Dienerian (Early Triassic) ammonoids from the Candelaria Hills (Nevada, USA) and their significance for palaeobiogeography and palaeoceanography

A well-preserved ammonoid fauna of Early Dienerian age has long been known from the lower portion of the Candelaria Formation in the old Candelaria silver mining district in Mineral and Esmeralda Counties, Nevada, but for a number of reasons, this fauna has never been studied in detail nor illustrated. Previous authors assigned this ammonoid fauna to the Early Dienerian Proptychites candidus Zone of Canada. In reality, it more closely resembles the Tethyan faunas than the higher palaeolatitude Canadian faunas, thus indicating the presence of some degree of equatorial faunal exchange between opposite sides of the Panthalassic Ocean during Early Dienerian time. It also indicates the onset of a provincialism, which contrasts with the cosmopolitan Griesbachian faunas. A rigorous taxonomic analysis of the Candelaria fauna allows us to differentiate the following ten species, which include two new species and one new genus (Mullericeras nov. gen.) belonging to the new family Mullericeratidae: Ambites lilangensis (Krafft, 1909), Ambites aff. radiatus (Brühwiler, Brayard, Bucher and Guodun, 2008), Ussuridiscus sp. indet., ??Koninckites?? aff. kraffti Spath, 1934, Mullericeras spitiense (Krafft, 1909), Mullericeras fergusoni nov. sp., Mullericeras sp. indet., Proptychites haydeni (Krafft, 1909), Proptychites pagei nov. sp., Vavilovites sp. indet. and Parahedenstroemia kiparisovae Shigeta and Zakharov, 2009. This Early Dienerian fauna correlates with the Ambites fauna known from the base of the Ceratite Marls in the Salt Range and from the base of the ??Meekoceras?? beds in Spiti (northern Gondwanian margin). The fauna also permits the precise dating of a shelfal anoxic episode on the equatorial North American margin. This anoxic event correlates in time with similar palaeoceanographic changes in the southern Tethys, which indicates that the Early Triassic biotic recovery was at least partly shaped by such discrete, short events rather than by pervasive and lingering adverse environmental conditions.     

Early Cretaceous (Valanginian) sea lilies (Echinodermata, Crinoidea) from Poland

Valanginian strata in central epicratonic Poland have recently yielded crinoids, not previously recorded from the area. The fauna comprises isocrinids (Balanocrinus subteres, B. gillieroni, “Isocrinus?” lissajouxi), millericrinids (Apiocrinites sp.) and comatulids (Comatulida indet.). For comparison, a few samples of isocrinids from Valanginian strata of Hungary (Tethyan province) were also analysed. The isocrinids, cyrtocrinids and roveacrinids (sensu Rasmussen 1978 inclusive of Saccocoma sp.) were already known from the Valanginian of the southernmost Tethyan regions of Poland (Pieniny Klippen Belt and Tatra Mountains). The current study demonstrates their occurrence in central epicratonic Poland, and suggests that many Jurassic to Cretaceous stalked crinoid taxa (mainly isocrinids) predominated in the shallow-water settings of this area. Thus, the hypothesis of migration (at least from mid-Cretaceous onwards) to deep-water areas, as a response to an increase of the number of predators during the Mesozoic marine revolution, seems not to be universally applicable.     

Toarcian–Aalenian (Early–Middle Jurassic) radiolarian fauna from the Los Molles Formation,Neuquén Basin,Argentina: Taxonomy and paleobiogeographic affinities

This paper reports the occurrence of a Toarcian–Aalenian (Early–Middle Jurassic) radiolarian fauna in the Los Molles Formation, Neuquén Basin, Argentina, as well as comments on its paleobiogeographic affinities. The micropaleontologic analysis was carried out in fine-grained rocks from a turbiditic section of the Los Molles Formation. These samples were first chemically processed using only hydrogen peroxide (H₂O₂), and afterward treated with acetic (CH₃COOH) and hydrofluoric (HF) acids. The first chemical procedure permitted the recovery of only few spongy spumellarians, while the second one enabled to recover more diversified radiolarian assemblages. In general, the studied fauna presents low diversity and abundance, with a strong dominance of spumellarians over nassellarians. The fauna is composed by the genera Paronaella, Homoeoparonaella, Praeconocaryomma, Archaeocenosphaera, Orbiculiformella, Praeparvicingula, and some unidentified spumellarians and nassellarians. According to paleobiogeographic models, the studied Toarcian–Aalenian fauna presents a mid to high latitude affinity. It is possible to infer from those data a bipolar distribution of some taxa, such as Praeparvicingula and probably Praeconocaryomma, between the Northern and Southern hemispheres since the Early Jurassic (Toarcian).     

A neoselachian shark fauna from the Late Cretaceous of Senegal

A selachian fauna is described for the first time from the Late Cretaceous (Campanian–Maastrichtian) of Senegal. So far, the Campanian Paki Formation has only yielded a single tooth of Rhombodus sp. whereas the Cap de Naze Formation has yielded a more diverse fauna including juvenile Cretalamna cf. Cretalamna biauriculata, Serratolamna serrata, Carcharias cf. Carcharias heathi, ?Carcharias sp., Squalicorax pristodontus, Schizorhiza stromeri, Parapaleobates sp., Rhombodus binkhorsti and Rhombodus andriesi. Teeth of juvenile Cretalamna largely dominate the assemblage. Such an assemblage confirms a Late Maastrichtian age for the unit 3 in the Cap de Naze Formation. The assemblage, although composed of cosmopolitan taxa, is similar to the contemporaneous selachian assemblage from the phosphates of Morocco.     

Soft-shelled turtles (Trionychidae) from the Bissekty Formation (Late Cretaceous: late Turonian) of Uzbekistan: Shell-based taxa

In this paper we describe previously unpublished trionychid turtle material, consisting of numerous shell fragments, from the Late Cretaceous (late Turonian) Bissekty Formation of the Dzharakuduk locality in Uzbekistan. This material is assigned to two shell-based taxa: Aspideretoides cf. riabinini and “Trionyx” cf. kansaiensis. The material which cannot be confidently attributed to these two taxa is identified as Trionychidae indet. In addition to these shell-based trionychid taxa, the Dzharakuduk turtle assemblage includes two skull-based taxa of trionychids (Khunnuchelys kizylkumensis and Trionychini indet.). The trionychids from the Bissekty Formation are most similar to trionychids from the younger (Santonian – early Campanian) Bostobe Formation of Kazakhstan, represented by three shell-based taxa (Aspideretoides riabinini, Paleotrionyx riabinini and “Trionyx” kansaiensis), and one skull-based taxon (Khunnuchelys sp.). We provide an improved understanding of the subtle similarities and differences between four closely related Cretaceous turtle assemblages of Middle Asia and Kazakhstan.     

Non-marine gastropods from the Cretaceous–Paleogene transition in the Pingyi Basin,eastern China

Our results present a taxonomic and palaeoecological study on non-marine gastropods from the latest Cretaceous to Paleocene deposits of the Pingyi Basin, Shandong Province, eastern China. These gastropods are systematically described: three species belonging to three genera including Physa dongtaiensis Gu, 1989, Hydrobia datangensis Yü, 1977, and the newly established species Cyathopoma pingyiensis sp. nov.; two indeterminata genera and species including Truncatelloidea gen. et sp. indet., and Pomatiopsidae gen. et sp. indet. Among them, Truncatelloidea gen. et sp. indet. and Cyathopoma pingyiensis sp. nov. are the dominant species with the longest record. Cyathopoma pingyiensis, sp. nov. is the earliest representative of this widely distributed Asian extant genus. Sedimentological facies analysis of the gastropod-bearing beds suggested that Truncatelloidea gen. et sp. indet. lived in a small pond with a river inlet and a shallow lake, while Ph. dongtaiensis, ?Pomatiopsidae gen. et sp. indet, and ?H. datangensis only thrived in the shallow lake. Cyathopoma pingyiensis sp. nov. inhabited the land area around the shallow lake. Our results showed that no significant species change of the gastropod fauna across the K/Pg (Cretaceous/Paleogene) boundary was observed in the Pingyi Basin.     

Neoselachians and Chimaeriformes (Chondrichthyes) from the latest Cretaceous–Paleogene of Sierra Baguales,southernmost Chile. Chronostratigraphic,paleobiogeographic and paleoenvironmental implications

This paper discusses a well-represented fossil record of cartilaginous fishes (Chondrichthyes) from southern South America. The recovered samples allow the recognition of three assemblages with chronostratigraphic and paleogeographic value: i) typical Maastrichtian sharks and rays with affinities to eastern Pacific fauna, including the taxa Ischyrhiza chilensis, Serratolamna serrata, Centrophoroides sp. associated to Carcharias sp., and Dasyatidae indet.; ii) a scarce reworked assemblage of Paleocene–Early Eocene age including the taxa Otodus obliquus and Megascyliorhinus cooperi; iii) a rich assemblage with reworked taxa of Early to Middle Eocene age, together with autochthonous deposited Middle to Late Eocene taxa with close affinities to paleoichthyofaunas recovered from the North Atlantic, represented by Carcharias ‘hopei’, Odontaspis winkleri, Carcharoides catticus, Macrorhizodus praecursor, Carcharocles auriculatus, Striatolamia sp., Striatolamia macrota, Hexanchus agassizi, Notorhynchus sp., Myliobatis sp., Abdounia sp., Pristiophorus sp., Squatina sp., cf. Rhizoprionodon sp., Ischyodus sp., and one new species, Jaekelotodus bagualensis sp. nov. The studied samples include for the first time taxa with well established chronostratigraphic resolutions as well as taphonomic information that help clarifying the age of the fossil-bearing units. In addition, they provide relevant information about the evolution of the Magallanes (=Austral) Basin from the Upper Cretaceous to the Paleogene, suggesting a probable connection with the Quiriquina Basin of south-central Chile during the latest Cretaceous. Finally, the studied assemblages indicate a latitudinal pattern of distribution that provides valuable data on the environmental evolution and temperature of southern South America during the Paleogene.     

Microvertebrates from the Wadhurst Clay Formation (Lower Cretaceous) of Ashdown Brickworks,East Sussex,UK

Ashdown Brickworks, near Bexhill, East Sussex, has produced a large number of vertebrate fossils from the Wadhurst Clay Formation, part of the Wealden Supergroup (Hastings Group; Valanginian; Lower Cretaceous). Here we describe the microvertebrate fauna of the ‘conglomerate bed’, representing a rich sample of taxa. While most of the recovered teeth and bones are abraded, some heavily, most can be identified to species level. The taxa include four species of hybodont sharks (Egertonodus basanus, Planohybodus ensis, Polyacrodus parvidens, P. brevicostatus), three taxa of bony fishes (an unidentified Lepidotes-like semionotiform, the pycnodontiform Ocloedus, and an albuliform), three taxa of crocodyliforms (the goniopholid Hulkepholis, a bernissartiid, and the atoposaurid Theriosuchus), and the theropod dinosaurs Baryonyx and an allosauroid. Sediments of the Wadhurst Clay Formation as a whole indicate freshwater to very slightly brackish-water environments of deposition, and the mainly aquatic time-averaged mixture of fishes and tetrapods recovered from the ‘conglomerate bed’, together with isolated terrestrial species, confirms this interpretation.     

Early Berriasian ammonites from Shal, Talesh region (NW Alborz Mountains, Iran)

An early Berriasian (Berriasella jacobi Zone) ammonite fauna is described for the first time from the Alborz Mountains in northwest Iran. It has been collected from a section located near the village of Shal (Talesh region); in addition to rare phylloceratids, lytoceratids and Neolissoceras, the majority of ammonites belong to the neocomitid subfamily Berriasellinae. With the exception of a new genus and species, Taleshites fuersichi, these taxa are common in European and North African Tethyan successions. Associated calpionellids confirm the early Berriasian age of the ammonite-bearing levels.     

Arctic ground squirrels of the mammoth-steppe: paleoecology of Late Pleistocene middens (∼24 000–29 450 14C yr BP), Yukon Territory,Canada

This paper presents paleoecological analyses of 48 fossil arctic ground squirrel (Spermophilus parryii) middens (nests and caches) recovered from ice-rich loess sediments in the Klondike region of west-central Yukon Territory. AMS radiocarbon dates and stratigraphic association of middens with Dawson tephra (∼25 300 ¹⁴C yr BP), indicate these paleoecological data reflect the onset of glacial conditions of early Marine Isotope Stage (MIS) 2 and terminal MIS 3 (∼24 000–29 450 ¹⁴C yr BP). Plant macrofossils include at least 60 plant taxa, including diverse graminoids (Poa, Elymus trachycaulus, Kobresia myosuroides), steppe forbs (Penstemon gormanii, Anemone patens var. multifida, Plantago cf. canescens), tundra forbs (Draba spp., Bistorta vivipara), dwarf shrubs (Salix cf. arctica, S. cf. polaris), sage (Artemisia frigida) and rare trees (Picea mariana). Many of these taxa identified in the middens represent the first recorded fossils for these plants in Eastern Beringia and add to our knowledge of the floristic composition of Pleistocene vegetation and biogeography in this region. Fossil beetles include typical members of the Eastern Beringian steppe–tundra fauna (Lepidophorus lineaticollis and Connatichela artemisiae) and others suggesting predominantly dry, open habitats. Cache forage selection is suggested by some plant taxa which were particularly frequent and abundant in the middens (Bistorta vivipara, Kobresia myosuroides, Ranunculus spp., Potentilla, Erysimum cf. cheiranthoides, Poa, Carex and Draba). Factors such as proximity of vegetation to burrows and abundance of fruits and seeds per plant were probably important in cache selection. Glacial conditions enabled arctic ground squirrels to form widespread and dense populations in regions such as the Klondike in which they are rare or absent at present. This fossil midden record supports previous hypotheses that suggest arctic ground squirrels evolved in and are well-adapted to the open, steppe–tundra vegetation, loessal soils and glacial climates of the mammoth-steppe biome.     

A new early Cambrian bradoriid (Arthropoda) assemblage from the northern Flinders Ranges,South Australia

A new assemblage of early Cambrian bivalved arthropods (Bradoriida) is described from the Arrowie Syncline in the northern Flinders Ranges, South Australia. The well preserved, largely endemic fauna comprises a total of six taxa (including five new species): Jiucunella phaseloa sp. nov., Jixinlingella daimonikoa sp. nov., Mongolitubulus anthelios sp. nov., Neokunmingella moroensis sp. nov., Phasoia cf. spicata (Öpik, 1968), and Sinskolutella cuspidata sp. nov. This assemblage is derived from a carbonate sedimentary package representing a high energy, shallow water archaeocyath-Renalcis biohermal facies of Terreneuvian, Stage 2 age which transitions up-section to a more restricted, low energy, intra-shelf lagoonal environment that correlates with a Cambrian Series 2, Stage 3 age. The new taxon J. phaseloa sp. nov., has a first appearance datum (FAD) in shallow water biohermal facies of the Hideaway Well Member of the Wilkawillina Limestone at a level 47 m below the FAD of Pelagiella subangulata which is taken to approximate the base of Series 2, Stage 3 in South Australia. Along with Liangshanella circumbolina, this makes J. phaseloa sp. nov. amongst the oldest bivalved arthropods in South Australia and potentially greater Gondwana. The presence of 25 bradoriid taxa from the early Cambrian of South Australia suggests East Gondwana represents a major centre of origin for the Bradoriida.     

A Changhsingian (Late Permian) brachiopod fauna from Son La, northwest Vietnam

A small brachiopod fauna is described from the upper part of the Yenduyet Formation near Son La, northwest Vietnam. The fauna includes the following species: Peltichia kwangtungensis (Zhan), Acosarina minuta (Abich), Rhipidomella hessensis King, Schuchertella cf. cooperi Grant, Derbyia sp., Waagenites soochowensis (Chao), Spinomarginifera chenyaoyanensis Huang, and a Marginiferinae gen. and sp. indet. The fauna is most closely correlated with brachiopods from the lower Changhsingian Stage of South China and the lyttoniid fauna from the Huai Tak Formation of northwest Thailand. In view of these correlations, an early Changhsingian (Late Permian) age is proposed for the Son La brachiopod assemblage.     

Middle Eocene calcareous algae from Southwestern Kachchh,Gujarat

The Fulra limestone (middle Eocene) exposed in the areas around Jhadwa and Harudi villages, southwestern, Kachchh reveals presence of a rich assemblage of calcareous algae belonging to the Chlorophyceae and Rhodophyceae along with abundant foraminifera. In the present paper, eight species belonging to eight genera of calcareous algae are described. These include Dissocladella longijangensis, Sporolithon keenani, Corallina crossmanni, Arthrocardia sp. Misra et al. 2001, Lithothamnion ishigakiensis, Melobesioideae gen. et spec. indet. 1, Melobesioideae gen. et spec. indet. 2 and Lithoporella melobesioides. Out of these, one taxon belongs to the family Dasycladaceae. Among the remaining taxa, one taxon to the family Sporolithaceae, three taxa to the family Corallinaceae and three to the family Hapalidiaceae. Two coralline species, Corallina crossmanni and Lithothamnion ishigakiensis, are recorded for the first time from India. Another species (Dissocladella longijangensis), though known from other areas of India, is new to the study area.     

Taxonomy and palaeoecology of thoracican cirripedes (Crustacea) from a Campanian rocky shoreline at Ivö Klack,southern Sweden

An abundant and diverse fauna of thoracican cirripedes is described on the basis of over 3000 isolated valves collected from a single lens of material representing in the order of a few thousand years of Campanian time (c. 80 Ma), from sediment formed on and close to a rocky shoreline at Ivö Klack in southern Sweden. At least twenty-eight species are present in the fauna, belonging to scalpelliform and basal sessile taxa. Two families (Myolepadidae and Titanolepadidae) and four genera are new (Myolepas, Bosquetlepas, Levelepas and Ivoelepas), as are seven species (Titanolepas spinifer, Levelepas roeperi, Ivoelepas nielseni, Arcoscalpellum scaniensis, Pollicipes vansyoci and P. (?) striatum). The palaeoecology is discussed with reference to extant cirripede faunas from rocky coastal environments. It is concluded that, although many taxa lived subtidally, the pollicipedids and pollicipedid-like forms (Capitulum, Pollicipes and Myolepas, respectively) lived in the intertidal zone, attached to rocks in a high-energy environment. Pachydiadema, a large basal balanomorph species, was adapted to living attached to boulders in the intertidal zone. The remarkable diversity significantly exceeds that of any known fossil thoracican assemblage, and is unknown in similar habitats at the present day, possibly challenging Darwin's famous quip that we currently live in the ‘Age of Barnacles’.     

Ammonites (Phylloceratina,Lytoceratina and Ancyloceratina) and organic-walled dinoflagellate cysts from the Late Barremian in Boljetin,eastern Serbia

Late Barremian ammonite fauna from the epipelagic marlstone and marly limestone interbeds of Boljetin Hill (Boljetinsko Brdo) of Danubic Unit (eastern Serbia) is described. The ammonite fauna includes representatives of three suborders (Phylloceratina, Lytoceratina and Ancyloceratina), specifically Hypophylloceras danubiense n. sp., Lepeniceras lepense Rabrenović, Holcophylloceras avrami n. sp., Phyllopachyceras baborense (Coquand), Phyllopachyceras petkovici n. sp., Phyllopachyceras eichwaldi eichwaldi (Karakash), Phyllopachyceras ectocostatum Drushchits, Protetragonites crebrisulcatus (Uhlig), Macroscaphites perforatus Avram, Acantholytoceras cf. subcirculare (Avram), Dissimilites cf. trinodosus (d'Orbigny) and Argvethites? sp. The taxonomic composition and percent abundance of the identified ammonites indicate that their taxa are predominantly confined to the Tethyan realm. Ammonites with smooth and slightly sculptured shells predominate among the studied fauna. The ammonite-bearing succession from Boljetin represents the lower part of the Upper Barremian, ranging in ammonite zonation from the Toxancyloceras vandenheckei Zone to the lower part of the Imerites giraudi Zone. The associated organic-walled dinoflagellate cysts confirm the Late Barremian age of the ammonite-bearing levels.     

Dinosaur eggshells from the Santonian Milk River Formation of Alberta,Canada

The North American fossil record of dinosaur eggshells for the Cretaceous is primarily restricted to formations of the middle (Albian–Cenomanian) and uppermost (Campanian–Maastrichtian) stages, with a large gap in the record for intermediate stages. Here we describe a dinosaur eggshell assemblage from a formation that represents an intermediate and poorly fossiliferous stage of the Upper Cretaceous, the Santonian Milk River Formation of southern Alberta, Canada. The Milk River eggshell assemblage contains five eggshell taxa: Continuoolithus, Porituberoolithus, Prismatoolithus, Spheroolithus, and Triprismatoolithus. These ootaxa are most similar to those reported from younger Campanian–Maastrichtian formations of the northern Western Interior than they are to ootaxa reported from older middle Cretaceous formations (i.e., predominantly Macroelongatoolithus). Characteristics of the Milk River ootaxa indicate that they are ascribable to at least one ornithopod and four small theropod species. The taxonomic affinity of the eggshell assemblage is consistent with the dinosaur fauna known based on isolated teeth and fragmentary skeletal remains from the formation, although most ornithischians and large theropods are not represented by eggshell. Relative to the Milk River Formation eggshell, similar oospecies occurring in younger Cretaceous deposits tend to be somewhat thicker, which may reflect an increase in body size of various dinosaur lineages during the Late Cretaceous.     

Shallow marine ostracode turnover in response to environmental change during the Paleocene–Eocene thermal maximum in northwest Tunisia

Two outcrop sections spanning the Paleocene–early Eocene boundary in the Sidi Nasseur–Wadi Mezaz area in northwest Tunisia provided rich ostracode assemblages, yielding 26 species of which three are newly described: Reymenticosta bassiounii, Reymenticosta nasseurensis and Buntonia? tunisiensis. The recorded ostracode fauna and associated foraminifera reflect deposition in a coastal to inner neritic environment. Many of the recorded taxa have a wide geographic distribution throughout the Middle East and North Africa. A correspondence is also observed with West African faunas, especially in the early Eocene fauna. These taxa seem to have originated in West Africa during the Paleocene and migrated northwards during the late Paleocene to early Eocene. Sea-level change and decrease in oxygenation associated with the Paleocene–Eocene thermal maximum (PETM) caused the local disappearance of the South Tethyan Paleocene fauna represented by Paracosta kefensis (morphotype-A), Paracosta aff. paleomokattamensis, Paracypris sp. B Esker, Loxoconcha saharaensis, Buntonia sp. 3 Donze et al., Protobuntonia nakkadii, and probably Reymenticosta bassiounii and R. nasseurensis. Simultaneously, a new but poorly diverse Afro-Tethyan fauna, mainly represented by Alocopocythere attitogonensis and Buntonia? tunisiensis, settled in the studied part of the basin. After the PETM, diversity increased again as various taxa (e.g. Bairdia aegyptiaca, Reticulina lamellata and Aegyptiana duwiensis) (re)appeared. Although detailed records across the P/E boundary are still sparse, it appears that the PETM exerted significant influence on the paleobiogeography and composition of Tethyan ostracode faunas.     

New record of coralline algae from the Holocene sediments of Agatti Island,Lakshadweep, India

The present paper examines the coralline algal assemblage recovered from the Holocene sediments of Agatti Island, Lakshadweep, India. The assemblage comprises nine species of coralline algae which include Amphiroa fragilisma, Lithophyllum nitorum, Lithophyllum incrustans, Lithoporella melobesioids, Spongites sp., Porolithon craspedium, Aethesolithon problematicum, Clathromorphum parcum, Melobesoideae gen. et spec. indet. Clathromorphum parcumis an endophytic coralline alga reported for the first time from the Indian subcontinent. Out of these, seven species belong to the family Corallinaceae and the two represent the family Hapalidiaceae of the class Rhodophyceae. Coralline algal association of this area is dominated by Lithophylloideae, Melobesioideae and Mastophoroideae which include encrusting to fragmented growth forms. The associated branching corals are well developed and adapted to shallow, warm water, low turbid, protected lagoonal environment.     

Cambrian trace fossils of the Cruziana ichnofacies from the Bikaner-Nagaur Basin,north western Indian Craton

The Marwar Supergroup of the Bikaner-Nagaur Basin is composed of sediments deposited from the late Neoproterozoic (Ediacaran) to Upper Cambrian. The Nagaur Sandstone Formation of the Nagaur Group (uppermost division of the Marwar Supergroup) preserves trace fossils significant for establishing Early Cambrian biostratigraphic zones and depositional facies. Fifteen ichnospecies (and eight ichnogenera) identified in the Nagaur Sandstone Formation include “Treptichnus” pedum, Cruziana cf. tenella, Cruziana isp., Diplichnites ispp. A, B, and C, Gyrophyllites isp., Lockeia isp., Merostomichnites isp., Monomorphichnus gregarius isp. nov., Monomorphichnus isp., Planolites isp., Psammichnites isp., Rusophycus bikanerus isp. nov., Rusophycus cf. carbonarius, Rusophycus isp. and radial trace fossils.These trace fossils belong to ethological categories pascichnia, repichnia, cubichnia, and fodinichnia and represent arthropod and worm-like burrowing biota. The assemblage and a regional comparison with contemporaneous trace fossils in the eastern Gondwanan realm suggest that the sequence in the study area belongs to the Cruziana tenella Ichnozone and to Stage 2 (upper part of Terreneuvian), however the Middle Cambrian is not excluded. The trace fossil assemblage belongs to the archetypal Cruziana ichnofacies. Cross bedded sandstone, mud cracks and rainprints in the ichniferous strata of the Nagaur Sandstone Formation indicate deposition in an intertidal sand flat with channels that was exposed episodically.