盐生荒漠土壤水稳定氢、氧同位素组成季节动态

对准噶尔盆地东南缘降水和土壤水稳定氢、氧同位素组成（δ¹⁸O和δD）进行了测定,分析了降水中δ¹⁸O和δD值的季节变化规律,表层土壤水中δ¹⁸O和δD值对降水脉冲的动态响应以及不同深度土壤水中δ¹⁸O和δD值的变化特征。结果表明：该区域大气降水线为δD=7.691δ¹⁸O+4.606;降水与表层土壤水中δ¹⁸O和δD值表现出明显的季节变化特征;表层土壤水中δ¹⁸O和δD值、质量含水量对降水脉冲响应显著,且不同量级的降水导致不同程度的响应。利用LSD法对0~300 cm土层内土壤水中δ¹⁸O和δD值进行多重比较分析,可将土壤在垂直方向上分为3层,表层（0~50 cm）土壤水分活跃,稳定同位素值随深度的增加而迅速减小;中间层（50~180 cm）土壤水分相对活跃,既受到降水入渗和蒸发作用的影响,也有地下水的补给;深层（180~300 cm）水分来源稳定,土壤水中δ¹⁸O和δD值基本不变。     

Mechanical shaking and soil water affect the growth of Psammochloa villosa in the Mu Us Sandland

The Mu Us Sandland is basically characterized by water shortage and high wind. Thus, wind-induced mechanical perturbation (MP) and soil water availability are likely to interact to affect plant growth. Since high water availability and MP can induce responses that are in the opposite direction, we hypothesized that MP effects on perennial grasses might be mitigated by increased soil water availability in the Mu Us Sandland. We counducted an experiment in which seedlings of Psammochloa villosa were subjected to two levels of MP (non-MP vs. MP 1 min d⁻¹) and two levels of water availability (200 ml d⁻¹vs. 400 ml d⁻¹) and measured three plant traits. MP significantly decreased plant height, total biomass, and root/shoot ratio. There were significant interactions between MP and soil water availability on plant height and root/shoot ratio. These findings imply that MP alone is a stressful factor for P. villosa and MP effects on its growth can be partially mitigated by increased soil water availability, and also suggest that P. villosa may respond to MP in a way that allows plants to survive in the windy semiarid environments.     

Plant productivity, predation, and the abundance of black-tailed jackrabbits in the Chihuahuan Desert of Mexico

The abundance of black-tailed jackrabbits (Lepus californicus) can fluctuate dramatically. We used data from the Chihuahuan Desert to test the relative strength of top-down (predation) or bottom-up (food availability) limiting forces. Predictions for the top-down hypothesis were, 1) a positive relationship between coyote (Canis latrans) and jackrabbit abundance (numerical response) and 2) a positive relationship between percent occurrence of jackrabbits in coyote scats and jackrabbit abundance (functional response). Predictions for the bottom-up hypothesis were, 1) plant productivity is directly related to precipitation, 2) jackrabbit abundance is positively related to precipitation and plant productivity, and 3) changes in abundance of jackrabbits over the reproductive season will be directly related to precipitation and plant productivity. We found a limited numerical response but no functional response of coyotes to jackrabbit abundance. Forb productivity was significantly related to annual precipitation levels (r² = 0.69, p = 0.002). Grass productivity was related to annual precipitation (r² = 0.34, p = 0.028). Jackrabbit abundance (r² = 0.38, p = 0.002) and changes in abundance (r² = 0.73, p < 0.001) were significantly related to precipitation and forb and grass productivity. We conclude that precipitation levels and plant productivity affect jackrabbit abundance more than predation levels.     

Water relations and the effects of clearing invasive Prosopis trees on groundwater in an arid environment in the Northern Cape,South Africa

Several Prosopis species have been introduced into South Africa in the last century and many of them have become invasive. This study investigates the water relations, effects of clearing, and the seasonal dynamics of groundwater use by invasive Prosopis trees. The trees were growing on deep sandy soils in the floodplain of an episodic river in the arid Northern Cape Province of South Africa. Data were collected on tree water uptake, evapotranspiration and water table depth over different seasons. Effects of tree clearing on groundwater were quantified by comparing data from a Prosopis invaded and an adjacent cleared area. Transpiration rates were less than 1.0 mm/d throughout the year and the trees showed structural and physiological adaptations to the combined low rainfall and low water holding capacity of the soils by developing very narrow sapwood areas and by closing their stomata. The trees abstracted groundwater as evidenced by the decline in borehole water levels in the Prosopis stand before the rainy season. Groundwater savings of up to 70 m³/month could be achieved in spring for each hectare of Prosopis cleared. The study suggests that clearing of invasive Prosopis would conserve groundwater in the arid parts of South Africa.     

Invasion by an exotic,perennial grass alters responses of a native woody species in an arid system

Grass growth has been encouraged in arid communities globally to improve grazing. Often exotic species are planted, subsequently spreading beyond their plantings to drastically alter cover and negatively impact native species. We evaluated the effects of the invasive, perennial grass Cenchrus ciliaris on the native woody species Cercidium microphyllum over two growing seasons: 2002 and 2003. To determine if C. ciliaris alters seasonal responses of C. microphyllum, we measured predawn water potential, photosynthesis, and branch sacrifice on C. microphyllum growing with either native cover or C. ciliaris. In 2003 a removal treatment of C. ciliaris was applied to assess responses in C. microphyllum during the summer. In both years, C. microphyllum water potentials closely tracked seasonal rainfall, suggesting dependence on shallow soil water. In 2002, we observed few negative impacts when C. microphyllum grew with C. ciliaris. However, the trees growing with C. ciliaris exhibited greater branch sacrifice, a typical response to drought, which likely complicated interpretation of the treatment effects. Removal of C. ciliaris resulted in increased water potentials and photosynthesis. This study suggests that exotic grasses, such as C. ciliaris, can reduce both water potential and photosynthetic rates in mature woody shrubs, like C. microphyllum, in aridland systems.     

Hydrogeochemistry of Groundwater and Its Suitability for Drinking and Agricultural Use in Nahavand,Western Iran

Groundwater is the major source of drinking water in Nahavand city. However, the groundwater quality at the agricultural areas has been deteriorating in recent years. Ground water quality monitoring is a tool which provides important information for water management and sustainable development of the water resources in Nahavand. Hydrochemical investigations were carried out in an agricultural area in Nahavand, western Iran, to assess chemical composition of groundwater. In this study, 64 representative groundwater samples were collected from different irrigation wells and analyzed for pH, electrical conductivity, major ions, and nitrate. The results of the chemical analysis of the groundwater showed that concentrations of ions vary widely and the most prevalent water type is Ca–Mg–HCO₃, followed by other water types: Ca–HCO₃, Ca–Na–HCO₃, and Na–Cl, which is in relation with their interactions with the geological formations of the basin, dissolution of feldspars and chloride and bicarbonate minerals, and anthropogenic activities. Thirty-seven percent of the water samples showed nitrate (NO₃ ⁻) concentrations above the human affected value (13 mg L⁻¹). The phosphorous (P) concentration in groundwater was between 0.11 and 0.90 mg L⁻¹, with an average value of 0.30 mg L⁻¹, with all of the samples over 0.05 mg L⁻¹. The most dominant class C2-S1 (76.5%) was found in the studied area, indicating that sodicity is very low and salinity is medium, and that these waters are suitable for irrigation in almost all soils. Agronomic practices, such as cultivation, cropping, and irrigation water management may decrease the average NO₃ ⁻ concentration in water draining from the soil zone.     

Comparative hydrological behaviour of two small catchments in semi-arid Zimbabwe

This paper describes and compares the hydrological responses of runoff, soil moisture and groundwater levels to rainfall events for two small semi-arid catchments over a 2-year period. Romwe received 1430 and 756 mm of rainfall in the 19999/00 and 2000/01 season, respectively. Mutangi received 756 and 615 mm of rainfall in the same years. Romwe generated 520 and 102 mm of runoff in the 19999/00 and 2000/01 seasons, respectively, while Mutangi generated 82 and 69 mm of runoff in the same years. The runoff response of the catchments was dominated by a relatively quick response to rainfall and with little or no significant contribution from regional groundwater or ‘old water’ sources. Total soil moisture storage to a depth of 120 cm was higher at Romwe than Mutangi for the entire study period reflecting the differences in the soil types. The groundwater level was closer to the surface and responded more quickly to rainfall at Romwe compared to Mutangi where water levels were between 12 and 16 m below the surface. There was a significant relationship between profile soil moisture and water table level at Romwe and none was observed at all in Mutangi. Significant (p<0.05) monthly rainfall runoff relationships were observed at both Romwe and Mutangi. At Romwe and Mutangi 91% and 76% of the runoff variation was accounted for by rainfall in the 1999/00 season, respectively. The rainfall–runoff relationship were different at Romwe for the two seasons, it was higher in the 1999/00 season than the 2000/01 season when 91% and 49% of the runoff variation was due to rainfall, respectively. The relationships were almost similar at Mutangi during the two seasons.     

An assessment of revegetation treatments following removal of invasive Pennisetum ciliare (buffelgrass)

In semi-arid regions of North America and Australia, Pennisetum ciliare (L.) Link (syn. Cenchrus ciliaris; buffelgrass) is highly invasive and has the potential to introduce fire to fire-intolerant ecosystems. Major efforts to remove P. ciliare continue and it is essential that P. ciliare be prevented from recolonizing. This study investigated potential methods to revegetate with native herbaceous plants: sowing seeds; sowing seeds and mulching; sowing seeds and transplanting seedlings; and relying on natural revegetation from the seedbank. The treatments were applied in 2009 and 2010 at sites in the Sonoran Desert which had undergone P. ciliare removal. Monsoon precipitation was below average each year and seedling emergence and establishment rates were low. There are indications that soil disturbance associated with planting seedlings promoted P. ciliare emergence and increased mortality of brittlebush (Encelia farinosa A. Gray ex Torr.), a common native perennial forb. Addition of mulch may have promoted P. ciliare over native grasses, and seeding had no effect. We did not find competition between herbaceous seedlings. Rather, native and exotic grass seedling densities were positively correlated across sites. Under prevailing conditions, low precipitation appeared to limit herbaceous plant establishment and none of the treatments reduced P. ciliare abundance.     

Differential seedling performance and environmental correlates in shrub canopy vs. interspace microsites

Shrubs in semi-arid ecosystems promote micro-environmental variation in a variety of soil properties and site characteristics. However, little is known regarding post-fire seedling performance and its association with environmental variation in former shrub canopy and interspace microsites. We compared post-fire seeding success and various soil properties important for seedling establishment between shrub canopy and interspace microsites in Wyoming big sagebrush (Artemisia tridentata) plant communities in southeast Oregon, U.S.A. We burned 5, 20 × 20 m sites and established paired canopy and interspace micro-transects seeded with bluebunch wheatgrass (Pseudoroegneria spicata, 193 seeds/m) or crested wheatgrass (Agropyron cristatum, 177 seeds/m). At one year post-fire, seedling density was 69% higher (p = 0.012) for crested wheatgrass (compared to bluebunch wheatgrass) and 75% higher (p = 0.019) for interspace microsites (compared to canopy). However, tiller and leaf area production were over twice as high (p < 0.05) in canopy microsites. Soil color and soil temperature, explained 19–32% of variation in seedling performance metrics. Shrub effects on seeding success are complex and interact with abiotic disturbances, but patterns of increased seedling performance in canopy microsites and their relationships to soil variables may suggest tactics for increasing success of restoration practices.     

NaCl stimulates growth and alleviates water stress in the xerophyte Zygophyllum xanthoxylum

Zygophyllum xanthoxylum is a succulent xerophyte that is recognized as one of the most drought-resistant plants found to date. Our previous investigations have shown that the strong drought resistance of Z. xanthoxylum might be related to the accumulation the Na⁺ in the leaves. In this study, we further investigated the effects of NaCl on the growth and drought resistance of Z. xanthoxylum under water stress. We found that 50 mM NaCl resulted in optimal plant growth and alleviated the deleterious impacts on leaf growth of different osmotic stress levels induced by PEG 6000. The positive effects of NaCl on the drought resistance of Z. xanthoxylum were also further confirmed by soil culture experiments. Ion analysis showed that the Na⁺ absorption of Z. xanthoxylum was specifically induced by water stress in a non-saline medium and that the addition of 50 mM NaCl increased the leaf Na⁺ concentration under water stress. These findings suggest that the positive roles of NaCl in the drought resistance of Z. xanthoxylum might be due to the ability of the plant to accumulate high concentrations of Na⁺ in the leaves and use the Na⁺ as an osmoregulator to cope with water stress.     

Reproductive phenology of Isolatocereus dumortieri (Cactaceae) in semiarid scrub in central Mexico: Effect of rain during the dry season

The giant cactus Isolatocereus dumortieri is a dominant species of the semiarid scrub of central Mexico. Its reproductive period is during the dry season, and it produces essential resources (pollen, nectar and fruits) for a great variety of animal species. We related the production of reproductive structures with water variables in the soil–plant system. The main goal of the present study was to evaluate the consequences of a change in rainfall pattern on the phenology of I. dumortieri. We watered some plants in the dry season to simulate heavy rain events, and to test the hypothesis that water availability during the reproductive season has a negative effect on fruiting and a positive effect on vegetative growth. The seasonality of rain events caused variations in soil water potential and plant osmotic potential, and we found that both variables influenced fruit production. The regression models relating the number of fruits with soil water potential, rainfall and osmotic potential were significant for all three study periods. The highest production of reproductive structures occurred in the driest year (2009), during which there was an ENSO event. Watering did not have a significant effect on osmotic potential or growth in the cacti. However, the watering × time interaction had a negative effect on the number of immature fruits. That is, at the end of the experiment, the plants that received the most water showed a decrease in the number of fruits. Plant growth during the rainy period was significantly greater than during the dry period. The results support the hypothesis that a change in rainfall pattern during the dry season has an effect on fruit production. Vegetative growth, however, occurred only during the wet season.     

Analysis of vegetation seasonality in Sahelian environments using MODIS LAI,in association with land cover and rainfall

Present-day Sahelian vegetation in a highly anthropized semi-arid region is assessed from local to regional scales, through the joint analysis of MODIS LAI (1 km² and 8-day resolutions), daily rainfall, morphopedological and land cover datasets covering the period 2000–2008. The study area is located in northwest Senegal and consists of the “Niayes” and the northwestern “Peanut Basin” eco-regions, characterized by market gardening and rain-fed cultivated crops, respectively. The objectives are i) to analyse at pixel scale LAI time series and their relation to vegetation and soil types, ii) the estimation of phenological metrics (start of season SOS, end of season EOS, growing season length GSL) and their inter-annual variability, iii) to recognize the vegetation responses to rainfall trends (mean annual precipitation, MAP; frequency of rainy events, K; combination of MAP and K, called F).Pixel-scale analyses show that LAI time series 1) describe the actual phenology (agreeing with ground-truth AGHRYMET data), and thus can be used as a proxy for Sahelian vegetation dynamics, 2) are strongly dependent on soil types. Median maps of SOS and EOS suggest an increase of the GSL from Saint-Louis to Dakar, in agreement with both the North-South rainfall gradient and the intensification of agricultural practices around Dakar. Significant correlations (R: 0.64) between annual variation coefficient of LAI and MAP for both herbaceous crops and natural vegetation are highlighted; this correlation is reinforced (R: 0.7) using the rainfall distribution factors K and F. Rainfall thresholds allowing the SOS can be defined for each type of vegetation. These thresholds are estimated at 0–5 mm, 20 mm and 40 mm for natural herbs, herbaceous crops and shrublands, respectively.If previous works revealed the close link between the MAP and the SOS, our results highlight that LAI dynamics are also controlled by rainfall distribution during the Monsoon season. In this study, climatic indicators are proposed for estimating vegetation dynamics and monitoring SOS. Coupling Earth Observation data, such as MODIS LAI, with rainfall data, vegetation and soil information is found to be a reliable method for vegetation monitoring and for assessing the impact of human pressure on vegetation degradation.     

Efficient fog harvesting by Stipagrostis sabulicola (Namib dune bushman grass)

Stipagrostis sabulicola, an endemic grass species of the central Namib Desert, grows on dune fields under conditions of very low annual precipitation punctuated by regular nocturnal fog events. The objective of this study is to determine to what extent S. sabulicola relies on water supply by fog harvesting. The following parameters were monitored: 1) climate, 2) stem runoff, 3) leaf water potential (LWP) and 4) soil water content (SWC). Collected fog water was 5.0 L (liter) per m⁻² leaf surface and therefore a total harvest of 4-5 L per fog event for a medium-sized mound of S. sabulicola. SWC close to a mound increased substantially during a fog event, with SWC at about 2.2% within a mound. LWP of S. sabulicola ranged between −1.7 MPa and −3.5 MPa. On days without fog, LWP was highest during the morning and decreased during the afternoon. No significant decrease of LWP occurred during days following a fog event. The increase of SWC at the plant base during a fog event indicates that fog harvest of S. sabulicola occurs mainly via stem flow with subsequent absorption by the root system and that fog catchment therefore represents a substantial water source for this species.     

Exploring the potential of perennial crops in reducing soil erosion: A GIS-based scenario analysis in southern Tuscany,Italy

Preserving soils is a major challenge in ensuring sustainable agriculture for the future. Soil erosion by water is a critical issue in the Mediterranean regions and usually occurs when high-erosive precipitation is in temporal association with poor vegetation cover and density. Modelling soil erosion risks over large spatial scales suffers from the scarcity of accurate information on land cover, rainfall erosivity and their intra-annual dynamics. We estimated the soil erosion risk on arable land in a Mediterranean area (Grosseto Province, southern Tuscany, Italy) and investigated its potential reduction as a response to the change in intra-annual distribution of land cover due to the increase of perennial forage crops. A GIS-based (R)USLE model was employed and a scenario analysis was performed by setting criteria for raising the performance of perennial forage crops. Statistical data on agricultural crops provided an insight into current intra-annual land cover dynamics. Rainfall erosivity was computed on the basis of 22-year hourly precipitation data. The model was used to: i) quantify the potential soil losses of arable land in the study area, ii) identify those areas highly affected by erosion risks iii) explore the potential for soil conservation of perennial crops, thereby enabling appropriate preventive measures to be identified. The erosion rates, averaged over an area of about 140’000 ha, are estimated to 33.42 Mg ha⁻¹ y⁻¹. More than 59% of the study area was subjected to soil losses higher than 11 Mg ha⁻¹ y⁻¹ (from moderate to severe erosion) and the highest rates are estimated for steep inland areas. Arable land with severe soil erosion rates (higher than 33 Mg ha⁻¹ y⁻¹) represent about 35% of the whole study area. The risk of soil loss by water erosion in the study area is estimated to be reduced on average by 36% if perennial crops are increased in terms of 35% of the total arable land. The soil erosion data produced compared well with the published local and regional data. This study thus provides useful preliminary information for landscape planning authorities and can be used as a decision support tool in quantifying the implications of management policies.     

Representative rainfall thresholds for landslides in the Nepal Himalaya

Measuring some 2400 km in length, the Himalaya accommodate millions of people in northern India and Pakistan, Nepal, Bhutan, and parts of other Asian nations. Every year, especially during monsoon rains, landslides and related natural events in these mountains cause tremendous damage to lives, property, infrastructure, and environment. In the context of the Himalaya, however, the rainfall thresholds for landslide initiation are not well understood. This paper describes regional aspects of rainfall thresholds for landslides in the Himalaya. Some 677 landslides occurring from 1951 to 2006 were studied to analyze rainfall thresholds. Out of the 677 landslides, however, only 193 associated with rainfall data were analyzed to yield a threshold relationship between rainfall intensity, rainfall duration, and landslide initiation. The threshold relationship fitted to the lower boundary of the field defined by landslide-triggering rainfall events is I = 73.90D^− 0.79 (I = rainfall intensity in mm h^− 1 and D = duration in hours), revealing that when the daily precipitation exceeds 144 mm, the risk of landslides on Himalayan mountain slopes is high. Normalized rainfall intensity–duration relationships and landslide initiation thresholds were established from the data after normalizing rainfall-intensity data with respect to mean annual precipitation (MAP) as an index in which N_I = 1.10D^− 0.59 (N_I = normalized intensity in h^− 1). Finally, the role of antecedent rainfall in causing landslides was also investigated by considering daily rainfall during failure and the cumulative rainfall to discover at what point antecedent rainfall plays an important role in Himalayan landslide processes. Rainfall thresholds presented in this paper are generalized so they can be used in landslide warning systems in the Nepal Himalaya.     

A simple method for estimating basin-scale groundwater discharge by vegetation in the basin and range province of Arizona using remote sensing information and geographic information systems

Groundwater is a vital water resource in the arid to semi-arid southwestern United States. Accurate accounting of inflows to and outflows from the groundwater system is necessary to effectively manage this shared resource, including the important outflow component of groundwater discharge by vegetation. A simple method for estimating basin-scale groundwater discharge by vegetation is presented that uses remote sensing data from satellites, geographic information systems (GIS) land cover and stream location information, and a regression equation developed within the Southern Arizona study area relating the Enhanced Vegetation Index from the MODIS sensors on the Terra satellite to measured evapotranspiration. Results computed for 16-day composited satellite passes over the study area during the 2000 through 2007 time period demonstrate a sinusoidal pattern of annual groundwater discharge by vegetation with median values ranging from around 0.3 mm per day in the cooler winter months to around 1.5 mm per day during summer. Maximum estimated annual volume of groundwater discharge by vegetation was between 1.4 and 1.9 billion m³ per year with an annual average of 1.6 billion m³. A simplified accounting of the contribution of precipitation to vegetation greenness was developed whereby monthly precipitation data were subtracted from computed vegetation discharge values, resulting in estimates of minimum groundwater discharge by vegetation. Basin-scale estimates of minimum and maximum groundwater discharge by vegetation produced by this simple method are useful bounding values for groundwater budgets and groundwater flow models, and the method may be applicable to other areas with similar vegetation types.     

Spatial carbon and nitrogen distribution and organic matter characteristics of biological soil crusts in the Negev desert (Israel) along a rainfall gradient

In arid and semi-arid areas biological soil crusts are main contributors to C and N-cycles and the origin of organic matter. Nevertheless systematic studies on the spatial distribution of total organic carbon (TOC) and nitrogen (N) and a characterization of crust organic matter composition are missing. To describe the spatial distribution of TOC and N we examined three soil depths and three relief positions along a steep rainfall gradient. In addition the molecular composition of organic matter was characterized by Pyrolysis-field ionization mass spectrometry.TOC and N concentrations decreased with increasing depth, the effects of the relief followed no clear trend. Surprisingly the amount of TOC and N decreased with increasing rainfall. Stable organic matter compounds were reduced with increasing rainfall. Topcrusts (0–2 mm) showed a relative enrichment in bacteria, as indicated by proportionally larger contents in N-acetylmuramic acid (m/z 167 + 276) than the subcrusts (2–40 mm). These were enrichment in cyanobacteria, as indicated by proportionally larger contents of hexadecadienoic acid (m/z 252).We conclude that the spatial distribution of TOC and N is related to sampling depth and annual precipitation. Organic matter composition and the main biomass contributors in crusts are successfully identified by pyrolysis-field ionization mass spectrometry.     

Effects of simulated storm sizes and nitrogen on three Chihuahuan Desert perennial herbs and a grass

Establishment and growth of three perennial herbs and a small tussock grass were studied in an experiment that provided simulated rainfall of 6 mm week⁻¹ or 25 mm once per month and nitrogen fertilization in combination with the different simulated rainfall regimes. Wild onion, Allium macropetalum, failed to establish in plots receiving 25 mm month⁻¹ simulated rainfall. The perennial composite, Bahia absinthifolia, occurred at higher densities in plots that were not irrigated but there were no differences in biomass in any of the irrigation or fertilization treatments. Desert holly, Perezia nana, failed to establish in nitrogen fertilized plots and developed higher abundance and biomass in plots receiving 25 mm month⁻¹. Nitrogen fertilization had either no effect or an adverse effect on the perennial herbs. The tussock grass, Dasychloa pulchella exhibited highest abundance and biomass with 6 mm week⁻¹ added water plus nitrogen. Since global climate change will affect both rain storm frequency and size and atmospheric nitrogen deposition, the results of this study are applicable to understanding vegetation responses climate change.     

Heliotropium europaeum only germinates following sufficient rainfall to allow reproduction

Regions with Mediterranean climates are seasonally arid and provide a niche for ephemeral species which germinate following discrete and variable rainfall during summer. These species must be able to detect when conditions are suitable for completion of their life cycle. Common heliotrope (Heliotropium europaeum) is one such species. It is considered a weed in its naturalised habitat in southern Australia as it uses resources that could be used by ensuing crops, and is toxic to livestock.We examined common heliotrope's germination responses to temperature and water potential, the effect of simulated rainfall on seedling emergence and plant growth in lysimeters.Fresh seeds of common heliotrope have relatively high optimum temperatures and water potentials for germination (around 35 °C and 0 MPa). Germination percentage of seeds extracted from the soil seed bank varies seasonally. In the soil types and climate of the study area, 24.2 mm is the minimum amount of rainfall resulting in emergence. Evapotranspiration required to reach minimal reproductive output was 19.4 mm.Common heliotrope's germination requirements have been selected to ensure that it will only germinate after sufficient rainfall to allow reproductive output. This is the mechanism through which common heliotrope detects its temporal niche.     

Hydrochemical Characteristics and Sodification of Groundwater in the Shirin Sou, Hamedan, Western Iran

Salinity and sodicity of groundwater are the principal water quality concerns in irrigated areas of arid and semi-arid regions. The hydrochemical characteristics and sodicity of groundwater in the Shirin Sou area, western Iran were investigated in this study by chemical analyses of groundwater samples from 49 wells. Chemical analysis of the groundwater showed that the mean concentration of the cations was in the order: Na⁺ > Ca²⁺ > Mg²⁺ > K⁺, while that for anions was SO₃ ²⁻ > Cl⁻ > HCO₃ ⁻ > NO₃ ⁻. The most prevalent water type is Na–SO₄ followed by water types Na–Cl and Ca–SO₄. The chemical evolution of groundwater is primarily controlled by water–rock interactions: mainly weathering of aluminosilicates, dissolution of sulfate minerals, and cation exchange reactions. Sulfate dissolution and pyrite weathering may both contribute to the SO₄ ²⁻ load of the groundwater. High Na⁺ concentrations in groundwater participate in ion-exchange processes, resulting in the displacement of base cations into solution and raised concentrations in groundwater. The principal component analysis (PCA) performed on groundwater identified three principal components controlling variability of groundwater chemistry. Electrical conductivity, Ca²⁺, Mg²⁺, Na⁺, SO₄ ²⁻, and Cl⁻ content were associated in the same component (PC1) (salinity), most likely linked to anthropogenic activities.