A new species of Vectiselachos (Chondrichthyes,Selachii) from the Early Cretaceous of southern England

Teeth of a new species of hybodont shark Vectiselachos (Chondrichthyes: Lonchidiidae) are described from the late Aptian (Early Cretaceous) of southern England. Vectiselachos gosslingi sp. nov. has very distinctive coarse striations that form raised ridges over the occlusal surfaces of the crown.     

New genera and species of bark-gnawing beetles (Coleoptera: Trogossitidae) from the Yixian Formation (Lower Cretaceous) of Western Liaoning,China

Two new genera from the Lower Cretaceous Yixian Formation of Beipiao City, Liaoning Province, northeastern China are described and illustrated. Paracretocateres gen. nov. (type species P. bellus, sp. nov.) and Yixianteres gen. nov. (type species Y. beipiaoensis sp. nov.) are assigned to the subfamily Lophocaterinae based on the state of the procoxae which are not projecting, the transverse procoxal cavities, and the exposed protochantins. The new taxa broaden the diversity of this family and provide further morphological characters for phylogenetic studies of Trogossitidae. They also provide evidence that lophocaterines well-diversified prior to the trogossitines and their biologies may have been similar to extant forms.     

A new species of Cyrtophyllitinae (Insecta: Ensifera) from the Cretaceous China

A new fossil species of Cyrtophyllitinae, Vitimoilus ovatus sp. nov., is described from the Dabeigou Formation and Yixian Formation of the Lower Cretaceous of China. It is clearly placed in Vitimoilus Gorochov, 1996 according to the following characters: R forking into RA and RP distally, M forking into MA and MP distally; CuA fused with CuPaα distal of the middle of the wing length; cross-veins in basal part of CuPb–CuPaβ area non-strongly curved.     

中国辽宁义县早白垩世燕鸟一新种

辽宁锦州义县早白垩世义县组发现了一种新型燕鸟(Yanornis)化石.该新型化石具有燕鸟的典型特征,如齿骨直且约占头骨全长的2/3、每侧约含20枚牙齿,颈椎细长,异凹型.愈合荐椎包括9枚脊椎,前肢约为后肢长的1.1倍.近端脚趾较其它趾节长和粗壮.然而,该新型化石部分特征又不同于燕鸟的模式种——马氏燕鸟(Yanornis martini),如其肱骨三角肌脊强烈发育,约为肱骨长度的一半.尺骨、桡骨及手部近等长,均长于肱骨.耻骨联合部较短,与尾综骨等长,不到耻骨长度的1/6.腓骨较长,约为股骨长度的4/5,约为胫跗骨长度的2/3.跗跖骨较短并完全愈合,不及胫跗骨长度一半.根据该新型化石的独特性及其与义县鸟、松岭鸟和马氏燕鸟的特征对比,笔者建立了燕鸟一新种——国章燕鸟(Yanornis guozhangisp.nov.).此外,国章燕鸟(新种)颈部还保存有3条鱼化石,这一特殊现象为研究燕鸟的生活习性和发育模式提供了重要信息.     

First Valanginian Polacanthus foxii (Dinosauria,Ankylosauria) from England,from the Lower Cretaceous of Bexhill,Sussex

A new partial skeleton of the armoured ornithischian dinosaur Polacanthus found in the Wadhurst Clay Formation (Valanginian stage) of Bexhill, Sussex is the oldest recorded occurrence of this taxon. Previous discoveries suggested that at least two armoured ornithischians occur in the Wealden succession: Polacanthus, which was mostly restricted to the Barremian, and Hylaeosaurus, which was recorded as present only in the Valanginian. The new discovery extends the stratigraphic range of Polacanthus into the Valanginian. Although these two taxa appear to be closely similar anatomically, their osteology now suggests they are not synonymous. The new specimen includes the first known jugal as well as a comparatively rare polacanthid plate/spine (splate) which probably comes from the shoulder (pectoral) area of these animals.     

New fossil species of the Venicoridae (Heteroptera: Pentatomomorpha) from the Lower Cretaceous of Northeast China

Specimens of the Venicoridae (Heteroptera) occur in the Lower Cretaceous Yixian Formation in northeastern China. We describe a new genus and two new species Halonatusivena shii gen. et sp. nov. and Halonatusivena nervosus sp. nov. from this formation. The veins of the membrane are distinctive; the edges of veins are not simple but resemble an irregular stain, with a reduced vasiform structure previously unknown in the Heteroptera. Remarkably, the antennae of one of the specimens of Halonatusivena shii exhibit morphological abnormality. An identification key for the known species of the Venicoridae is provided.     

A new species of Mirovia (Coniferales, Miroviaceae) from the Lower Cretaceous of the Iberian Ranges (Spain)

The Lower–Middle Albian coaly clay bed of the Escucha Formation, which is exposed at Rubielos de Mora (eastern Iberian Ranges, Spain), contains a diverse fossil plant assemblage. Among the taxa present in this layer, Mirovia gothanii Gomez sp. nov. differs from other species of the genus by its greater leaf length, margins typically overhanging the depressed stomatal groove, a single short, blunt, papilla borne by each subsidiary cell, non-stomatal cells inside the groove and margins, and a higher number of resin ducts in the mesophyll. Morphological study of the well-preserved cuticles demonstrates that the species also occurs in Lower Cretaceous coals of Santa Maria de Meià (Pyrenees, Spain) where Gothan (1954) described it as Sciadopitytes sp. Both localities constitute the southernmost extent of the genus in Laurasia when the family was likely to have reached its climax in terms of abundance and diversity.     

A new species of Taxodioxylon from the Lower Cretaceous of the Jixi Basin, eastern Heilongjiang, China

Newly discovered silicified wood from the Jixi Basin, Heilongjiang Province, northeast China, represents the first record of Taxodioxylon sensu Gothan from the Lower Cretaceous of China. Taxodioxylon szei sp. nov. is characterized by abundant wood parenchyma, uniseriate rays 1–50 cells in height, 1–2 rows of separately or contiguously arranged bordered pits, taxodioid-type pits in cross field, no normal resin canals and an absence of spiral thickenings in tracheids. It agrees well with the diagnostic characters of Taxodioxylon, but is distinguished from previously described species by higher rays, the presence of idioblasts in some rays, and by having black granules, which seem to be clusters of a needle-like substance (crystals?) in axial wood parenchyma cells, and traumatic resin canals. The anatomy of T. szei, and other palaeobotanical and sedimentological evidence indicate that the Early Cretaceous climate of the Jixi Basin was seasonal and very favourable to tree growth.     

Ptychodus decurrens Agassiz (Elasmobranchii: Ptychodontidae) from the Upper Cretaceous of India

Although a very high invertebrate faunal diversity is known from the outcrops of the Ariyalur group in the Cauvery Basin, southern India, little is known about its vertebrate fauna. Recent fieldwork in the badland exposures of the Karai Formation (Upper Cenomanian-Lower Turonian) near Garudamangalam in the basin has yielded two teeth belonging to the Late Cretaceous shark Ptychodus decurrens (Ptychodontidae). The fossil record of Ptychodus decurrens from the southern continents is very poor, being known from a single Late/Middle Albian occurrence in Australia. This finding documents the first record of fossil P. decurrens in India and second from a Gondwanan landmass, and provides the first evidence of a cosmopolitan, Pangaean, distribution of the species during the Albian-Turonian and additional insights into the palaeoecology of the Cauvery Basin during the deposition of the Karai Formation.     

Discovery of three new mordellids (Coleoptera, Tenebrionoidea) from the Yixian Formation of western Liaoning, China

In this paper one new genus and three new species Bellimordella capitulifera gen. et sp. nov., B. longispina gen. et sp. nov., B. robusta gen. et sp. nov., all placed within the subfamily Praemordellinae of the family Mordellidae, are described and illustrated. The specimens were recovered from the Yixian Formation of Beipiao City, Liaoning Province, NE China. The key to the species of this new genus is given. Furthermore, we discuss evolution of the pygidium and possible food for those ancient mordellids.     

A New Troodontid (Theropoda: Troodontidae) from the Lower Cretaceous Yixian Formation of Western Liaoning, China

A specimen collected from the Lower Cretaceous Yixian Formation of western Liaoning, China, represents a new genus and species of troodontid theropod. The new taxon is named and described on the basis of the holotype and the only known specimen, which comprises an articulated skeleton with the presacral vertebral, shoulder girdle and forelimbs missing as preserved. Diagnostic features of the new species include nasals that are sinusoid in lateral view, absence of a passage connecting the antorbital and maxillary fenestrae, relatively large teeth, plate-like chevrons forming a band along most of the length of the tail, and a long neck between the femoral head and shaft. The temporal constraints of the three paravian groups (Troodontidae, Dromaeosauridae and Aves) combined with the character distributions among the earliest known troodontids indicate a rapid evolution at the base of the Troodontidae.     

The status of Dollodon and other basal iguanodonts (Dinosauria: Ornithischia) from the Lower Cretaceous of Europe

Dollodon bampingi was recently named based upon a specimen from the Bernissart Quarry that had previously been referred to Mantellisaurus atherfieldensis. The initial diagnosis of Dollodon did not adequately distinguish it from Mantellisaurus or from other basal iguanodonts, necessitating a reassessment of the material. Firsthand examination of the holotypes of the two taxa and numerous other basal iguanodont specimens, as well as a principal components analysis of basal iguanodont dentaries, did not find any morphological features to justify the distinction of Dollodon from Mantellisaurus. D. bampingi is thus best considered a junior synonym of M. atherfieldensis. Furthermore, the recent referral of the species Iguanodon seelyi to the genus Dollodon is not supported; I. seelyi is indistinguishable from Iguanodon bernissartensis, and is considered a junior synonym of that species. Finally, the recently named taxon Proplanicoxa galtoni, also based upon a specimen formerly attributed to M. atherfieldensis, is considered a nomen dubium and probable junior synonym of M. atherfieldensis. Thus, only two species of large-bodied basal iguanodont should be recognized from the Barremian-Aptian of England and Belgium: M. atherfieldensis and I. bernissartensis.     

Systematic revision of the fossil snakefly family Baissopteridae (Insecta: Raphidioptera) from the Lower Cretaceous of China,with description of a new genus and three new species

The fossil snakefly family Baissopteridae from the Lower Cretaceous (upper Barremian) of the Yixian Formation of Liaoning Province, China is reviewed. Two genera and five species of Baissopteridae are recorded, including a new genus, namely Microbaissoptera gen. nov., and three new species, i.e., Baissoptera bicolor sp. nov., Baissoptera sinica sp. nov., and Microbaissoptera monosticha sp. nov. Moreover, Baissoptera euneura Ren, 1997 is herein treated as a new junior synonymy of Baissoptera grandis Ren in Ren et al., 1995. Baissoptera minima Ponomarenko, 1993 is transferred to the presently described new genus Microbaissoptera gen. nov. based on the small body-size and the presence of a single gradate series of crossveins in radial and medial areas of posterior part of both fore- and hind wings. A key to genera of Baissopteridae is provided.     

The Olduvai buffalo Pelorovis and the origin of Bos

The origin of the genus Bos is a debated issue. From ∼ 0.5 Ma until historic times, the genus is well known in the Eurasian large mammal assemblages, where it is represented by Bos primigenius. This species has a highly derived cranial anatomy that shows important morphological differences from other Plio-Pleistocene Eurasian genera of the tribe Bovini such as Leptobos, Bison, Proamphibos-Hemibos, and Bubalus. The oldest clear evidence of Bos is the skull fragment ASB-198-1 from the middle Pleistocene (∼ 0.6-0.8 Ma) site of Asbole (Lower Awash Valley, Ethiopia). The first appearance of Bos in Europe is at the site of Venosa-Notarchirico, Italy (∼ 0.5-0.6 Ma). Although the origin of Bos has traditionally been connected with Leptobos and Bison, after a detailed anatomical and morphometric study we propose here a different origin, connecting the middle Pleistocene Eurasian forms of B. primigenius with the African Late Pliocene and early Pleistocene large size member of the tribe Bovini Pelorovis sensu stricto. The dispersal of the Bos lineage in Western Europe during middle Pleistocene times seems to coincide with the arrival of the Acheulean tool technology in this continent.     

A New Specimen of Liaoceratops yanzigouensis (Dinosauria:Neoceratopsia) from the Early Cretaceous of Liaoning Province, China

A new specimen of basal neoceratopsian dinosaur Liaoceratops yanzigouensis is described. The specimen comes from the Lujiatun Bed of the Lower Cretaceous Yixian Formation in Beipiao City of western Liaoning Province, and is represented by a very well preserved three-dimensional partial skull and mandible. It is also the smallest among the three specimens of L. yanzigouensis, and several features in the new specimen, such as the short preorbital length and the round rostroventral orbital rim, can be ontogenetically-related. The superb exposure of the palatal complex may be caused by the removing of its brain by a small predator in the contemporary Jehol Biota.     

The postglacial history of three Picea species in New England, USA

Given the difficulty of separating the three Picea species—P. glauca, P. mariana, and P. rubens (white, black, and red spruce)—in the pollen record, little is known about their unique histories in eastern North America following deglaciation. Here we report the first use of a classification tree analysis (CART) to distinguish pollen grains of these species. It was successfully applied to fossil pollen from eight sites in Maine and one in Massachusetts. We focused on the late glacial/early Holocene (14,000 to 8000 cal yr B.P.) and the late Holocene (1400 cal yr B.P. to present)—the two key periods since deglaciation when Picea has been abundant in the region. The result shows a shift from a Picea forest of P. glauca and P. mariana in the late glacial to a forest of P. rubens and P. mariana in the late Holocene. The small number of P. rubens grains identified from the late glacial/early Holocene samples (<5%) suggests that that species was either absent or rare at most of the sites. The occurrence and distribution of the three species do not reveal any geographic or temporal trend during late glacial time, but the data suggest that they were distributed in local patches on the landscape. The results of this study indicate that the recent population expansion of Picea (1000 to 500 cal yr B.P.) was likely the first time since deglaciation that P. rubens was abundant in the region.     

Latest Pleistocene paleoecology of Jefferson's ground sloth (Megalonyx jeffersonii) and elk-moose (Cervalces scotti) in northern Illinois

New records of Jefferson's ground sloth (Megalonyx jeffersonii) and elk-moose (Cervalces scotti) from Lang Farm provide the first precise temporal correlation of these taxa with the specific environments inhabited by them near the time of their extinction. Six AMS ¹⁴C measurements establish an age of 11,405 ± 50 ¹⁴C yr B.P. for Lang Farm Cervalces and an age of 11,430 ± 60 or 11,485 ± 40 ¹⁴C yr B.P. for the Megalonyx. These measurements represent the youngest ¹⁴C dates for these two genera based on direct dating. Comparison of the dates with pollen data from northern Illinois indicates that these species inhabited a nonanalog environment that was transitional from mid-latitude tundra to mixed conifer and deciduous woodland. Although spruce (Picea sp.) was dominant, it was less abundant than prior to 12,500 ¹⁴C yr B.P. The presence of black ash (Fraxinus nigra) and fir (Abies sp.) indicates a wet climate and heavy winter precipitation. This may have been the preferred habitat for Cervalces because of its narrow geographic range. However, this habitat type was only one of many occupied by Megalonyx as indicated by its broad geographic distribution.     

Teredolites Leymerie in the Lower Greensand Group (Cretaceous) of the Isle of Wight and the problematic ichnology of reworked clasts

Nineteenth-century references to clavate borings in woody substrates in the Lower Greensand of the Isle of Wight used a variety of names, but Teredo (a wood-boring bivalve, not a boring), Teredolithes (a junior synonym of Teredolites) and Gastrochaena (a bivalve borer of rock and shelly substrates, not a boring in wood) are all nomenclatorially incorrect. Borings in a beach clast derived from the Lower Greensand Group and recently collected from Sandown Bay, Isle of Wight, are referred to Teredolites isp. cf. T. longissimus Kelly and Bromley. This specimen confirms the presence of Teredolites in the Lower Greensand Group and demonstrates a common ichnological problem of beach clasts; borings, either fossil or modern, are incompletely preserved, making confident classification below the level of ichnogenus problematic.