Association of macro- and microfossils in the Vendian (Ediacaran) postglacial successions in Western Mongolia

The Vendian (Ediacaran) beds of the Zavkhan Basin, in the upper part of the Tsagaanolom Formation (<632 ± 14 Ma), yielded a new “Zavkhan” association of algae, microfossils, and problematic organisms, which is established in the series of alternating chert-carbonate shale with phosphorite interbeds. This association is distinct in the predomination of large (250 μm and over) sphaeromorphic microfossils of the genera Tasmanites, Archaeooides, and Leiosphaeridia, whereas acanthomorph acritarchs are represented by rarely found Cavaspina sp. and Tanarium sp. Multicellular algae included fragments of encrusting or foliate thalli with pseudoparenchymatous structure of polygonal cells characteristic of Rhodophyta algae (Thallophycoides sp.), and cordlike thalli of Vendotaenid algae Tyrasotaenia podolica. These layers of siltstone contain imprints of the problematic Vendian macrofossil Beltanelliformis brunsae. In their stratigraphic position, chemostratigraphic data, and fossil assemblage, the “Zavkhan” association can be assigned to the Upper Vendian.     

Early Triassic palynomorphs from Kuraloi Block,Belphar area,Ib-River coalfield,Mahanadi basin,Odisha

A rich assemblage of palynomorphs including miospores and megaspores has been recorded from the bore core no. IBKAN-2 drilled at a depth of 17.80m in Kuraloi Block-A, south-west part of the Ib-River Coalfield, Jharsuguda district, Odisha. The miofloral assemblage recovered shows prominence of cingulate, zonate, taeniate, non-striate and non saccate palynotaxa which indicate an early Triassic age and is comparable with the palynoassemblages of the same age known from the Damodar basin, India. The megaspore assemblage includes eight genera and fifteen species namely, Banksisporites karanpuraensis, B. indicus, B. utkalensis, Banksisporites sp., Barakarella pantii, Barakarella sp., Biharisporites spinosus, Biharisporites sp., Erlansonisporites triassicus, Erlansonisporites cf. erlansonii, Erlansonisporites sp. Jhariatriletes sp., Singhisporites sp., Ramispinatispora sp. and Talchirella trivedii. Records of megaspores are sparse in the early Triassic of Gondwana basins of India. The present study records the first occurrence of miospores and megaspores from the early Triassic of Mahanadi basin besides substantiating the earlier data.     

Smaller foraminifers from type sections of the Bolorian stage,the Lower Permian of Darvaz

Smaller foraminifers from Upper Yakhtashian and Bolorian deposits of the stratotype area (Pamir, Darvaz, Tajikistan) are investigated. Four assemblages are defined. The first assemblage is from Chalaroschwagerina vulgaris-Pamirina darvasica Zone. The second assemblage found in the transitional Yakhtashian-Bolorian beds includes Globivalvulina, Palaeotextulariidae, Hemigordiidae, and Glomospira, associated with the first Pachyphloia and Langella forms. Characteristic taxa of third assemblage from the Misellina (Brevaxina) dyhrenfurthi Zone are Geinitzinidae, Globivalvulina, Palaeotextulariidae, Glomospira, and rare Pachyphloia. The forth assemblage of Hemigordiidae, Pachyphloia, Palaeotextulariidae, Geinitzinidae, Pseudoagathammina is identified in the M. (Misellina) parvicostata Zone. The assemblages were compared with concurrent analogs from China, Japan, and Russia. New species and subspecies Glomospira paleograndis sp. nov., G. darvasica sp. nov., Agathammina darvasica sp. nov., Pachyphloia darvasica sp. nov., Nodosinelloides cubanicus elongatus subsp. nov., and Hemigordius saranensis darvasicus subsp. nov. are described.     

Foraminiferal evidence of basin submergence in part of Sundarban mangrove delta,India

Sundarban is a largest mangrove forest delta developed along the NE-SW direction covering parts of India and Bangladesh. Little work has been done on Indian part of Sundarban in respect of heterogeneity in channel morphology which could be mostly due to the effect of tilting and basin subsidence. These changes might have played a major role on development of high marshes, which offers a congenial environment for survival of Haplophragmoides wilberti, Jadammina macrescens, Trochammina inflata, and Miliammina fusca. These marsh benthonic foraminiferal assemblages provide a direct evidence of recent past sea level changes. To establish the depositional pattern and their effects rendered by merciless changing environment, 11 pit sections have been excavated along three E-W transects from Indian Sundarban. Depth of these pit sections varies from 2 to 3 m. Generally, top 20 cm sediment (in pit section W-1 to W-11) deposited under the intertidal environment, as indicated by the presence of Ammonia tepida. However, sediment below 20 cm in some of the pit sections (W-3 and W-5) exhibits fresh water signatures as indicated by the presence of Charophytes algae. In other pit sections (W-1, W-2, W-6, W-7, W-8, W-9 and W-11), the intertidal assemblage is noticed just above the upper marshes assemblage and vice versa, signifying that depositional environment is in proximity to the mangrove dominated area as indicated by the presence of marsh benthonic foraminiferal assemblage containing T. inflata, H. wilberti, Haplophragmoides sp., J. macrescens and M. fusca. Bottom sediment in most of the pit sections from south to north have different depositional environment with alternate presence of intertidal to subtidal faunal assemblages. The peculiar presence of intertidal assemblage above the upper marshes assemblages in recent sediment points towards the theory of submergence due to relative rise in the sea level. But the effect of relative sea level rise is not uniform throughout the area because of differential subsidence due to varied rate of sediment supply (0.5 to 3.3 cm/year) and eastward tilting of the basin. Based on the upper marshes benthonic foraminiferal assemblage and radiocarbon age (in W-1 at 100 cm ~?150 years age), the average subsidence rate as recorded is approximately 0.3 to 0.5 cm/year. Hence, such depositional sequence conjectures that the Indian part of Sundarban is undergoing a phase of submergence concomitant to basin subsidence.     

Microbiostratigraphy of the Upper Paleocene to Middle Eocene Jahrum Formation in the Folded Zagros Zone,SW Iran

The Jahrum Formation (Upper Paleocene to Middle Eocene) is composed of carbonate and dolomitic carbonate rocks in the Zagros Basin. The Zagros is located at the boundary between the Arabian and Eurasian lithosphere plates and represent the orogenic response to a collision between Eurasia and advancing Arabia during the Cenozoic. The study area is located in the northern part of Kuh-E-Tudej, ~175 km southeast of Shiraz in the Folded Zagros Zone. The Jahrum Formation at Kuh-E-Tudej, with a thickness of 190 m, consists of medium to massive bedded limestone. The following foraminiferal index species are identified in the studied section: Fallotella alavensis, Kathina sp., Miscellanea sp., Lockhartia sp.,Orbitolites shirazeinsis, Nummulites sp., Opertorbitolites sp., Dictyoconus cf. egyptiensis, Orbitolites cf. complanatus, Dictyoconus sp., Coskinolina sp., Somalina stefaninii, Discocyclina sp., Praerhapydionina sp., Coskinolina cf. liburnica, Nummulites cf. globulus, Nummulites cf. aturicus, and Alveolina sp. The age of the studied sediments ranges from Upper Paleocene to Middle Eocene. The microbiostratigraphic studies revealed four biozones based on the foraminifers identified in the studied section.     

Age and palynological characteristic of the Dongning Formation (Eastern Heilongjiang,China)

The taxonomic composition of the palynological assemblage of the Dongning Formation is supplemented. The palynological assemblage corresponds to those from the Lipovtsy Formation (Aptian) in the Razdolnaya Basin of Primorye (Russia) and the Muling Formation (Aptian) in the Jixi Basin of eastern Heilongjiang (China). It is found that the age of the Dongning Formation is Aptian. The palynological assemblage is characterized by dominance of spores of Gleicheniaceae; they are accompanied by spores of Cyatheaceae. The most important feature of the palynological assemblage of the Dongning Formation is the presence of angiosperm pollen (Tricolpites sp., T. micromunus, T. vulgaris, Retitricolpites georgiensis, R. vulgaris, Clavatipollenites hughesii, Quercites sparsus, Fraxiniopollenites variabilis).     

Biostratigraphy and depositional environment of the Miocene limestone bed of Baripada,Mayurbhanj district,Odisha: Foraminiferal,sedimentological and bulk organic geochemical evidences

Baripada Marine Beds (BMB) have been studied extensively in terms of its mega fossil content. However, not much has been discussed about the foraminiferal content and the organic matter assemblage in these beds. The fossiliferous sequence of BMB consists of sandstone, shale and limestone units. The present study is persued on the limestone unit of BMB. Foraminiferal, thin section and rock-eval pyrolysis studies were performed on the 33 surface samples collected from five sections of Jamdapal and Mukurmatia region along the Budhabalang river bank. Foraminifera are less abundant in the samples of Jamdapal, whereas Mukurmatia is comparatively rich. First appearance datum of Globorotalia menardii[ranges since middle Miocene (12.6 Ma); FAD at planktic foraminiferal zone N12] and last appearance datum of Neogloboquadrina continuosa [ranges between early Miocene (23.2 Ma) to late Miocene (8.3 Ma); FAD at planktic foraminiferal zone N4B and LAD at N16] together suggest that the limestone unit was deposited in between 12.6 to 8.3 Ma within the upper Miocene. Also, the association of shallow water benthic foraminifera (Species of Ammonia, Asterorotalia, Bolivina, Buliminella, Cibicides, Challengerella, Criboelphidium, Cribononion, Elphidium, Hanzawaia, Nonionella) and planktic foraminifera (Globigerina falcoensis, Globigerina woodi, Globorotalia menardii, Neogloboquadrina continuosa) together with oyster bank and shark teeth suggest deposition of limestone within well oxygenated, tropical, shallow water, open marine condition (within 40m water depth). Lithological and thin section studies together with global sea level fluctuation history advocate that upper Miocene marine transgression promoted the formation of this unit. Bulk organic geochemical data obtained by the rock-eval pyrolysis studies on selected samples indicate a low total organic carbon (TOC), with low hydrogen index (HI), high oxygen index (OI). The organic facies is characterised by type-IV kerogen with major contribution from near shore terrestrial plants. This also suggests deposition in shallow, oxygenated environments that did not promote significant accumulation and preservation of organic content in sediments.     

New record of coralline algae from the Holocene sediments of Agatti Island,Lakshadweep, India

The present paper examines the coralline algal assemblage recovered from the Holocene sediments of Agatti Island, Lakshadweep, India. The assemblage comprises nine species of coralline algae which include Amphiroa fragilisma, Lithophyllum nitorum, Lithophyllum incrustans, Lithoporella melobesioids, Spongites sp., Porolithon craspedium, Aethesolithon problematicum, Clathromorphum parcum, Melobesoideae gen. et spec. indet. Clathromorphum parcumis an endophytic coralline alga reported for the first time from the Indian subcontinent. Out of these, seven species belong to the family Corallinaceae and the two represent the family Hapalidiaceae of the class Rhodophyceae. Coralline algal association of this area is dominated by Lithophylloideae, Melobesioideae and Mastophoroideae which include encrusting to fragmented growth forms. The associated branching corals are well developed and adapted to shallow, warm water, low turbid, protected lagoonal environment.     

Peri-Gondwana Late Early–Middle Cambrian trilobites from the Kuhbanan Formation in Dahu section,Kerman Province,Iran

Outcropped of the Kuhbanan Formation at Dahu, near Zarand, about 63 km north of Kerman, Iran contains peri-Gondwana trilobites. In this study, 185 trilobite samples including six species and genera were identified and described from Dahu section. This trilobite’s assemblage including Redlichia noetlingi, Redlichia sp., Kermanella kuhbananensis, Kermanella lata lata, Kermanella lata minuta, Iranoleesia pisiformis, and Iranaspis sp. based on occurrence of the trilobite fauna a late Early to Middle Cambrian (Series 2–3) is suggested for this strata. These trilobite fauna help confirm conclusions from recent geological studies that place the Kerman Basin of Iran during the Cambrian.     

Finds of Ediacaran-type fossils in Vendian deposits of the Yudoma Group,Eastern Siberia

It has been established that terrigenous units of mainly carbonate deposits of the Yudoma Group in the Uchur–Maya Region of the Siberian Platform contain an assemblage of Late Vendian Ediacaran-type macrofossils. The assemblage contains species such as Aspidella terranovica, Beltanelliformis brunsae, Nenoxites curvus, and Palaeopascichnus sp., which had widespread geographic distribution. Finds of Ediacaran fossils in fossil-rich deposits of the Yudoma Group have expanded the possibilities of using the classic biostratigraphic approach applied for subdivision and global correlation of carbonate deposits in Late Precambrian paleobasins.     

<Emphasis Type="Italic">Suraqalatia brasieri</Emphasis> n.gen., n.sp. (larger foraminifera) from the Maastrichtian of Sulaimani area in northern Iraq

Suraqalatia brasieri n.gen., n.sp. from the family Dicyclinidae Loeblich and Tappan 1964 occurs on the Maastrichtian carbonate platform of northern Iraq. The new genus is recognizable by its large very compressed conical test, up to 55–70 mm in diameter, to 0.3–1.6 mm in thickness, planspiral cooling having very small proloculus in the initial part and later circular chambers including numerous chamberlets with an agglutinated wall. Suraqalatia brasieri n.gen., n.sp. is associated with textulariids, miliolids and rotaliids as Loftusia elongata Cox, L. morgani Douvillé, Orbitoides medius d’Archiac, O. megaloformis Papp & Kupper, O. gruenbachensis Papp, O. apiculatus Schlumberger, Omphalocyclus macroporus (Lamarck), Siderolites calcitrapoides Lamarck, Sirtina orbitoidiformis Brönnimann & Wirz. The associated macrofauna comprises large and rich giant rudists (Preradiolites sp.), other bivalves (Gryphaea sp. and Glycymeris sp.), gastropods (Acteonella sp.), echinoderms and corals. The fauna indicates shallow marine carbonate platform conditions within the Maastrichtian green house. It is also worth mentioning that the new genus has only been recorded from the Maastrichtian age.     

Refinement of stratigraphy according to the first finds of planktonic species of <Emphasis Type="Italic">Orbulina</Emphasis> and <Emphasis Type="Italic">Praeorbulina</Emphasis> from the Guri Limestone of the Mishan Formation in northwest of Bandar Abbas,South Iran

The Zagros Basin is one of the most universal oil and gas basins that is located in the west to south of Iran and in north of the Arabian Plate. The Guri Member at the bottom of the Mishan Formation, in some areas such as Bandar Abbas hinterland, contains a significant amount of gas. The Bandar Abbas hinterland is located in the southeast of Zagros. The Guri Limestone is the youngest hydrocarbon reservoirs of the Zagros Sedimentary Basin. In this study, a total of 178 samples from the Guri Limestone in the Handun section are investigated for foraminiferal biostratigraphy. The study of foraminifers led to a recognition of 43 genera and 57 species of benthic and planktonic foraminifera. For the first time, planktonic foraminiferal species including Praeorbulina glomerosa, Praeorbulina transitoria, Orbulina suturalis, and Orbulina universa are reported, and based on the identified benthic and planktonic foraminifera taxa, the age of the Guri Member at Handun section is estimated as late Burdigalian to Langhian.     

Mississippian foraminifers from the Onimaru Formation in the Hikoroichi area, South Kitakami Belt, Northeast Japan

Abundant foraminifers were found from the Mississippian Onimaru Formation distributed in the Hikoroichi area of the central part of the South Kitakami Belt, Northeast Japan. They include Brunsia pulchra, Archaediscus sp., Paraarchaediscus? sp., Neoarchaediscus? sp., Palaeotextularia spp., Climacammina, spp., Tetrataxis spp., Haplophragmella sp., Lituotubella? sp., Forschiella sp., Cribrospira sp., Bradyina spp., Janischewskina sp., Endothyra spp., Planoendothyra aljutovica., Endothyranopsis compressa, E. crassa, Omphalotis samarica, Eo-staffella spp., Mediocris breviscula, and several others. The Onimaru foraminiferal fauna is similar to those re-ported from MFZ (Mamet Foraminiferal Zone) 15-16. This supports a late Visean (V3b-V3c) age of the formation, which has been proposed previously by rugose corals.     

New Riphean microbiotas of the Billyakh Group,the north Anabar region (Fomich River basin): To Riphean biostratigraphy of the Siberian platform

Excellently preserved organic-walled and silicified microfossils are first found in the Lower Riphean Ust-Il’ya and Kotuikan formations of the Billyakh Group in the northern slope of the Anabar Uplift (the Fomich River basin). Similar assemblages were previously known only from sections located southward in the Kotuikan River basin, and taxonomic composition of organic-walled microbiotas from the Ust-Il’ya and Kotuikan formations became a corner stone in competitive microphytological models that are based on different approaches. In their composition and general appearance, microbiotas from the Kotuikan and Ust-Il’ya formations in the Fomich River basin are similar to microbiotas reported from the Kotuikan River basin, although northern sections of the above formations characterize deeper sedimentation settings than in localities known before. The Ust-Il’ya and Kotuikan assemblages of organic-walled microfossils include sphaeromorphic Chuaria circularis and Leiosphaeridia, two-layer vesicles the genus Simia, filamentous Plicatidium and Taenitrichoides, and some others. The silicified microbiota from the lower Kotuikan Subformation is largely composed of akinetes of Anabaena-like cyanobacteria Archaeoellipsoides, spherical Myxococcoides grandis, and short trichomes Filiconstrictosus and Orculiphycus representing initial germination stages of Anabaena-like cyanobacterial spores. Acanthomorphic acritarchs known from lithology-similar Lower and Middle Riphean (Mesoproterozoic) formations of Australia and China have not been observed in the Ust-Il’ya and Kotuikan microbiotas, which are probably of older age. The found microbiotas outline substantially wider distribution area of organic-walled and silicified microfossils, supplement microphytological characteristics of Riphean sediments in the Anabar Uplift, provide information on taxonomic composition of microbiotas from a wider spectrum of facies, and specify relationships between Early and Middle Riphean assemblages of microorganisms from different continents.     

Late Carboniferous lycopsids of the Karantrav locality (the Southern Urals)

Numerous plant remains as the imprints of lycopsid strobili and stems and the sporadic imprints of articulate stems were first found in the Upper Carboniferous deposits from the Western Slope of the Southern Urals (Karantrav Settlement area). The brief stratigraphic characteristic of the studied locality is given. The studied plant assemblage is represented by Lepidodendron ophiurus Brongniart, L. vaselgense Anikeeva et O. Orlova, sp. nov., Lepidodendron sp., Lepidostrobus tevelevii O. Orlova, Mamontov et Anikeeva, sp. nov., L. ronnaensis Bek et Oplustil, Knorria sp., Calamites sp., and Mesocalamites ramifer (Stur) Hirmer. In-situ microspores of the Lycospora type were discovered in the sporangia of some strobili of the genus Lepidostrobus. Four lycopsid species (two are new) were described.     

The Bajocian–Bathonian plant fossil from the Hojedk formation,Lenjan section,Kerman, Iran

The Jurassic successions represent a wide distribution in North of the Kerman province. These successions include Ab-Haji, Badamu, and Hojedk formations. The Hojedk Formation contains the plant fossils. The Lenjan section is one of the suitable areas for paleontological studies on the Hojedk Formation. The study section is mostly composed of green sandstone and shale with several interbedded coal veins with different thicknesses. The thickness of the Hojedk Formation is about 200 m in the Lenjan section. In this study, seven genera and 13 species of macro plant fossils were identified and described, including Nilssonia undulata, Nilssonia bozorga, Nilssonia berriesi, Nilssonia sp., Klukia cf. exilis, Klukia exilis, Cladophlebis antarctica, Coniopteris lobata, Coniopteris murrayana, Elatocladus confertus, Podozamites sp., Equisetites sp., and Coniopteris sp. The Bajocian–Bathonian can be attributed to the Lenjan section based on the recognized flora.     

Benthic foraminiferal turnover across the Paleocene–Eocene Thermal Maximum (PETM) event in Eastern Desert,Egypt

Benthic foraminiferal fauna are analyzed quantitatively and qualitatively at three stratigraphic sections in Eastern Desert of Egypt (Serai, Duwi, and Um El Huetat). These sections embrace the Paleocene–Eocene Thermal Maximum (PETM) interval which is represented by the occurrence of five distinctive beds. These beds constitute the Dababiya Quarry Member at the lower part of Esna Formation. The occurrence of them indicates an expanded and relatively continuous record across the P/E boundary. The organic-rich clay layer (bed no. 1 of the Dababiya Quarry Member) marks the start of the PETM event. This bed is characterized by the extinction of all benthic foraminiferal fauna except for the occurrence of rare agglutinated foraminiferal species. The presence of these species indicates an oceanic anoxic event at the sea floor. High concentration of phosphatic contents including fish remains occurred in the middle part of the PETM (bed nos. 2 and 3 of the Dababiya Quarry Member) with the continuous absence of benthic foraminiferal fauna except for few specimens at the top of bed 3. Bed nos. 4 and 5 of the Dababiya Quarry Member represent the upper part of the PETM and the initial stage of sea floor recovery. Low diversity and abundance of benthic foraminiferal taxa occurred within these beds, represented by Valvulineria scrobiculata, Lenticulina midwayensis, Loxostomoides appliane, and Siphogenerinoides eleganta. This phenomenon continues upward during the post-PETM event. The Paleocene velasco-type benthic foraminifera Angulogavelinella avnimelechi and Coryphostoma midwayensis species are extinct within the advent of the PETM event. The benthic foraminiferal assemblages at the studied sections are dominated by midway-type fauna with little representative of velasco-type fauna. The velasco-type species are represented with high abundance at Serai section and with low densities at Um El Huetat section, while at Duwi section, they rarely occurred. This suggests outer neritic-upper bathyal (150–400 m) setting at Serai section and mostly middle-outer neritic (50–150 m) setting at Um El Huetat and Duwi sections.     

Facies analysis of Pleistocene limestones from Neil West Coast Formation,Neil Island,Ritchie’s archipelago of South Andaman,India

Petrographic thin section analysis of the samples collected from the type section of Neil West Coast Formation, situated in the west coast of Neil Island yielded moderately preserved coralline red algae, benthic and planktic foraminifers, coral fragments, echinoid spines and gastropod shells. The coralline red algae are represented by both non-geniculate and geniculate forms. The non-geniculate forms belong to melobesids, lithophylloids and mastophoroides. The geniculate forms are represented by species of Amphiroa, Corallina, and Jania. However, the diversity and abundance of coralline algal forms are less in comparison to the benthic foraminifers those are represented by Amphistegina, Neorotalia, Ammonia, Elphidium, Operculina, Assilina, Amphisorus and texularids. Planktic foraminifers like Globigerinoides and other biogenic components viz., gastropod shells, echinoid spines and coral fragments are also common. A foraminiferal-algal grainstone facies has been recognized as observed in the field as well as in thin section analysis. The overall assemblage of the biogenic components and facies analysis indicate intertidal to near shore environment of deposition with high energy condition and increased hydrodynamic activity.     

A candidate for the Global Stratotype Section and Point at the base of the Serpukhovian in the South Urals,Russia

The Verkhnyaya Kardailovka section is one of the best candidates for the GSSP (Global Stratotype Section and Point) at the base of the Stage (Mississippian). For boundary definition, the first appearance of the conodont Lochriea ziegleri Nemirovskaya, Perret et Meischner, 1994 in the lineage Lochriea nodosa (Bischoff, 1957)?L. ziegleri is used. L. ziegleri appears in the Venevian Substage somewhat below the base of the Serpukhovian in the Moscow Basin. The position of the FAD of L. ziegleri within the Hypergoniatites?Ferganoceras Genozone is confirmed and lies between 19.53 and 19.63 m above the section’s base. Before 2010, deep-water stylonodular limestone containing the boundary in unnamed formation C at Kardailovka was well exposed but only 3 m of Viséan strata cropped out immediately below. Recent trenching exposed another 10 m of underlying Viséan carbonates in formation C and older Viséan siliciclastics and volcanics in unnamed formation B. The contact between formation B and underlying crinoidal limestones in unnamed formation A representing the middle Viséan Zhukovian (Tulian) regional Substage was excavated. The boundary succession, situated in the Magnitogorsk tectonic zone above the Devonian Magnitogorsk arc and Mississippian magmatic and sedimentary rift succession, was deposited west of the Kazakhstanian continent during closure of the Ural Ocean. In the lower part of the section, Viséan tuffaceous siliciclastics and volcanics of formation B record rapid deepening after deposition of neritic middle Viséan crinoid lime grainstone of formation A and subsequent subaerial exposure. The overlying condensed upper Viséan to Serpukhovian succession in formation C comprises deep-water limestone deposited in a sediment-starved basin recording minor turbidite influx and carbonate-mound development. The δ¹³C_carb plot shows a positive shift of 1‰ V-PDB (from +2 to +3‰) between 17.0 and 17.75 m (3.05 and 1.97 m below FAD L. ziegleri). The δ¹⁸O_apatite graph displays a prominent upward shift from 19.9 to 21.1‰ V-SMOW (at 19.15 to 19.51 m) in the nodosa Zone below FAD of Lochriea ziegleri.