Solemys (Chelonii,Solemydidae) remains from the Maastrichtian of Pyrenees: evidence for a semi-aquatic lifestyle

We report on a turtle from the Mina Esquirol site (Vallcebre basin), a new locality of early Maastrichtian age in the south-eastern Pyrenees. Fossils were located in the basal Tremp Formation, which was deposited in a littoral marsh. The material consists of a cast of a carapace including peripheral fragments and partial neural plates. The carapace exhibits a vermiculate ornamentation that is characteristic of genus Solemys and a histological bone structure similar to that of terrestrial taxa. However, taphonomic data indicates little transport and a short biostratinomic history, whereas palaeontological and sedimentological context indicates that the specimen was preserved in a shallow brackish water environment. Based on this taphonomic and sedimentological evidence, we suggest that at least some species of genus Solemys had a lifestyle similar to extant fresh or brackish water turtles (terrapins) and that the histological evidence alone for a terrestrial lifestyle is misleading.     

The chronology of dinosaur oospecies in south-western Europe: Refinements from the Maastrichtian succession of the eastern Pyrenees

The Late Cretaceous deposits of the continental Tremp Formation in the Vallcebre Syncline (South-eastern Pyrenees) provide an extensive egg record of dinosaurs. The parataxonomical study and analysis of multiple eggshell samples, the precise stratigraphical control of several sections and the time calibration of the abundant egg levels enable the establishment of a robust oospecies succession. The successive occurrence of three megaloolithid oospecies (Megaloolithus siruguei-Megaloolithus mamillare-Megaloolithus sp.) in the Early and Late Maastrichtian is well correlated with the magnetic polarity time scale throughout the chron 31. The replacement of Megaloolithus siruguei with Megaloolithus mamillare occurs around the reversal of chrons 31r-31n. A comparison with oospecies successions from Arc basin localities (France) allows the age calibration for such oospecies replacement to be confirmed. This age refinement implies that some of the stated boundaries for the proposed oospecies assemblages may change and that an in-depth revision of the age and magnetic calibration of some south European egg localities is required.     

Cretaceous–Tertiary tectonic inversion of the Cotiella Basin (southern Pyrenees,Spain)

The Cotiella Nappe includes one of the most important Mesozoic basins of the southern Pyrenees, which was subsequently inverted during the Tertiary compression. The Late Cretaceous Cotiella Basin is here interpreted as the western sector of the Cretaceous Cotiella-Bóixols basin (100᎜ km wide), located in the central part of the southern Pyrenees. The present-day complex structure of the Cotiella Nappe is the result of the inversion process, linked to the emplacement of basement thrust sheets of the Axial zone. In its western sector, the Cotiella Nappe consists of several superimposed thrust sheets, with complex geometry, becoming simpler towards the east, with a single thrust surface and smaller displacements. The Cotiella-Bóixols basin underwent strong subsidence during the Early Cretaceous at its eastern sector, and its depocentre migrated westward during the Late Cretaceous. The reconstruction of the sedimentary basin to the pre-compressional stage shows that during the Mesozoic the Cotiella-Bóixols basin was located to the south of a basement high, which later became the Pyrenean Axial Zone. From a balanced cross section, it can be inferred that the Cotiella, north-verging extensional system was connected with the north-Pyrenean rift by means of a 10-km deep horizontal detachment. The compressional Tertiary detachment within the upper crust was shallower than the extensional detachment, and individualised four basement thrust sheets, which form the Axial Zone antiform.     

Identification of an underfilled foreland basin system in the Upper Devonian of the Central Pyrenees: implications for the Hercynian orogeny

A foreland basin succession has been identified in the Frasnian of the Central Pyrenees. This succession comprises a carbonate-dominated transgressive system which recorded the cratonward migration of the foreland basin subsidence, and siliciclastic depocenters which recorded the progression of the thrust-fold deformation. The foreland basin system has always been maintained in deep-marine environments, i.e., at an underfilled depositional state. It was associated with a thrust wedge which descended toward a deep-marine hinterland, i.e., with a type of orogenic wedge usually related to subduction zones. The Frasnian foreland basin system differs from the one known in the Carboniferous which evolved to overfilled depositional state and was associated with a thrust wedge rising toward a mountainous hinterland. Consequently, the Hercynian orogeny in the Pyrenees seems to result first, from a Frasnian thrusting controlled by a subduction zone located north of the Pyrenees, and second, from a Carboniferous thrusting controlled by the surrection of a frontal thrust belt in the Pyrenees. The association of underfilled foreland basin systems and hinterland-dipping thrust wedges, as exemplified in the Frasnian of the Pyrenees, can be interpreted as illustrative of the initial stages of thrust-wedge growth in deep-marine settings.     

Transverse fold evolution in the External Sierra,southern Pyrenees,Spain

Fault-slip data are used to reconstruct varying tectonic regimes associated with transverse fold development along the eastern and southern margins of the Jaca basin, southern Pyrenees, Spain. The Spanish Pyrenean foreland consists of thrust sheets and leading-edge décollement folds which developed within piggyback basins. Guara Formation limestones on the margins of the Jaca basin were deposited synchronously with deformation and are exposed in the External Sierra. Within the transverse folds, principal shortening axes determined from P and T dihedra plots of fault-slip data show a shift from steep shortening in stratigraphically older beds to NNE–SSW horizontal shortening in younger beds. Older strata are characterized by extensional faults interpreted to result from halotectonic (salt tectonics) deformation, whereas younger strata are characterized by contraction and strike-slip faults interpreted to result from thrust sheet emplacement. The interpretation of the timing for the shortening axes in the younger strata is supported by the observation that these axes are parallel to shortening axes determined from finite strain analysis, calcite twins, and regional thrusting directions determined from fault-related folds and slickenlines. This study shows that fault population analysis in syntectonic strata provides an opportunity to constrain kinematic evolution during orogeny.     

Braided stream and flood plain architecture: the Rio Vero Formation,Spanish Pyrenees

Early- to middle-Miocene fluvial sandstones of the Rio Vero Formation were studied, in an area around the town of Barbastro, south central Pyrenees Spain. The outstanding quality of outcrops in this area allows a three-dimensional study of architectural elements.Six architectural elements are recognised, described in detail, and interpreted from three key localities. Seven main lithofacies were identified and sub-divided into gravelly, sandy and fine-grained lithofacies. The architectural elements and lithofacies have been combined with a hierarchy of depositional bounding surfaces to fully interpret the evolution of the depositional system at the meso- and macro-scale. Not only the different architectural elements and lithofacies of the complete braided fluvial system, but also the lateral variation of the architectural elements were emphasised in this study. Differential tectonic movements, seasonal climate change, and their effect on vertical and lateral evolution of the area were the main control on basin sedimentation, channel interconnection, palaeocurrent patterns, and consequently the fluvial architecture.The presence of lateral ramp anticlines caused the fluvial system to be laterally restricted, with the main channel-belts being located in the areas of highest subsidence and lowest topography. Intervening topographic highs acted as both flood plains and lateral barriers between the main channel systems. The proposed depositional model comprises broad, low-sinuosity, perennial, but seasonal moderate-energy streams. The sandstone architecture is dominated by channel-fill and sheet sands, and associated simple and more complex bars. Adjacent to the main channel-belts fine-grained sandstones, siltstones and immature paleosols occur.The along-strike relationship between major fluvial systems and their outlets into a foreland basin has important implications for the infill of the basin and the modelling of fluvial systems along mountain belt fronts.     

Constraining the age of the last marine sediments in the late Cretaceous of central south Pyrenees (NE Spain): Insights from larger benthic foraminifera and strontium isotope stratigraphy

The uppermost Cretaceous (upper Campanian–Maastrichtian) marine deposits of the central south Pyrenees host a rich larger benthic foraminiferal fauna and several rudist-rich levels. These marine deposits are directly overlain by the continental facies of the Arén and Tremp Formations, which are famous for their fossil dinosaur remains. Larger benthic foraminiferal distribution documents an important faunal turnover in all the carbonate platform environments within the photic zone, from open marine to littoral areas. Biostratigraphy indicates that this turnover occurred close to the Campanian-Maastrichtian boundary. This is also confirmed by strontium isotope stratigraphy which indicates an earliest Maastrichtian age for the appearance of the larger benthic foraminiferal assemblage constituted by Lepidorbitoides socialis, Clypeorbis mammillata, Wannierina cataluniensis, Orbitoides gruenbachensis, Siderolites aff. calcitrapoides, Fascispira colomi, Omphalocyclus macroporus and Laffiteina mengaudi. In particular, a numerical age of 71 Ma is obtained for the Hippurites radiosus level, just a few meters below the first continental deposits of the Arén sensu stricto Formation. The youngest marine sediments of the central south Pyrenees are early Maastrichtian in age. This is also an important constraint for the age of the end-Cretaceous dinosaur fossil localities of the Tremp basin.     

Geophysical methods for mapping Quaternary sediment thickness: Application to the Saint-Lary basin (French Pyrenees)

This study aims at mapping the sediment infill thickness in the Saint-Lary basin (Aure valley, French Pyrenees). For this purpose, we combine passive seismic and gravity surveys. The resonance frequencies of the sediment body are retrieved from seismic ambient measurements, while the gravimetric survey shows negative residual anomaly of about −3 mGal in the basin. Both methods reveal unexpected but consistent bedrock shape. The southern Saint-Lary basin appears deeper than its northern part, with maximal infill thickness of about 300 m and 150 m, respectively. Valley cross sections show regular and smooth “U”-shape in the southern Saint-Lary basin, in contrast to an irregular and asymmetric pattern in the northern basin. This basin shape may be related to Quaternary fluvio-glacial carving processes especially controlled by a regional fault (the Soulan fault), variations in bedrock hardness, and preferential ice flow paths.     

Petrography and temporal changes in petrofacies of deep-marine Ainsa–Jaca basin sandstone systems, Early and Middle Eocene, Spanish Pyrenees

A petrographic study of 157 samples from the Early to Middle Eocene deep-marine sandy systems, Ainsa–Jaca basin, Spanish Pyrenees, shows that each system has a characteristic petrofacies. Three main petrofacies are recognized. Petrofacies 1 sandstones comprise mainly siliciclastic grains (≥80%), subordinate terrigenous carbonate grains and negligible intrabasinal grains. Petrofacies 2 hybrid arenites are characterized by significant amounts (≥10%) of intrabasinal carbonate grains. Petrofacies 3 calcilithites contain relatively abundant (≥10%) extrabasinal carbonate grains. On the basis of these petrofacies, a revised correlation of the sandy systems is proposed between the more proximal Ainsa basin, and the more distal Jaca basin sediments, now separated by the Boltaña anticline, across which it is impossible to actually trace out individual beds or sandstone packages between both basins. The arenite composition in the Ainsa and Jaca basins is interpreted as being controlled mainly by synsedimentary tectonic processes that led to changes in sediment sources during basin evolution.     

A paleomagnetic study of the Basque Arc (Basque-Cantabrian Basin,Western Pyrenees)

The Basque Arc constitutes the northern segment of the Basque-Cantabrian basin, in the western part of the Pyrenees. The main goal of the present study was to find out by means of paleomagnetic analysis if the arched shape of the Basque Arc has a primary origin, due to the development of sedimentary basins related to the opening of the Bay of Biscay or a secondary origin due to rotations about vertical axes. Nine volcanic flows of late Albian to Santonian age (100–83.5 My) were sampled together with 10 sedimentary sites (marls, limestones, calcarenites and sandstones) of lower Jurassic to early Eocene age in order to carry out paleomagnetic and rock-magnetic experiments, 15 paleodirections being obtained. In order to analyse these results together with data from previous studies, the studied area was subdivided into a western, a central and an eastern sector. While inclinations of all three sectors show a similar value, declinations differ. The western sector displays a 37 ± 16° clockwise rotation of its mean paleodeclination, the central sector is not rotated (4 ± 9°) and both sedimentary sites which make up the eastern sector show counter-clockwise rotations (−25 ± 11° and −68 ± 9°). These results suggest that the shape of the Basque Arc does not have a primary origin, but a secondary origin due to rotations about vertical axes as a result of differential shortening related to post-Lutetian compressive tectonics which resulted in the formation of the Pyrenees.     

The upper Maastrichtian dinosaur fossil record from the southern Pyrenees and its contribution to the topic of the Cretaceous–Palaeogene mass extinction event

In the present paper, the fossil record of the archosaurs (dinosaurs, crocodylomorphs and pterosaurs) of the southern Pyrenees before the Cretaceous–Palaeogene (K–Pg) transition is revised. On the basis of this fossil record, a well-dated succession of dinosaurs and other archosaurs is established within polarity magnetochrons C30 and C29r. Almost 150 sites with dinosaur remains have been identified, containing hadrosauroid ornithopods, titanosaur sauropods and theropods, as well as egg sites and tracks. Fossil remains of dinosaurs and other archosaurs are abundant in C29r, disappearing abruptly near the top of the “Lower Red Garumnian” unit of the Tremp Formation. Thus this should be located very close to, or coinciding with the K–Pg boundary. These data suggest that the disappearance of the dinosaurs and other archosaurs was geologically abrupt in the southern Pyrenees, but to date there is no incontrovertible evidence of the presence of the impact level that marks the Cretaceous–Palaeogene boundary. Interestingly, what is highlighted in the southern Pyrenees is that the vertebrate-rich upper Maastrichtian continental sites were replaced by similar sedimentological facies characterized by the virtual absence not only of dinosaurs but also of any vertebrate remain throughout the lower Palaeocene. This could mean that the Danian terrestrial ecosystems of the southern Pyrenees took longer than other areas of the world to recover their biodiversity after the K−Pg extinction event.     

Tectonic controls on drainage evolution and development of terminal alluvial fans, southern Pyrenees, Spain

An understanding of how drainage patterns respond to tectonics can provide an insight into past deformational events within mountain belts and where sediment flux is supplied to depositional basins. The transverse rivers draining the Spanish Pyrenees show sudden diversions to axial courses and the capture of lateral systems producing large trunk rivers that break through the thrust front at structurally controlled points. The drainage reorganization from the initial regularly spaced pattern in the Late Eocene caused by the growth of thrust‐controlled topography influenced the location of outlets into the Ebro basin. The headward capture and merger of rivers as a result of structural diversion formed two large terminal fan systems during the Oligo‐Miocene along the Pyrenean thrust front. The early structural topographic controls on drainage evolution will have long‐term effects on sedimentation and stratigraphic architecture of foreland basins. This will only be maintained as long as there is tectonic uplift and the river systems strive to re‐attain a regular drainage spacing across the orogenic belt as partly achieved in the Pyrenees.     

On the consistency of proposed models for the Pyrenees with the structure of the eastern parts of the belt

The structure of the eastern Pyrenees consists mainly of south-directed thrusts involving basement and cover rocks. An antiformal stack developed by the piling up of basement thrust sheets which outcrop in the Axial zone. These structures account for a thin-skinned thrust model rather than a vertical fault model in which the Axial zone would be essentially autochthonous, and the North-Pyrenean fault the axial plane of a fan thrust system. New data from the Eastern Pyrenees and the thin-skinned model suggest that(1) the structure east of the Pedraforca nappe is similar to that of the Central Pyrenees; (2) the cover rocks of the South-Pyrenean units and of the Axial zone-after restoration—built up a northwards-thickening prism consistent with the existence of a unique Pyrenean sedimentary basin during Mesozoic time; (3) the Axial zone is only a complex antiformal stack developed as a part of South-Pyrenean system related to the Paleogene thrusting-tectonics. The Axial zone palaeogeographic area had no special meaning during Mesozoic time.     

Asynchroneity of maximum glacier advances in the central Spanish Pyrenees

The deglaciation history of the Escarra and Lana Mayor glaciers (Upper Gállego valley, central Spanish Pyrenees) had been reconstructed on the basis of detailed geomorphological studies of glacier deposits, sedimentological and palynological analyses of glacial lake sediments and an accelerator mass spectrometry (AMS) ¹⁴C chronology based on minimum ages from glacial lake deposits. The maximum extent of the Pyrenean glaciers during the last glaciation was before 30 000 yr BP and pre‐dated the maximum advances of the Scandinavian Ice Sheet and some Alpine glaciers. A later advance occurred during the coldest period (around 20 000 yr BP), synchronous with the maximum global ice extent, but in the Pyrenees it was less extensive than the previous one. Later, there were minor advances followed by a stage of debris‐covered glaciers and a phase of moraine formation near cirque backwalls. The deglaciation chronology of the Upper Gállego valley provides more examples of the general asynchroneity between mountain and continental glaciers. The asynchroneity of maximum advances may be explained by different regional responses to climatic forcing and by the southern latitude of the Pyrenees. Copyright © 2002 John Wiley & Sons, Ltd.     

The variscan geology of the Pyrenees

The main outlines of the geology of the Variscan part of the Pyrenees are discussed. Rocks involved in this cycle are high-grade basement gneisses, Palaeozoic sediments and their metamorphic equivalents, late intrusive granodiorites and early, pre-Variscan granites. The main features of the stratigraphy of the Palaeozoic are given.Structures fall into two domains: a low-grade suprastructure, essentially with steep folds and cleavages, and a high-grade infrastructure with dominantly low-dipping foliations. An important phase of early, pre-cleavage folding occurs in low-grade rocks mainly along the southern border of the Axial zone. In high-grade rocks most structures and the metamorphism postdate the main cleavage phase in low-grade rocks. The influence of the Alpine orogeny on the Variscan structures consists mainly of faults, steep, reverse faults in the northern, and south-directed thrusts in the southern part of the Pyrenees. Metamorphism took place under high geothermal gradients and low pressures, as indicated by the abundant occurrence of andalusite and cordierite     

End-signature of deep-marine basin-fill, as a structurally confined low-gradient clastic system: the Middle Eocene Guaso system, South-central Spanish Pyrenees

The ca 300 m thick Guaso system is the youngest part of the ca 4 km thick deep-marine fill of the Middle Eocene Ainsa basin, Spanish Pyrenees. It is overlain by 150 to 200 m of fine-grained slope, prodelta and deltaic sediments. The ca 25 discrete deep-marine sandbodies within the Ainsa basin accumulated over ca 10 Myr, making eustasy the most likely control for coarse sand deposition (probably the ca 400 kyr Milankovitch mode). The first-order control on basin-scale accommodation, however, was tectonically-driven subsidence. Previously, the Guaso sandbodies were interpreted as linked to deep erosional, canyon-like features, but here it is argued that they are laterally extensive sandbodies, built by lateral-switching of 3 to 10 m deep erosional channels, and confined only by basin structure during deposition. The Guaso system represents the end of deep-marine deposition in a structurally-confined, delta-fed, low-gradient clastic system. The critical end-signature of deep-marine deposition was a phase of differential tectonic uplift above the underlying (Boltaña) thrust creating a narrower and shallower basin morphology, thus allowing sedimentation to create a low-gradient clastic system. Then, the next eustatic sea-level fall was insufficient to permit the cutting of canyons or deeply-incised slope channels, as had been the case earlier when the topographic relief between shelf and basin was at least several hundred metres greater. Such low-gradient clastic systems may characterize the end-signature for the infill of other shallowing-up deep-marine basins where a tectonic driver on subsidence is removed and/or differential uplift/subsidence leads to reduced sea floor gradients, leaving eustasy and sediment flux as the principal control on sediment supply.     

The Last Pterosaurs: First Record from the Uppermost Maastrichtian of the Tremp Syncline (Northern Spain)

Pterosaur bones are for the first time reported from the Tremp Syncline(Spain) and the southern Pyrenees.They come from the Torrebilles-2 site(Isona,Lleida Province) in the upper Maastrichtian part of the Tremp Formation,which falls within the C29r magnetochron.Fragmentary long bones are referred to the Pterosauria based on their cortical thinness,micro-architecture,and histological features.They belong to very large to possibly giant individuals.The most complete specimen is possibly a partial right femur.The review of the Maastrichtian pterosaur record shows that only a few specimens can be referred to the C29r(which contains the K/Pg boundary).The identification of fragmentary remains could improve our knowledge about the abundance and distribution of these flying sauropsids just before the end-Cretaceous mass extinction.     

First biostratigraphical constraints on the pre-Upper Ordovician sequences of the Pyrenees based on organic-walled microfossils

The first age constraint on the upper part of the pre-Upper Ordovician series of the Pyrenees is presented. Data are based on acritarchs from the Jujols Group in the La Molina area on the southern slope of the Canigó massif, near the Upper Ordovician unconformity. The new data allow us to attribute a Late Cambrian (Furongian) to Early Ordovician age to the upper part of the Jujols Group and to support the absence of well developed Middle Ordovician strata in the Pyrenees. Our results confirm the occurrence of an Upper Ordovician unconformity and highlight the “Sardic” affinity of the pre-Upper Ordovician sequences in the Pyrenees. These sequences differ markedly from the pre-Upper Ordovician series of the Iberian Massif.     

Prediction of ground displacements and velocities from groundwater level changes at the Vallcebre landslide (Eastern Pyrenees, Spain)

In active landslides, the prediction of acceleration of movement is a crucial issue for the design and performance of warning systems. The landslide of Vallcebre in the Eastern Pyreenes, Spain, has been monitored since 1996 and data on rainfall, groundwater levels and ground displacements are measured on a regular basis. Displacements observed in borehole wire extensometers have shown an immediate response of the landslide to rainfall episodes. This rapid response is likely due to the presence of preferential drainage ways. The occurrence of nearly constant rates of displacement in coincidence with steady groundwater levels suggests the presence of viscous forces developed during the movement. An attempt to predict both landslide displacements and velocities was performed at Vallcebre by solving the momentum equation in which a viscous term (Bingham and power law) was added. Results show that, using similar rheological parameters for the entire landslide, computed displacements reproduce quite accurately the displacements observed at three selected wire extensometers. These results indicate that prediction of displacements from groundwater level changes is feasible.     

Trace fossils as diagnostic indicators of deep-marine environments, Middle Eocene Ainsa-Jaca basin, Spanish Pyrenees

A quantitative study of trace fossil abundance in the Middle Eocene deep-marine clastic systems, Ainsa-Jaca basin, Spanish Pyrenees, shows that they are powerful discriminators of submarine fan and related environments. Sixteen fan and related environments are recognized from upper-slope gully to the distal basin-floor. For each environment, the degree of bioturbation (density), trace-fossil diversity, number of pre-depositional and post-depositional trace fossils, as well as the number of graphoglyptid ichnospecies were quantified. In the more laterally confined and channel-dominated Ainsa basin, there is a trend of increasing bioturbation intensity and trace-fossil diversity away from channel-axis to off-axis environments. In the more unconfined and distal Jaca basin, there is a trend of increasing trace-fossil diversity and number of pre-depositional trace fossils including graphoglyptids from the channel-lobe transition to the fan-fringe. The trace-fossil assemblages of the Ainsa-Jaca basin are characteristic of a number of sub-ichnofacies of the Nereites ichnofacies. In the distal Jaca basin, the Paleodictyon sub-ichnofacies occurs in the lobe-fringe and fan-fringe, whereas the distal basin-floor has a trace-fossil assemblage typical of the Paleodictyon sub-ichnofacies, but with a high proportion of post-depositional fodinichnia . Trace-fossil assemblages of proximal basin, axial, environments are characteristic of the Ophiomorpha rudis sub-ichnofacies, whilst proximal off-axis environments, have a mixed Paleodictyon-Ophiomorpha rudis sub-ichnofacies trace-fossil assemblage.