扬子板块西北缘下泥盆统平驿铺组双壳类遗迹化石

四川北部广元地区下泥盆统平驿铺组产出丰富的双壳类遗迹化石,但尚未开展过系统的遗迹学工作,也未进行过行为学与古生态学分析。文中针对四川广元马家剖面平驿铺组中部地层开展沉积学和系统遗迹学研究,在陆棚至近滨带下部沉积中共识别出3个属种的双壳类遗迹化石,包括Lockeia siliquaria,Protovirgularia rugosa,Ptychoplasma vagans,以及其他无脊椎动物门类遗迹化石Cruziana problematica,Dimorphichnus isp.,Lophoctenium isp.等。对双壳类遗迹化石的出现方式、形态以及保存特征的分析表明,平驿铺组中部曾出现过2种类型的双壳类,分属具分叉足(原鳃目)和楔状足的类群。原鳃目分布于过渡带和陆棚区,营沉积物摄食生活;受高能事件或沉积物掩埋影响,它们由停息状态变为“逃逸”状态的行为产生了Pro.rugosa。具楔状足的双壳类依生态类型可分为2类：在过渡带与陆棚区,漫游的、营沉积物摄食的类型产生了Pty.vagans;而在近滨下部,滤食性双壳类居群则在不同期次高能事件的间隙对沉积物进行大规模殖居,产生了L.sil...     

Zoophycos and related trace fossils from the Chefar El Ahmar Formation,Upper Emsian-Frasnian Ia-Ib (Ougarta,SW Algeria)

The Upper Emsian to Frasnian Ia-Ib strata of the Marhouma area (or “km 30” outcrop), exposed in the Ougarta Range (SW Algeria) belong to the Chefar El Ahmar Formation. On the basis of distinct lithological and palaeontological features, this formation is subdivided into three members (Lower Marly Limestones Member, Middle Marly Limestones Member, and Upper Marly Limestones Member). The studied beds show low to moderate diversity of trace fossil assemblage which contains thirteen ichnotaxa: Chondrites intricatus, Chondrites isp., Chondrites cf. targionii, Circulichnis cf. montanus, Cochlichnus isp., Neonereites biserialis, Neonereites multiserialis, Nereites isp., Palaeophycus isp., Planolites isp., Thalassinoides isp., Zoophycos aff. cauda-galli, and Zoophycos isp. A. The two latter ichnotaxa are the most common trace fossils in the assemblage and occur at three different levels showing different bioturbation intensities. The first Zoophycos-bearing level (Zl 1) is characterised by an overall high bioturbation intensity reflecting a very high oxygenation rate and nutrient supply, allowing the development of large and dense Zoophycos specimens. The second Zoophycos-bearing level (Zl 2) has a considerable reduction of bioturbation intensity as compared to the previous level, with an abundance of Chondrites, which is probably due to radical palaeoecological changes that suggests dysoxic and stressful conditions. The third Zoophycos-bearing level (Zl 3) is characterised by an overall moderate bioturbation intensity. The distribution of trace fossils was influenced by lithology, sedimentation rate, energy level (storm events), bottom oxygenation, and nutrient supply. The lithofacies and trace fossils of the Chefar El Ahmar Formation both indicate a depositional environment fluctuating from the lower shoreface to lower offshore zone.     

Gastrochaenolites ornatus Kelly and Bromley and other trace fossils on a Middle Miocene omission surface from the Aures Massif,Algeria

Trace fossils provide detailed palaeoenvironmental and palaeoecological information of both ancient and modern sedimentary systems. During middle Miocene times the Aures Massif located in the northeastern part of Algeria, was affected by, at least, one marine transgression. The latter led to the installation of a carbonate platform, which is placed, for the first time, in a Mediterranean context. In the Rhassira basin, the Middle Miocene marine succession is characterised by carbonate platform deposits dominated by rhodolith beds, typical of those known throughout the Mediterranean area. This succession can be divided into many units separated by discontinuities interpreted here as omission surfaces. The Djebel Arhane section shows two omission surfaces characterised by a pre-omission suite (firmground) represented by Balanoglossites burrows for the first surface and Gastrochaenolites ornatus burrows/borings for the second one, and an omission suite (hardground) as evidenced by the bioerosive structures Trypanites and Caulostrepsis, in both surfaces, respectively. Gastrochaenolites ornatus traces were formed and preserved in firm, compact, semi-lithified and fine-grained substrates (firm- to hardground), indicating the Glossifungites ichnofacies. They show bioglyphs which have been formed during contraction of the posterior adductor muscles. These suggest that their tracemakers were represented by suspension-feeding bivalves, most probably Pholadidae or Mytilidae, which rotated during penetration. The fill of these traces is composed of marine deposits related to a transgressive lag. The omission suite is divided into two ichnocoenoses: (i) pre-lithification burrows/borings, and (ii) post-lithification borings. This is the first report of the ichnotaxon G. ornatus from Algeria.     

准噶尔西北部塔尔巴哈台组早石炭世植物和遗迹化石的发现及其古地理意义*

首次在准噶尔西北部塔尔巴哈台组上部发现了早石炭世维宪期植物化石 Archaeocalamites scrobiculatus,Sublepidodendron grabaui,S. mirabile,S. cf. mirabile,Sublepidodendron sp., Lepidodendropsis sp., Mesocalamites sp., Sigillaria sp., Rhodiopteridium?sp.,Knorria sp.,证实了该组的时代可延续到早石炭世。根据研究区塔尔巴哈台组和上覆黑山头组的生物组合面貌及二者的接触关系,论证了塔尔巴哈台组顶部和黑山头组在准噶尔西北部具有明显的穿时性,前者从杜内期延续到维宪期,后者从杜内期延续到谢尔普霍夫期。与植物化石共同保存的深水相遗迹化石指示了塔尔巴哈台地区在早石炭世早中期为半深海-深海沉积环境,不同于东部吉木乃地区同期的滨浅海沉积环境,说明准噶尔西北部在早石炭世存在着不同的沉积体系。北疆地区早石炭世广泛的海侵活动以及额尔齐斯-斋桑洋向南侧哈萨克斯坦板块俯冲是造成这种沉积差异的主控因素,同时也造成了塔尔巴哈台组和黑山头组在区域上具穿时性。     

华南泥盆纪遗迹化石及遗迹相

通过对华南泥盆系6条剖面(四川北川甘溪、广西横县六景、广西桂林杨堤、贵州独山大河口—白虎坡、贵州贵阳乌当、四川广元后高坪)的遗迹沉积学系统研究,鉴定和描述遗迹化石15属24种,包括Arenicolites carbonaria,Arenicolites isp.,Chondrites cf.intricatus,Cho...     

Arenig trace fossils from the Salamanca area,western Spain

Trace fossils are recorded at two localities from the ‘Armorican Quartzite’ (Lower Ordovician) in the Salamanca area, western Spain. Sixteen ichnospecies, belonging to twelve ichnogenera, are described; all ichnospecies occur in the upper part of the unit, only four in the lower part. The ichnogenera comprise Arthrophycus, Cruziana, Daedalus, Didymaulichnus, Diplichnites, Diplocraterion, cf. Gor-. dia, Monomorphichnus, Muensteria, Palaeophycus, Planolites, and Rusophycus. The presence of Cruziana furcifera, C. goldfussi, C. rugosa, and C? cf. imbricata indicate that the upper part of the unit is of Arenig age. The lower part has yielded only the two former ichnospecies and may extend down into the Tremadoc. The species of Cruziana and their stratigraphical significance are discussed.     

Misplaced trace fossils in unlikely environments

Graphoglyptids are trace fossils that are generally acknowledged to have had a deep-sea origin, displaying complex, nutrient gathering strategies in response to stable environments with low productivity. However, two recently discovered graphoglyptids in the Permian of New Mexico seem to challenge previous assumptions. Rather than in deep-sea sediments, Spirorhaphe azteca and Augerinoichnus helicoidalis were discovered in tidal flat palaeoenvironments, showing that these behavioural patterns are also carried out by animals in tidal settings. Fine sediments of the region are recognized as the key factor in the preservation of these trace fossils. Presence of graphoglyptids in a transitional environment demonstrates that behaviour is not solely defined by depth, but many intricate factors are involved.     

天山南侧第三系遗迹化石新发现

遗迹化石研究具有多方面的意义,而对沉积环境的解释是其中重要的方面。在天山南侧的诸多新生界剖面上首次发现了大量遗迹化石,为地层界面的认识和进一步分析当时的生物面貌、沉积环境提供了重要的证据。发现了3个遗迹化石相,即Scoyenia(斯柯茵迹)遗迹相、Skolithos(石针迹)遗迹相和Glossifungites(舌菌迹)遗迹相,共计有10余个遗迹化石属,这在以陆相为主的地层中是不多见的。另外,在一些剖面上还发现了大量哺乳类足迹化石,对确定当时生物群的面貌和生态环境具积极意义。     

Upper bathyal trace fossils document palaeoclimate changes

Marine trace fossils were investigated in a 102-m cored section that covers −14.379 to −14.142 Ma (Middle Miocene). Long-term climate trends are perfectly reflected by changing trace fossil communities of the upper bathyal (−200 to −300 m). The community structure is expressed in abundance of taxa based on probabilities of time concordance between the deposition of the sediment and the time, when the traces were produced. Changes in community structure precisely parallel the increasing δ¹³C isotopes during the Middle Miocene climate transition and perfectly mirror oscillations in solar insolation in both amplitudes and phase. Furthermore, the density stratification of the seawater caused by wind-induced currents is reflected in trace fossil communities. Additionally, the dependence of ichnotaxa abundance from threshold values in the sedimentation rate could be demonstrated.     

Upper proterozoic volcanic graywackes from northwestern Hoggar (Algeria) — geology and geochemistry

Late-Proterozoic volcanic graywackes from the NW Hoggar (Algeria) have been investigated from geological and petrological points of view. Thirty-five samples have been analysed for major elements and the following trace elements: Li, Rb, Sr, Ba, Ni, Co, Cr, V, Zn, Cu.These graywackes constitutes a thick flysch-like formation deposited in a marine environment. They are composed of volcanic (mainly andesites) and plutonic detritus. The chemical analyses confirm this observation and demonstrate the very immature character of these rocks.The composition of the NW Hoggar graywackes differs chemically from other investigated graywackes in that they have a higher Ca, Na content and lower SiO₂, Ni content.Several geological observations suggest that the average Hoggar graywacke composition approaches the composition of the W Hoggar Late Precambrian upper crust. The graywackes were formed as a result of erosion of penecontemporaneous calc-alkali volcanic suite, emplaced after widespread mafic intrusions in the Upper Proterozoic shelf deposits.     

Trilobite tracks and other trace fossils from the Upper Cambrian of North Wales

Trace fossils are recorded from eight localities within Upper Cambrian (Ffestiniog Stage) strata in North Wales. Trilobite furrows (Cruziana), resting impressions (Rusophycus), walking and striding tracks (Diplichnites), sideways grazing tracks (Dimorphichnus) and swimming grazing marks (Monomorphichnus, gen. nov.) are described, largely on the basis of over 500 collected specimens, and are figured together with examples of other trace fossil genera including Rouaultia, Phycodes and Monocraterion. The trilobite traces are used to investigate the varied methods of locomotion of the animais. They also indicate the morphology of some of the trilobite hard and soft parts from which it is inferred that most of these traces were probably made by a population consisting of a small number of closely allied species of trilobites belonging to the Olenid family.     

西藏定结、定日三叠系遗迹化石及其地质意义

西藏定结县、定日县一带的三叠系产大量遗迹化石,可划分为4个遗迹化石组合,对应于Skolithos遗迹相,Cruziana遗迹相和Zoophycos遗迹相.根据遗迹相及其它相标志判别,该区早中三叠世主要为陆棚环境,晚三叠世主要为滨岸环境.对遗迹化石的分析结果表明该区早三叠世存在短暂的缺氧事件.     

Cambro‐Ordovician trace fossils from the SW‐Norwegian Caledonides

A diverse trace fossil association is described for the first time from low‐grade metamorphic rocks of the SW‐Norwegian Caledonides. The investigated cliff sections with autochthonous to parautochthonous metasediments comprise a coarsening‐ and thickening‐upward succession interpreted as prograding delta deposits. Sedimentary features indicate a tide‐influenced environment. Twenty‐one ichnospecies have been identified and assigned to the Cruziana and Skolithos ichnofacies, including the oldest record of Beaconites capronus and Macaronichnus segregatis. Cruziana ichnostratigraphy (sensu lato), previously rarely used in other palaeocontinents than Gondwana, allows an age determination for these metasediments of Middle Cambrian to Lower Ordovician as indicated by the presence of Cruziana barbata, C. furcifera, C. rugosa, C. semiplicata, Didymaulichnus rouaulti and Rusophycus ramellensis. Baltica was geographically the most isolated from the other three large continents (Gondwana, Laurentia and Siberia) during Cambro‐Ordovician time, and provinciality of faunal assemblages (e.g. brachiopods, conodonts) has been proved and is also supposed for trilobites by some authors. However, although the Cruziana ichnospecies result from a high specialization of their tracemakers, and therefore only a small group of trilobite species is eligible for its origin, the ichnospecies reported from Baltica occur also on other palaeocontinents and do not support the assumption of trilobite provincialism. Copyright © 2004 John Wiley & Sons, Ltd.     

Possible animal body fossils from the Late Neoproterozoic interglacial successions in the Kimberley region, northwestern Australia

New specimens of the enigmatic Ediacara-type fossil Palaeopascichnus have been identified from the upper part of the Neoproterozoic Ranford Formation in the Kimberley region, northwest Australia. New material is morphologically similar to Palaeopascichnus and represents the largest species of this genus. They resemble the present-day xenophyophore protists in chamber morphology and growth patterns, supporting the interpretation that Palaeopascichnus is possibly a xenophyophore body fossil rather than a trace fossil. Stratigraphic correlation reveals that the new Palaeopascichnus specimens are preserved in the interglacial successions between the Landrigan/Marinoan and Egan/Ediacaran glaciations. If correlation with the early Ediacaran formations of South Australia is accepted, this represents the earliest known identifiable member of the Ediacara biota. New fossil record fills the evolutionary gap between the Cryogenian and Ediacaran animal assemblages and well-known Ediacaran biota. The new Palaeopascichnus specimens represent the first record of Ediacara-type fossils in Kimberley, and suggest the probability that additional Ediacaran fossils may be found in northwestern Australia.     

新疆吐哈盆地侏罗纪湖相动物遗迹化石的发现及古环境意义

对首次在吐哈盆地台北凹陷侏罗系发现的分布于不同剖面、不同层位中的动物遗迹化石进行了系统研究。主要分析了研究区内动物遗迹化石的特征 ,将之划分为六大类型 :( 1 )垂直居住迹 ;( 2 )垂直觅食迹 ;( 3 )“U”形管 ;( 4)潜穴系统 ;( 5)逃逸迹 ;( 6)停息迹。较详细地分析了各类型的形态特征和保存特点 ,简要分析了造迹生物的行为习性及环境意义。在此基础上 ,建立了本区动物遗迹化石的环境模式。同时指出 ,遗迹化石在陆相含油气盆地确实大量存在 ,并在储层研究中具广泛的实践意义。     

Diminutive trace fossils in the Chengjiang Lagerstätte

Diminutive trace fossils described here have been well preserved in connection with certain pliable carapaces or skeletons of the Chengjiang animals. Taphonomic evidence suggests that these traces were formed in situ. The peculiar preservation indicates that some bilaterians of uncertain affinities seem to have been active at the burial site, constituting an important group of the hidden majority of Cambrian marine community. This may be an essential feature closely associated with the palaeoecology and exceptional preservation of the Cambrian fossil Lagerstätten. The rare preservation of such millimetre‐sized trace fossils raises further doubts about the timing of the initial benthic bilaterians by applying trace fossil evidence.     

下扬子区下三叠统遗迹化石及其环境意义

<正> 遗迹化石是地质时期各类生物在生活期间进行居住、运动、寻食、代谢和生殖等行为习性活动,在沉积物表面或其内部留下的各种痕迹。作为重要的生物标志,遗迹化石可用来判别和恢复古代沉积环境。近年来,笔者在下扬子区的宿松、贵池、泾县、宁国、宜兴等地(图1)的下三叠统,发现大量遗迹化石。本文从遗迹化石组合特征出发,讨论遗迹化石与沉积环境之间的关系。承杨式溥教授鉴定遗迹化石标本,在此深表谢忱。     

Pliocene trace fossils from oyster substrates in the Nijar Basin,Betic Cordillera,southern Spain

The Almería-Níjar Basin is a Neogene, intermontane depression marginal to the Mediterranean in southern Spain in the vicinity of El Argamasón, Andalucia. The Pliocene Cuevas Formation rests unconformably on the Upper Messinian rock succession in the Carboneras Fault Zone. The Cuevas Formation is a coarse-grained, bioclastic-rich, calcarenite to calcirudite shoreface deposit. Oysters, namely Saccostrea cucullata (Born), are locally common and preserve a moderate diversity of borings: Caulostrepsis taeniola Clarke; Entobia isp.; Gastrochaenolites isp. aff. G. lapidicus Kelly and Bromley; Maeandropolydora isp. cf. M. sulcans Voigt; Oichnus paraboloides Bromley; and Talpina isp. aff. T. hirsuta Voigt. All represent domiciles except for the predatory O. paraboloides trace. This suite of ichnotaxa is assigned to the Entobia ichnofacies sensu Bromley and Asgaard; they are comparable, particularly, with the Boulder Assemblage of the Pliocene of Rhodes, Greece. Physical disturbance is an important parameter in favouring this pattern of infestation, whether the bored clasts are boulders or oyster valves.     

Palaeoenvironmental significance of trace fossils from the palaeocene sediments of the Pondicherry area,South India

Sediments of Manaveli Formation (Palaeocene) are studied from the Thondaimanattam quarry section and Manaveli section of the Pondicherry area, which comprises of bioturbated arenaceous limestone-shale and shaly-sandstone sequences respectively. These sediments consist of high density and low diversity trace fossil genera which includes Lanicodichnus, Ophiomorpha, Planolites, Palaeophycus, Skolithos, Thalassinoides, and Polychaetes tubes. The abundance of horizontal structures of the Thondaimanattam quarry section indicate low to moderate wave and current energy in fully marine subtidal environments while the presences of vertical structures of the Manaveli section indicate moderate to high wave and current energy in the marginal marine environment of deposition for the Palaeocene sediments of the Pondicherry area.