Latest Campanian and Maastrichtian Siderolitidae (larger benthic foraminifera) from the Pyrenees (S France and NE Spain)

The siderolitids from the uppermost Campanian and Maastrichtian deposits of the Pyrenees have been re-studied. This has revealed a high diversity and rapid replacement of taxa, confirming the group as a good tool for high resolution biostratigraphy. Two genera have been found in the uppermost Campanian–Maastrichtian interval in the Pyrenean deposits: Siderolites Lamarck, and Wannierina Robles-Salcedo. Siderolites, with canaliferous spines or denticulate periphery, is represented by four species replacing each other from the latest Campanian to Maastrichtian: Siderolites praecalcitrapoides (latest Campanian), S. pyrenaicus sp. nov. (early Maastrichtian), Siderolites calcitrapoides (late Maastrichtian) and Siderolites denticulatus (late Maastrichtian). Wannierina is characterised by well-developed keels and ramified marginal canals. Two species of Wannierina have been identified and they succeeded one another from latest Campanian to early Maastrichtian: Wannierina vilavellensis sp. nov. (latest Campanian) and Wannierina cataluniensis (early Maastrichtian). The species of the genus Siderolites inhabited shallow waters of tropical to subtropical platforms with moderate-to-high water-energy conditions and those of the genus Wannierina are typical of deep–water low-energy environments but still in the eutrophic zone.     

Upper cretaceous biostratigraphy of the south-central Pyrenees (Lleida,Spain)

Abstract

Biostratigraphical data using larger foraminifera and planktonic foraminifera permitted us to establish the correlation between shallow platform sediments rich in larger foraminifera (Montsec and Serres Marginals thrust sheets) and deeper ones containing planktonic foraminifera (Boixols thrust sheet).

Consequently, the Santa Fe limestones containing Ovalveolina-Praealveolinaassemblage represent the Cenomanian. Early Turonian ( ‘IT~ archaeocretacea and P. helvetica zones) exist in both, Montsec and Boixols thrust sheets and it is constituted by Pithonella limestones. Late Turonian (M. schneegansi zone) is only present in Boixols thrust sheet (Reguard Fm.), the Montsec thrust sheet having an erosive hiatus.

Late Coniacian-Early Santonian (D. Concavata zone) is represented in the Montsec thrust sheet (Cova Limestones) and in the eastern part of the Boixols thrust sheet (St. Corneli Fm.) by two differents facies giving two different microfaunal assemblages; the firts one, characterized by Ophtalmidiidae s.l indicate a restricted lagoonal environment while the second one, characterized by diverses species of complex agglutinated, Fabulariidae, Meandropsinidae and Rotaliidae, represents an open shallow platform. In the Boixols thrust sheet (Anseroles Fm.) dominate the planktonic foraminifera and small benthic.

In the late Santonian (D. asyrnetrica zone) the sea reached as far as Serres Marginales thrust sheet where sediments (Tragó de Noguera unit) are terrigenous and deposited in a very shallow platform. In the Montsec thrust sheet (Montsec marls) the larger foraminifera indicate a platform deeper than that of the Serres Marginals thrust sheet. In the Boixols thrust sheet the sediments are deposited in an outer platform (Herbasavina Fm.) or turbiditic basin (Mascarell Mb.).

During Campanian times the transgresion reaches the maximum. In the Serres Marginals sediments are deposited in a restricted shallow environment rich in Meandropsinidae (Serres Limestones). In the Montsec thrust sheet they are deposited in a platform with patch reefs and shoals (Terradets limestones) and in the Boixols one in an outer platform, talus and/or basin.

During Early Maastrichtian times (C. falsostuarti zone) terrigenous materials arrived in the basin, the rate of sedimentation increased outstripping the subsidence rate and the retreat of the sea to the north. Late Maastrichtian (C. gansseri zone) is only present in the Boixols thrust sheet.     

Sedimentology and depositional environments of the Maastrichtian Patti Formation, southeastern Bida Basin, Nigeria

The Maastrichtian Patti Formation, which consists of shale - claystone and sandstone members, constitutes one of the three Upper Cretaceous lithostratigraphic units of the intracratonic southeastern Bida Basin, in central Nigeria. Well exposed outcrops of this formation were investigated at various locations around the confluence of the Niger and Benue Rivers. The lithostratigraphic sections were measured and their peculiar sedimentological features such as textures, physical and biogenic sedimentary structures, facies variations and associations were documented and used to interpret the depositional environments and develop a paleogeographic model. Some selected representative samples of the sedimentary depositional facies were also subjected to grain size analysis.Three shoreline sedimentary depositional facies composed of shoreface, tidal channel, and tidal marsh to coastal swamp facies were recognized in the study area. Continental sedimentary depositional facies such as fluvial channel, swamp, and overbank were also documented. The sandstones of the shoreface and tidal channel facies are medium- to coarse-grained, moderately sorted (standard deviation ranges from 0.45–1.28 averaging 0.72), and quartzarenitic. The fluvial channel sandstone facies are coarse- to very coarse-grained, mostly poorly sorted (standard deviation ranges from 0.6–1.56 averaging 1.17), and subarkosic. Typical sedimentary structures displayed by the shoreface and tidal channel facies include burrows, clay drapes, hummocky and herringbone cross stratifications, whereas the fluvial channel sandstone facies are dominated by massive and planar cross beddings. The tidal marsh to coastal swamp shales and ferruginised siltstone facies are fossiliferous and bioturbated, whereas the nonmarine swamp siltstones contain vegetal imprints and lignite interbeds. The overbank claystone facies are massive and kaolinitic.In the study area, a regressive to transgressive model is proposed for the Patti Formation. This model correlates with stratigraphically equivalent sediments of the Ajali and Mamu Formations in the adjacent Anambra Basin to a great extent.     

Fluid flow record from fracture-fill calcite in the Eocene limestones from the South-Pyrenean Basin (NE Spain) and its relationship to oil shows

The presence of oil shows associated with fractures provides a significant opportunity to a) unravel the type, origin and evolution of fluids involved in fracture-fills, and b) examine how they relate to oil migration. Two stages of calcite cement (C1 and C2) were distinguished in the fractures of the Eocene Armàncies platform carbonates; C1 is characterised by fence-like crystals, exhibits dull red luminescence and contains abundant twin planes, inclusions and δ¹⁸O values that range from − 6.2‰ to − 4.8‰ VPDB. C2 consists of blocky clean crystals, is characterized by dark brown-red luminescence that alternates with yellow bands, and contains hydrocarbon fluid inclusions with homogenisation temperatures of approximately 120 °C. δ¹⁸O values range from − 9.6‰ to − 8.9‰ VPDB. The remaining porosity after C2 precipitation is filled with liquid oil that reached 115 °C. This would seem to indicate that free oil and fluid inclusions oil probably come from the same migration pulse. Oil migration timing was coeval with C2 and continued after calcite cementation was completed.     

Lepidorbitoides (Foraminifera) from the lower Maastrichtian Kallankuruchchi Formation,Cauvery Basin,India: Morphometry and paleobiogeography

The southernmost occurrence of the early Maastrichtian larger benthic foraminifera (LBF) in Tethys is known from the Kallankuruchchi Formation in the Cauvery Basin, SE India, represented by Lepidorbitoides-Siderolites assemblages. The systematics, age and paleobiogeography of Lepidorbitoides here have as yet remained unresolved due to lack of information particularly on the nepionic arrangement, whereas their links with the Western Tethyan and Caribbean biogeographic domains were speculated. Lepidorbitoides, studied from the same level in seven samples in two separate areas, invariably possess quadriserial nepionts and adauxiliary chamberlets, whose mean number ranges from 3.79 to 4.67. The ratio between the sample means of the internal diameter of deuteroconch and protoconch varies between 1.72 and 1.86. The equatorial layer in the early stage consists of arcuate chambers with basal stolons, and ogival-to spatulate chamberlets with annular and oblique stolons in the later stages. These features are consistent with the phylogenetically advanced members of the Western Tethyan Lepidorbitoides lineage, such as L. minor (Schlumberger) and L. socialis (Leymerie), and all samples were assigned to the transitional development stages of these species based on the morphometry. The taxonomic status of some Lepidorbitoides species, originally described from the Kallankuruchchi Formation and widely adopted in previous works, such as L. blanfordi (Rao) and L. inornata (Rao), are not justified. We extend the geographic range of Western Tethyan Lepidorbitoides to southern India.     

The chronology of dinosaur oospecies in south-western Europe: Refinements from the Maastrichtian succession of the eastern Pyrenees

The Late Cretaceous deposits of the continental Tremp Formation in the Vallcebre Syncline (South-eastern Pyrenees) provide an extensive egg record of dinosaurs. The parataxonomical study and analysis of multiple eggshell samples, the precise stratigraphical control of several sections and the time calibration of the abundant egg levels enable the establishment of a robust oospecies succession. The successive occurrence of three megaloolithid oospecies (Megaloolithus siruguei-Megaloolithus mamillare-Megaloolithus sp.) in the Early and Late Maastrichtian is well correlated with the magnetic polarity time scale throughout the chron 31. The replacement of Megaloolithus siruguei with Megaloolithus mamillare occurs around the reversal of chrons 31r-31n. A comparison with oospecies successions from Arc basin localities (France) allows the age calibration for such oospecies replacement to be confirmed. This age refinement implies that some of the stated boundaries for the proposed oospecies assemblages may change and that an in-depth revision of the age and magnetic calibration of some south European egg localities is required.     

A riparian plant community from the upper Maastrichtian of the Pyrenees (Catalonia,NE Spain)

Angiosperms began to colonise riparian habitats very soon in their evolutionary history, probably already in the Aptian, but it is still poorly known when flowering plants finally dominated entirely these kind of communities as they do in the present. A new fossil plant locality (Molí del Baró-1) from the upper Maastrichtian of the Southern Pyrenees is described in which meandering river facies represent one of the first riparian communities formed only by angiosperms. The fossil assemblage consists of abundant leaves, seeds, logs and sporomorphs. Angiosperms remains dominate in all these cases and the leaf sample is mostly composed of a new eudicot willow-like species, Saliciphyllum gaetei sp. nov., the palm Sabalites longirhachis and an helophytic monocot. Pollen remains suggest that the later belonged to Typhaceae. Most of these plant remains were parautochthonous and deposited in a pond formed in the accretional part of a meander loop. The locality of Molí del Baró-1 represents an unique plant fossil assemblage in the uppermost Cretaceous of southern Europe. It clearly differs from those reported in other Maastrichtian localities of the Pyrenees (Fumanya and South Isona) and from the Campanian-Maastrichtian of Austria and Romania. In addition, it reflects a surprisingly modern physiognomy for a Late Cretaceous riverine plant assemblage that was built up with willow-like plants, palms and reeds.     

Tectonically induced climate and its control on the distribution of depositional systems in a continental foreland basin, Cloverly and Lakota Formations (Lower Cretaceous) of Wyoming, U.S.A.

Continental sediments of the Cloverly and Lakota Formations (Lower Cretaceous) in Wyoming are subdivided into three depositional systems: perennial to intermittent alluvial, intermittent to ephemeral alluvial, and playa. Chert-bearing sandstones, conglomerates, carbonaceous mudrocks, blocky mudrocks, and skeletal limestones were deposited by perennial to intermittent alluvial systems. Carbonaceous mudrocks contain abundant wood fragments, cuticle and cortical debris, and other vascular plant remains representing deposition in oxbow lakes, abandoned channels, and on floodplains under humid to seasonal conditions. Intraformational conglomerates, sandstones, bioturbated and blocky mudrocks with caliche nodules, and bioturbated limestones characterize deposition in intermittent to ephemeral alluvial systems. Bioturbated limestones are encased in bioturbated mudrocks with abundant pseudo-slickensides. The presence of caliche nodules in some of the blocky to bioturbated mudrocks is consistent with supersaturation and precipitation of calcium carbonate from groundwater under semi-arid conditions. Caliche nodules, pseudo-slickensides, and carbonate-rich floodplain sediments are interpreted to have been deposited by intermittent to ephemeral alluvial systems under seasonal to semi-arid climatic conditions. Laminated mudrocks, siltstones, vuggy carbonates, bedded to nodular evaporites, pebbly mudrocks, and diamictites were deposited in evaporative alkaline lakes or playas. Pebbly mudrocks and diamictites are interpreted to represent deposition from channelized and unchannelized hyperconcentrated flows on a playa, resulting from intense rain events within the basin.The areal abundance and distribution of these depositional systems change systematically across the overfilled portion of the Early Cretaceous Cordilleran foreland basin in Wyoming. The lower part (A-interval) of the Cloverly and Lakota Formations is characterized by deposits of perennial to intermittent rivers that existed 300 to 1000 km east of the Sevier fold-and-thrust belt. Proximal to the Sevier fold-and-thrust belt, the A-interval of the Cloverly Formation and upper Ephraim Formation of the Gannett Group are typified by deposits of intermittent to ephemeral rivers and their associated floodplains. In the middle part (B-interval) of the Cloverly Formation, intermittent to ephemeral alluvial systems expand to 600 km into the basin. The upper part (C-interval) of the Cloverly Formation is characterized by playa deposits in the Bighorn and Wind River Basins and intermittent to ephemeral alluvial deposits along the front of the ancestral Sevier Mountains. Deposits of perennial to intermittent alluvial systems in the C-interval of the Cloverly and Lakota Formations are restricted to the Black Hills region, almost 900 km to the east of the Sevier Mountains. The change in the areal distribution of depositional systems through time within this continental foreland basin may be attributed to the development of a rain shadow associated with the uplift of the Sevier Mountains in the Early Cretaceous.     

Sedimentology and palynology of the Calafate Formation (Maastrichtian), Austral Basin, Southern Patagonia, Argentina

The Calafate Formation crops out in south-western Santa Cruz Province, Argentina, and displays a stacking of asymmetrical coarsening–fining-upward cycles. These cycles are interpreted as the product of short-lived transgressive-regressive events in which the coarsening upward part represents sedimentary aggradation with a stable or decreasing sea level. Sedimentological and palynological analyses indicate nearshore marine conditions. Even though the existence of an estuary or incised valley cannot be determined, this is the most probable palaeogeographic model. Based on dinoflagellate cysts, the base of the section is considered to be not older than Maastrichtian. The presence of the oyster Ambigostrea clarae (Ihering) occurring together with the dinoflagellate cyst species Manumiella druggii (Stover) Bujak and Davies and Eisenackia circumtabulata Drugg in the middle part of the section indicates an age no older than late Maastrichtian. According to sedimentological data, deposits representing the Cretaceous–Palaeogene transition would have been eroded, which is confirmed by the presence of Grapnelispora loncochensis Papú. This megaspore is a consistent component of the Maastrichtian assemblages from Patagonia.     

Provenance constraints on the Tremp Formation paleogeography (southern Pyrenees): Ebro Massif VS Pyrenees sources

A detailed petrological study has been performed for the end-Cretaceous clastic deposits of the southern Pyrenees. Provenance results indicate that the Maastrichtian systems from both the Àger and the Vallcebre synclines show compositional features that mainly consist of a high proportion of single and polycrystalline quartz grains, feldspar and plutonic fragments. By contrast, the sandstone systems of the Tremp syncline exhibit minor contributions from igneous source areas and higher amounts of carbonatic components. These results reveal that the Tremp basin had a source area interpreted as situated to the North in the uplifting Pyrenees. The fact that this basin does not show a high plutonic source signal indicates that the Àger and the Vallcebre basins had been fed from a distinct source area located to the South, here interpreted as the Ebro Massif. Thus, the differences mentioned above might imply that the Montsec High acted as a barrier, avoiding a southern influence in the Tremp basin.     

A paleomagnetic study of the Basque Arc (Basque-Cantabrian Basin,Western Pyrenees)

The Basque Arc constitutes the northern segment of the Basque-Cantabrian basin, in the western part of the Pyrenees. The main goal of the present study was to find out by means of paleomagnetic analysis if the arched shape of the Basque Arc has a primary origin, due to the development of sedimentary basins related to the opening of the Bay of Biscay or a secondary origin due to rotations about vertical axes. Nine volcanic flows of late Albian to Santonian age (100–83.5 My) were sampled together with 10 sedimentary sites (marls, limestones, calcarenites and sandstones) of lower Jurassic to early Eocene age in order to carry out paleomagnetic and rock-magnetic experiments, 15 paleodirections being obtained. In order to analyse these results together with data from previous studies, the studied area was subdivided into a western, a central and an eastern sector. While inclinations of all three sectors show a similar value, declinations differ. The western sector displays a 37 ± 16° clockwise rotation of its mean paleodeclination, the central sector is not rotated (4 ± 9°) and both sedimentary sites which make up the eastern sector show counter-clockwise rotations (−25 ± 11° and −68 ± 9°). These results suggest that the shape of the Basque Arc does not have a primary origin, but a secondary origin due to rotations about vertical axes as a result of differential shortening related to post-Lutetian compressive tectonics which resulted in the formation of the Pyrenees.     

Constraining the age of the last marine sediments in the late Cretaceous of central south Pyrenees (NE Spain): Insights from larger benthic foraminifera and strontium isotope stratigraphy

The uppermost Cretaceous (upper Campanian–Maastrichtian) marine deposits of the central south Pyrenees host a rich larger benthic foraminiferal fauna and several rudist-rich levels. These marine deposits are directly overlain by the continental facies of the Arén and Tremp Formations, which are famous for their fossil dinosaur remains. Larger benthic foraminiferal distribution documents an important faunal turnover in all the carbonate platform environments within the photic zone, from open marine to littoral areas. Biostratigraphy indicates that this turnover occurred close to the Campanian-Maastrichtian boundary. This is also confirmed by strontium isotope stratigraphy which indicates an earliest Maastrichtian age for the appearance of the larger benthic foraminiferal assemblage constituted by Lepidorbitoides socialis, Clypeorbis mammillata, Wannierina cataluniensis, Orbitoides gruenbachensis, Siderolites aff. calcitrapoides, Fascispira colomi, Omphalocyclus macroporus and Laffiteina mengaudi. In particular, a numerical age of 71 Ma is obtained for the Hippurites radiosus level, just a few meters below the first continental deposits of the Arén sensu stricto Formation. The youngest marine sediments of the central south Pyrenees are early Maastrichtian in age. This is also an important constraint for the age of the end-Cretaceous dinosaur fossil localities of the Tremp basin.     

Further evidence of marine Cenomanian,lower Turonian and Maastrichtian in the Upper Benue basin of Nigeria (West Africa)

Field investigations in the Upper Benue rift basin of Nigeria highlight the necessity for a review of the Cenomanian-Turonian biostratigraphy of the area. Three ammonite zones of Vascoceras bulbosum (Reyment), Paravascoceras costatum (Reyment) and Pseudotissotia (Bauchioceras) nigeriensis (Woods) have been recognized at sections exposed at the quarry of the cement factory near the village of Ashaka. The sediments within the Vascoceras bulbosum zone also contain specimens of Exogyra olisiponensis (Sharpe), Kanabiceras septemseriatum (Cragin) and Epengonoceras dumbli (Cragin). These are all indicative of an uppermost Cenomanian age and reveal that at least the lower portion of the Gongila Formation as exposed at Ashaka is faunistically equivalent to the Pindiga Formation as exposed in the type section at Pindiga.The shales in the Gombe region in which Libycoceras ismaeli (Zittel) was found are probably part of the Maastrichtian Fika shales outcropping extensively in the northern part of the area. The discovery of Cyclolithes nov. sp. within Gombe sandstones at Dakiti near Kumo confirm that they are at least partially marine in origin and Maastrichtian in age. This could be considered as an additional evidence in favour of the presence of marine waters in the Upper Benue during the last part of the Cretaceous.     

Facies model of a mixed clastic–carbonate,wave‐dominated open‐coast tidal flat (Tithonian–Berriasian,north‐east Spain)

Detailed logging and analysis of the facies architecture of the upper Tithonian to middle Berriasian Aguilar del Alfambra Formation (Galve sub‐basin, north‐east Spain) have made it possible to characterize a wide variety of clastic, mixed clastic–carbonate and carbonate facies, which were deposited in coastal mudflats to shallow subtidal areas of an open‐coast tidal flat. The sedimentary model proposed improves what is known about mixed coastal systems, both concerning facies and sedimentary processes. This sedimentary system was located in an embayed, non‐protected area of a wide C‐shaped coast that was seasonally dominated by wave storms. Clastic and mixed clastic–carbonate muds accumulated in poorly drained to well‐drained, marine‐influenced coastal mudflat areas, with local fluvial sandstones (tide‐influenced fluvial channels and sheet‐flood deposits) and conglomerate tsunami deposits. Carbonate‐dominated tidal flat areas were the loci of deposition of fenestral‐laminated carbonate muds and grainy (peloidal) sediments with hummocky cross‐stratification. Laterally, the tidal flat was clastic‐dominated and characterized by heterolithic sediments with hummocky cross‐stratification and local tidal sandy bars. Peloidal and heterolithic sediments with hummocky cross‐stratification are the key facies for interpreting the wave (storm) dominance in the tidal flat. Subsidence and high rates of sedimentation controlled the rapid burial of the storm features and thus preserved them from reworking by fair‐weather waves and tides.     

Turtles from the Lower Cretaceous of the Cameros Basin (Iberian Range, Spain)

Knowledge of the turtle fauna from the Lower Cretaceous of the Iberian Peninsula has been very limited until now. There are several fossil sites where Lower Cretaceous associations of continental vertebrates have been found. Although turtles have been identified in some of them, most of these specimens have not been studied, so the diversity is unknown. Among all these findings, the turtles from the Cameros Basin are considered particularly relevant, both in their abundance and diversity. Their study has allowed the identification of several taxa. At least one representative of Solemydidae and three taxa of Eucryptodira are recognized. This study establishes kinship and biogeographic relationships between the taxa in Cameros with those found in other Spanish fossil sites and with those of other European regions.     

First pachycephalosaurid (Dinosauria: Ornithischia) from the Frenchman Formation (upper Maastrichtian) of Saskatchewan,Canada

Compared to earlier representatives of the family, pachycephalosaurids are less well known from upper Maastrichtian deposits around the world. Here, we report on a nearly complete left postorbital attributable to the pachycephalosaurid Sphaerotholus cf. Sphaerotholus buchholtzae from the upper Maastrichtian Frenchman Formation of Saskatchewan. This marks a probable northern range extension for the species into Canada, and the first occurrence of a pachycephalosaurid from this formation. We further demonstrate the taxonomic distinction between Stegoceras edmontonense and S. buchholtzae, which has been debated, based on postorbital sutural proportions. The northerly occurrence of Sphaerotholus cf. S. buchholtzae is consistent with the hypothesis of low beta diversity during the late Maastrichtian of North America, and its high stratigraphic incidence documents the persistence of non-pachycephalosaurin pachycephalosaurines in a critical interval for understanding the end-Cretaceous mass extinction event.     

Titanosaur (Dinosauria: Sauropoda) Osteoderms from the Maastrichtian of Uberaba, Minas Gerais State, Brazil

Titanosaur remains are common findings at the paleontological site of Peirópolis in Uberaba. Among those remains, two osteoderms referred to titanosaur sauropods were reported. Both dermal bones share many features, such as the coarse texture, parallel ventral grooves, lack of the cingulum and presence of a ventral ridge (crest). Armored titanosaurs had wide distribution in the Upper Cretaceous, and have been reported from Argentina, Brazil, France, Madagascar, Malawi, Romania and Spain. In the present work we describe and compare the Brazilian titanosaur osteoderms found thus far.     

The lower carboniferous (dinantian) of the Swords area: Sedimentation and tectonics in the Dublin Basin,Ireland

The litho- and biostratigraphy of the Lower Dinantian succession in a deeper part of the Dublin Basin is described. The sub-Waulsortian Malahide Limestone Formation (emended) is described fully for the first time, and has proved to be very much thicker than was previously suspected, in excess of 1200 m. Succeeding the ‘Lower Limestone Shale’ unit, which is transitional from the underlying Old Red Sandstone facies, the following six new members are recognized: Turvey Micrite Member, Swords Argillaceous Bioclastic Member, St. Margaret's Banded Member, Huntstown Laminated Member, Dunsoghly Massive Crinoidal Member and Barberstown Nodular Member (top). The Malahide Limestone Formation is overlain by ‘Waulsortian’ limestones of the Feltrim Limestone Formation (new name) which form overlapping and isolated mudmounds with complex relationships with their enclosing non-mound facies. Though very much thicker, the Courceyan succession is comparable with that elsewhere on the south side of the Basin, and is part of the Kildare Province (Strogen and Somerville 1984). Isopach maps for the region show that this province and the North Midlands are separated by the deepest part of the Dublin Basin, named the ‘East Midlands Depocentre’, in which a shale-dominant facies is present. The top of the ‘Waulsortian’ is of early Chadian age. Formations younger than this are dominated by basinal calcareous shales (Tober Colleen Formation) and by storm deposits and calciturbidites with appreciable terrigenous input from the east (Rush Formation). The Courceyan main shelf conodont biozones are also greatly thickened in this area. The Pseudopolygnathus multistriatus Biozone (> 300 m thick) is succeeded by a very thick (> 900 m) Polygnathus mehli Biozone. The base of the Chadian is considered to occur below the top of the Feltrim Limestone Formation and, although equivocal, may be diagnosed in the Dublin Basin by the first appearance of the problematic microfossil Sphaerinvia piai and a primitive form of the calcareous alga Koninckopora. In the late Courceyan, the Swords area was part of a gently sloping shelf extending northwards into the basin. During deposition of the Feltrim Limestone Formation there was major deepening and there is evidence of initial break up of the Dublin Basin by faulting into separate blocks. By Chadian time the Basin was definitely subsiding by fault displacements and basinal limestones contain shallow water faunas and littoral sand and pebbles derived by turbidite flows from the margins of the higher blocks. The early subsidence was apparently by pure flexure, but in the Viséan the Dublin Basin was fault-controlled, differing from the adjacent Shannon Basin in having both margins strongly faulted.