐类矿物均以方解石为主,较少含有其他盐类矿物,并且植物钙质根管中碳酸盐类矿物百分含量明显高于地表风积砂中碳酸盐类矿物的百分含量。因此,单纯的地下水重结晶作用无法解释本研究区内植物钙质根管中盐类矿物的组成以及碳酸盐类矿物的来源,并且在研究区内现代气候条件不利于植物根系的钙化和钙质根管的形成。植物钙质根管应是地质历史时期的产物,指示了相对湿润的气候条件。     

Pre-drilling reflection survey of the Black Forest, SW Germany

Summary. The unified seismic exploration program, consisting of 345 km of deep reflection profiling, a 200 km refraction profile, an expanding spread profile and near-surface high resolution reflection meaasurements, revealed a strongly differentiated crust beneath the Black Forest. The highly reflective lower crust contains numerous horizontal and dipping reflectors at depths of 13-14 km down to the crust-mantle boundary (Moho). The Moho appears as a flat horizontal first order discontinuity at a relatively shallow level of 25–27 km above a transparent upper mantle. From modelling of synthetic near-vertical and wide-angle seismograms using the reflectivity method the lower crust is supposed to be composed of laminae with an average thickness of about 100 m and velocity differences of greater than 10% increasing from top to bottom. The upper crust is characterised by mostly dipping reflectors, associated with bivergent underthrusting and accretion tectonics of Variscan age and with extensional faults of Mesozoic age. A bright spot at 9.5 km depth is characterised by low velocity material suggesting a fluid trap. It appears on all of the three profiles in the centre of the intersection region. The upper crust seems to be decoupled from the lowest crust by a relatively transparent zone which is' also identified as a low-velocity zone. This low velocity channel is situated directly above the laminated lower crust. The laminae in the Rhinegraben area are displaced vertically to greater depths indicating an origin before Tertiary rift formation and a subsidence of the whole graben wedge.     

准噶尔盆地古尔班通古特沙漠生物结皮蓝藻研究

通过对古尔班通古特沙漠中9个样地73份生物土壤结皮蓝藻的研究,分析该沙漠生物结皮中蓝藻的区系组成、生态分布特点和蓝藻在生物结皮中的作用机制。研究表明,该沙漠中有蓝藻门植物6科25属77种,其中以丝状种类占优势;蓝藻在不同地貌部位分布略有差异,在丘间低地种类最为丰富;不同种类的蓝藻对地貌部位的选择性不同,大部分对地貌部位具有选择性;蓝藻种类和数量在结皮层较多,在结皮下层明显减少。生物结皮的显微观察揭示了丝状蓝藻和胞外多糖形成错综复杂的网状结构,起着捆绑、黏结沙粒和固定沙粒的作用。     

浙江瑶琳洞风化碳酸钙景观复生试验

目前,世界上80%以上的风景旅游洞穴,开放后,大量游客涌入和高能景观灯的应用,洞穴环境及能量埸和化学埸发生了巨大变化,洞穴碳酸钙景观遭受严重破坏,其景观的美学价值大大降低。文中论述了在已遭风化破坏的碳酸钙景观上滴喷高Ca⁺²碱液,吸收洞穴空气CO₂,形成CaCO₃沉积层的试验。试验结果表明,Ca⁺²碱液滴喷技术可以恢复风化碳酸钙景观的美学度,同时又降低洞穴空气中的浓度,改善洞穴环境。     

Soil physico-chemical properties affecting the distribution of biological soil crusts along an environmental transect at Zapotitlán drylands, Mexico

The composition of cyanobacteria, mosses, and lichens of biological soil crusts were correlated with soil characteristics and sun exposure along an environmental transect. The study was conducted in Zapotitlán drylands, a locality within the Tehuacán–Cuicatlán Valley, central Mexico, where a great variety of environments for crust development exist due to landscape fragmentation. Data were analyzed with redundancy and Sorensen analysis. Soil crusts consisted of different combinations of cyanobacteria (7 species), mosses (19 species), and lichens (8 species). The relative frequencies of these groups were positively correlated with soil apparent density and lichens were also positively correlated with soil pH. However, there were no significant correlations with sun exposure.     

The last 500 year of sedimentation in Shkodra Lake (Albania/Montenegro): paleoenvironmental evolution and potential for paleoseismicity studies

Lake Shkodra (northern Albania, southern Montenegro) is a large (45 km length, 15 km width) and shallow (5 m mean depth) lake, developed on a mainly karstic carbonate substratum. Its recent sedimentary fill (mixed calcareous/siliciclastic clayey silts) was analysed through short gravity cores representing five centuries of environmental archive. A combination of high resolution texture analysis (particle size), magnetic susceptibility, carbon content, and radionuclide activity profiles (²¹⁰Pb, ¹³⁷Cs, ²⁴¹Am) permitted us to detect and date environmental changes, such as the Little Ice Age and the 1962–1963 catastrophic floods. Anthropogenic influences on the watershed, such as damming of the Drin River in the late 1970s, also appear to be recorded. Impacts of known strong earthquakes (1905, 1979), however, could not be clearly detected in the sediment record. The potential of the Shkodra lacustrine archive for paleoseismic investigations on long time intervals (such as Holocene) is discussed. This paper is a preliminary contribution to Shkodra Lake’s Holocene paleolimnology.     

Aeolian influences on the soils and landforms of the southern Yilgarn Craton of semi-arid, southwestern Australia

Within a landscape developed on the deeply weathered, dominantly granitic rocks of the Yilgarn Craton in southwestern Australia, there is strong evidence of prior aeolian activity. Multiple arrays of clayey saltation deposits occur as clay dunes or lunettes extending up to 5 km to the southeast of playas, and quartzose sand dunes and sheets occur in a 2-km wide band that extends 10 km southeast of an ephemeral creek line. Parabolic dunes occur within some clayey lunettes. The parallel orientation of these diverse features and the elliptical shape of the playas suggest that the winds that have created these geomorphic entities have been from the northwest. Multiple lunette arrays with up to seven members have not been previously reported in this region.There is evidence for a more widespread, but more subtle, aeolian influence on the soils. Dust deposits appear to mantle parts of the landscape and are inferred to be coeval with the clayey saltation deposits. Although these do not occur as a discrete layer, evidence includes a plume of calcareous and illitic soils southeast of the major playa, in a landscape otherwise characterised by acidic, kaolinitic soils. Similarly, the occurrence of deep sandy soils on many slopes with a southeasterly aspect may indicate the interference of topography on the transport of saltating sands during an arid climatic phase. The presence of such aeolian deposits has implications for agricultural management, mineral exploration and the interpretation of ecological gradients in similar undisturbed landscapes.Current desertification, in the form of salinization and wind erosion, has precedence in this landscape. The aeolian deposits provide a means of interpreting landscape-scale responses to previous climate change and a key to predicting possible outcomes of both desertification and future climate change. Assuming that Bowler's theory that clay dunes require an adjacent salinized playa bed to form is correct, the arrays of clayey lunettes together with vegetated former playas indicate that the landscape has undergone multiple cycles of salinization and recovery. The lunettes may thus provide a means of predicting landscape hydrological responses to the widespread removal of natural vegetation for farming. Dating of the various sequences is required to provide a time scale for these landscape processes.     

丹霞山顺层洞穴风化特征的试验研究

以丹霞盆地丹霞组2段顺层洞穴为研究对象,探讨不同岩性红层抗风化能力的差异,重点关注软岩夹层的风化特征及其对顺层洞穴发育的影响。在野外考察的基础上,选择3个典型顺层洞穴为案例,采集了洞穴剖面不同岩性红层的岩样及夹层水样品,进行一系列的岩矿理化测试和水样分析实验。结果表明：洞穴剖面岩性的不均一性及软岩夹层的风化凹进是形成丹霞山顺层洞穴的主要原因。顺层洞穴发育的软岩夹层一般是粉砂质泥岩,粘土矿物含量高,以泥质胶结为主,吸水性较强,开型空隙率较大,抗压强度小,易于风化破碎;上下岩层主要是砂岩和砂砾岩,以钙质和铁质胶结为主,吸水性较弱,开型孔隙率较小,抗压强度较大,风化速度较慢。在洞穴岩层风化过程中,粘土矿物的胀缩、钙质胶结物的溶解, 及铁质胶结物的水化作用等是导致岩体结构破坏的重要因素,但促使不同岩性红层风化的主导因素有所差别。粉砂质泥岩的风化以粘土矿物的胀缩作用为主,砂岩和砂砾岩的风化则以钙质胶结物的溶解作用为主。此外,酸性雨水渗入岩体后与各种矿物进行的一系列离子交换作用也促进了顺层洞穴岩层的风化。     

Salt tolerance of Microcoleus vaginatus Gom., a cyanobacterium isolated from desert algal crust, was enhanced by exogenous carbohydrates

Microcoleus vaginatus isolated from a desert algal crust of Shapotou was cultured in BG-11 medium containing 0.2 mol l⁻¹ NaCl or 0.2 mol l⁻¹ NaCl plus 100 mg l⁻¹ sucrose, extracellular polymeric substances (EPS) or hot water-soluble polysaccharides (HWP), respectively. Photosynthetic oxygen evolution rates, photosystem II activity (Fv/Fm) and dark respiration of NaCl-stressed cells were enhanced significantly by the added sucrose or EPS under salt stress conditions (0.2 mol l⁻¹ NaCl). Compared with cells treated with salt alone, sodium contents in cells reduced significantly; the content of cellular total carbohydrate did not change, and intracellular sucrose, water-soluble sugar increased significantly following the addition of exogenous carbohydrates. Sucrose synthase (SS) activity of NaCl-stressed cells increased following the addition of sucrose, and sucrose phosphate synthase (SPS) activity of NaCl-stressed cells increased following the addition of exogenous sucrose, EPS or HWP compared with cells stressed with NaCl only. The results suggested that the extruded EPS might be re-absorbed by cells of M. vaginatus as carbon source, they could increase salt tolerance of M. vaginatus through the changes of carbohydrate metabolism and the selective uptake of sodium ions.     

Recently induced anoxia leading to the preservation of seasonal laminae in two NE-German lakes

The recent sediments of two lakes in the NE German lowland became seasonally laminated at different times. Anoxic bottom conditions resulted from a surplus of organic matter (OM), in the early stage indicated by irregularly laminated sediments comprising abundant iron-sulfide framboids. Their diagenetic formation predates the preservation of biochemical calcite varves. In the larger, deeper Lake Tiefer See near Klocksin, anoxia developed stepwise. A first anoxic pulse was contemporary with inflow narrowing by railway-dam construction and accumulation of OM. It was favored by a decrease of the intensity of lake circulation (turnover). Nutrients introduced from artificial fertilizer then increased the primary production (diatoms) to the point of OM surplus and seasonal laminae formation started 40 years later in 1924. In the smaller, shallower Lake Tiefer See in the Uckermark, a massive pulse of iron sulfide was centered around 1960, seven years after installation of piped field drainage into the lake. Anoxia developed rapidly with the nutrients drained from a fertilized groundwater catchment that is 10 times larger than the surface catchment, while surface erosion was reduced. Reducing bottom conditions became regular and the seasonal lamination was preserved after 1967. Morphological criteria to screen lakes for varved sediments should include reductions of natural lake inflow and catchment increase, such as by inflow of field drainage. Similar developments of increased nutrient input or intensity decrease of lake circulation may result from historical human activities but also from natural processes.     

藓类结皮斑块面积与环境因子的关系

古尔班通古特沙漠生物结皮占该沙漠总面积的30%左右,苔藓结皮是其中的优势者,在防风固沙、捕获隐匿降水和水土保持方面发挥着重要的生态作用。自然状态下,该沙漠苔藓结皮多以镶嵌式或纯群状斑块分布,本文从斑块尺度出发,以斑块面积为指标,分析了该沙漠藓类结皮斑块的分布格局和特征。结果表明:（1）在调查的7 372个藓类结皮斑块中,有6 930个斑块的面积在100 cm2以内,占总斑块数目的94%,面积在100 cm2以上的斑块约为442个,占总斑块数的6%。（2）本研究共调查了143 223 cm2苔藓结皮,其中100 cm2以下的小斑块面积约为57 289 cm2,占总调查面积的40%;其次是大小在100~200 cm2的斑块,面积约为32 941 cm2,占总调查面积的23%;大小在200 cm2以上的斑块面积约为50 128 cm2,占总面积的35%。面积较大的斑块主要分布在沙漠东南部,西北部藓类结皮斑块面积较小,这一变化特征与苔藓结皮在区域尺度的优势发育区一致。（3）藓类结皮斑块面积大小与粒径在0.2 mm以下的土壤含量正相关,大粒径的土壤不利于藓类结皮形成较大的斑块。（4）藓类结皮斑块面积与主要灌丛盖度呈微弱负相关,与土壤有机质、全氮、速效钾、可溶性钙含量显著正相关,这与区域尺度的报道一致。     

青藏高原古岩溶的性质、发育时代和环境特征

青藏高原目前多处所见岩溶地貌主要属第三纪古岩溶之地下部分经后期剥蚀而出露于地表的，风化壳红土和洞穴次生化学沉积等古岩溶相关沉积也多以残留形态出露在已经发生解体的高原主夷平面的南和东南缘，风化壳红土中所含粘粒部分的主要化学成分为SiO2,Al2O3和Fe2O3；粘土矿物多属“伊利石-高岭石”型组合，少数样品属“高岭石-伊利石”型组合，据硅酸系数和粘土矿物组合判断，古岩溶风化壳红土的发育阶段处在化学风化的初期，但由于目前所见红土仅反映当时风化壳剖面根部的化学风化状况，故其较弱的风化指数仍能间接指示古岩溶发育时期湿热的地表环境，扫描电镜观测结果亦表明，风化壳红土中石英砂的表面结构特征以化学溶蚀形成的为主，机械侵蚀形成的为辅，反映了高原风化壳红土垢长期残留特征，对应风化壳发育时期的湿热环境。     

Late Holocene linkages between decade–century scale climate variability and productivity at Lake Tanganyika, Africa

Microlaminated sediment cores from the Kalya slope region of Lake Tanganyika provide a near-annually resolved paleoclimate record between ∼∼2,840 and 1,420 cal. yr B.P. demonstrating strong linkages between climate variability and lacustrine productivity. Laminae couplets comprise dark, terrigenous-dominated half couplets, interpreted as low density underflows deposited from riverine sources during the rainy season, alternating with light, planktonic diatomaceous ooze, with little terrigenous component, interpreted as windy/dry season deposits. Laminated portions of the studied cores consist of conspicuous dark and light colored bundles of laminae couplets. Light and dark bundles alternate at decadal time scales. Within dark bundles, both light and dark half couplets are significantly thinner than within light bundles, implying slower sediment accumulation rates during both seasons over those intervals.Time series analyses of laminae thickness patterns demonstrate significant periodicities at interannual–centennial time scales. Longer time scale periodicities (multidecadal to centennial scale) of light and dark half couplet thicknesses are coherent and in some cases are similar to solar cycle periods on these time scales. Although laminae thickness cycles do not strongly covary with the actual Δ¹⁴C record for this same time period, two large Δ¹⁴C anomalies are associated with substantial decreases in both light and dark laminae thickness. In contrast to the multidecadal– centennial time scale, significant annual to decadal periodicities, which are broadly consistent with ENSO/PDO forcing and their impact on East African climate, are not coherent between light and dark half couplets. The coherency of light–dark couplets at decadal–centennial time scales, but not at shorter time scales, is consistent with a model of a long-term relationship between precipitation (recorded in wet season dark laminae thickness) and productivity (light laminae thickness), which is not manifest at shorter time scales. We hypothesize that this coupling results from long-term recharging of internal nutrient loading during wet periods (higher erosion of soil P) and reduced loading during drought intervals. The relationship is not expressed on short time scales during which the dominant control on productivity is wind-driven, dry season upwelling, which is uncorrelated with wet-season precipitation. Our record greatly extends the temporal record of this quasi-periodic behavior throughout the late Holocene and provides the first evidence linking decade- to century-scale episodes of enhanced productivity to enhanced precipitation levels and nutrient recharge in a productive tropical lake.     

辽东半岛红色风化壳粒度分布特征浅析

运用贝克曼激光衍射粒度分析仪对辽东半岛大连市棋盘子村和龙王村的红色风化壳作粒度特征分析,粒度频率分布曲线的差异指示出两地风化壳风化程度较弱,风化不完全.结合X射线荧光光谱仪进行化学分析进一步探讨两地的风化特征,发现两地各自发育的母岩都具有不均一性,棋盘子村风化壳的发育与母岩具有同源关系,龙王村风化壳发育与母岩非同源.与南方红色风化壳比较,南方红色风化壳风化程度高于棋盘子村,且母岩具有均一性.     

Climate signal in varve thickness: Lake La Cruz (Spain), a case study

Lake La Cruz is a meromictic, karstic lake with annually laminated sediment formed by summer pulses of calcite deposition. The aim of this study was to explore the potential use of the laminated sediment from Lake La Cruz as a quantitative climate proxy, by calibrating lamina thickness against instrumental climate data. Statistical analysis of the relation between lamina thickness and the meteorological dataset indicated a high correlation between calcium carbonate lamina thickness and rainfall from December to March (r  = 0.725, P < 0.01, n = 35). Winter rainfall anomalies in the area are, in turn, highly negatively correlated with the North Atlantic Oscillation index (NAO, r = −0.832; P < 0.01; n = 53). We propose a regression model to infer past winter rainfall from calcium carbonate laminae thickness. These results highlight new possibilities for paleoenvironmental research using calcite laminated sediment records as climate proxies, especially to study past rainfall variability.     

黄河中游降雨特性对泥沙粒径的影响

以黄河中游36个有泥沙粒径资料的水文测站流域为样本，在不同自然地理类型流域划分的基础上，建立了流域泥沙粒径特片与降雨特性的关系。结果表明，在不同类型流域，流域泥沙的粗细与降雨季节性变率和年际变率之间存在着相当好的线性正相关关系；而与年降雨量和降雨不均匀系数之间呈明显的非线性关系。流域地面物质、植被和地貌发育程序等下垫面环境因素对泥沙粒径特性与降雨特性之间关系起着十分重要的控制作用。不同类型流域曲线的斜率各不相同，在图中所处的位置也不相同。     

Taking varves to bits: Scanning electron microscopy in the study of laminated sediments and varves

Conventional high resolution studies of varved sediments are able to identify clastic and biogenic laminae, but are often unable to resolve the nature of fine-scale lamination contained therein. This intra-annual signal provides us with the highest potential resolution from the sedimentary record and can be resolved using scanning electron microscopy (SEM). Six case studies from lacustrine and marine settings are presented to illustrate the combination of clastic and biogenic fabric types typically found in laminated sediments. Clastic laminae fabrics include those which originate through grain settling and those which are event deposits. The correct identification of event deposits is essential if varves are to be used chronologically. SEM-based biogenic laminae fabric studies have identified seasonal faunal successions where individual laminae may be less than 100 thick and most recently, deep chlorophyll maxima (DCM) summer diatom floras, providing an insight into seasonal scale processes. High resolution lamina fabric studies can provide a basis for generating records of seasonal and inter-annual variability, thus contributing to our understanding of lacustrine and marine processes and palaeoenvironmental interpretation.     

Biological soil crusts exhibit a dynamic response to seasonal rain and release from grazing with implications for soil stability

In Northern Mexico, long-term grazing has substantially degraded semiarid landscapes. In semiarid systems, ecological and hydrological processes are strongly coupled by patchy plant distribution and biological soil crust (BSC) cover in plant-free interspaces. In this study, we asked: 1) how responsive are BSC cover/composition to a drying/wetting cycle and two-year grazing removal, and 2) what are the implications for soil erosion? We characterized BSC morphotypes and their influence on soil stability under grazed/non-grazed conditions during a dry and wet season. Light- and dark-colored cyanobacteria were dominant at the plant tussock and community level. Cover changes in these two groups differed after a rainy season and in response to grazing removal. Lichens with continuous thalli were more vulnerable to grazing than those with semi-continuous/discontinuous thalli after the dry season. Microsites around tussocks facilitated BSC colonization compared to interspaces. Lichen and cyanobacteria morphotypes differentially enhanced resistance to soil erosion; consequently, surface soil stability depends on the spatial distribution of BSC morphotypes, suggesting soil stability may be as dynamic as changes in the type of BSC cover. Longer-term spatially detailed studies are necessary to elicit spatiotemporal dynamics of BSC communities and their functional role in biotically and abiotically variable environments.