Collection of radiocarbon dates on the mammoths (Mammuthus primigenius) and other genera of Wrangel Island, northeast Siberia, Russia

We present and discuss a full list of radiocarbon dates for woolly mammoth and other species of the Mammoth fauna available from Wrangel Island, northeast Siberia, Russia. Most of the radiocarbon dates are published here for the first time. Of the124 radiocarbon dates on mammoth bone, 106 fall between 3700 and 9000 yr ago. We believe these dates bracket the period of mammoth isolation on Wrangel Island and their ultimate extinction, which we attribute to natural causes. The absence of dates between 9–12 ka probably indicates a period when mammoths were absent from Wrangel Island. Long bone dimensions of Holocene mammoths from Wrangel Island indicate that these animals were comparable in size to those on the mainland; although they were not large animals, neither can they be classified as dwarfs. Occurrence of mammoth Holocene refugia on the mainland is suggested. Based on other species of the Mammoth fauna that have also been radiocarbon on Wrangel Island, including horse, bison, musk ox and woolly rhinoceros, it appears that the mammoth was the only species of that fauna that inhabited Wrangel Island in the mid-Holocene.     

The North of Eastern Siberia: Refuge of Mammoth Fauna in the Holocene

The global climate changings at the end of Pleistocene led to extinction of the typical representatives of Mammoth fauna–mammoth, woolly rhinoceros, wild horse, bison, muskox, cave lion, etc.–on the huge territories of Northern Eurasia. Undoubtedly the Mammoth fauna underwent pressure from the Upper Paleolithic Man, whose hunting activity also could play the role in decreasing the number of mammoths and other representatives of megafauna (large mammals). Archaeological data testify that the typical representatives of Mammoth fauna were the Man's hunting objects only till the end of the Pleistocene. Their bone remains are not usually found on the settlements of Mesolithic Man. Formerly it was supposed that the megafauna of ‘Mammoth complex’ was extinct by the beginning of Holocene. Nevertheless the latest data testify that the global extinction of the Mammoth fauna was sufficiently delayed in the north of Eastern Siberia. In the 1990s some radiocarbon data testified that the mammoths on the Wrangel Island existed for a long time during the Holocene from 8000 till 3700 y. BP. The present radiocarbon data show that wild horses inhabited the north of Eastern Siberia (the lower stream of the Enissey river, the Novosibirskie Islands, the East Siberian sea-shore) 3000–2000 y. BP. Musk-oxen lived on the Taimyr Peninsula and the Lena River delta about 3000 y. BP. Some bison remains from Eastern Siberia belong to the Holocene. The following circumstances could promote the process of preservation of the Mammoth fauna representatives. The cool and dry climate of this region promotes the maintenance of steppe associations – habitats of those mammals. The Late Paleolithic and Mesolithic settlements are not found in the Arctic zone of Eastern Siberia from the Taimyr Peninsula to a lower stream of the Yana River; they are very rare in the basins of the Indigirka and Kolyma Rivers. So, the small number of the Stone Age hunting tribes on the North of Eastern Siberia was another factor in the long-term preservation of some Mammoth fauna representatives.     

Last straw versus Blitzkrieg overkill: Climate-driven changes in the Arctic Siberian mammoth population and the Late Pleistocene extinction problem

A set of radiocarbon dates on woolly mammoth were obtained from several regions of Arctic Siberia: the New Siberian Islands (n = 68), north of the Yana-Indigirka Lowland (n = 43), and the Taimyr Peninsula (n = 18). Based on these and earlier published dates (n = 201) from the East Arctic, a comparative analysis of the time-related density distribution of ¹⁴C dates was conducted. It was shown that the frequencies of ¹⁴C dates under certain conditions reflect temporal fluctuations in mammoth numbers. At the end of the Pleistocene the number of mammoths in the East Arctic changed in a cyclic manner in keeping with a general “Milankovitch-like” trend. The fluctuations in numbers at the end of the Pleistocene occurred synchronously with paleoenvironmental changes controlled by global climatic change. There were three minima of relative mammoth numbers during the last 50 000 years: 22 000, 14 500–19 000, and 9500 radiocarbon years ago, or around 26 000, 16–20 000, and 10 500 calendar years respectively. The last mammoths lived on the New Siberian Islands, which were connected to the continent at that time, 9470 ± 40 radiocarbon years ago (10 700 ± 70 calendar years BP). This new youngest date approximates the extinction time of mammoths in the last continental refugium of the Holarctic. The adverse combination of environmental parameters was apparently a major factor in the critical reduction in mammoth numbers. The dispersal of humans into the Arctic areas of Siberia no later than 28 000 radiocarbon years ago did not overtly influence animal numbers. Humans were not responsible for the destruction of a sustainable mammoth population. The expanding human population could have become fatal to mammoths during strong the minima of their numbers, one of which occurred at the very beginning of the Holocene.     

Patterns of faunal extinction and paleoclimatic change from mid-Holocene mammoth and polar bear remains, Pribilof Islands, Alaska

Qagnaxˆ Cave, a lava tube cave on St. Paul Island in the Pribilofs, has recently produced a mid-Holocene vertebrate faunal assemblage including woolly mammoth, polar bear, caribou, and Arctic fox. Several dates on the mammoth remains converge on 5700 ¹⁴C yr BP. These dates, ~ 2300 yr younger than mammoth dates previously published from the Pribilof Islands, make these the youngest remains of proboscideans, and of non-extinct Quaternary megafauna, recovered from North America. Persistence of mammoths on the Pribilofs is most parsimoniously explained by the isolation of the Pribilofs and the lack of human presence in pre-Russian contact times, but an additional factor may have been the local existence of high-quality forage in the form of grasses enriched by nutrients derived from local Holocene tephras. This interpretation is reinforced by stable carbon and nitrogen isotope values obtained from the mammoth remains. The endpoint of mammoth survival in the Pribilofs is unknown, but maybe coterminous with the arrival of polar bears whose remains in the cave date to the Neoglacial cold period of ~ 4500 to 3500 ¹⁴C yr BP. The polar bear record corroborates a widespread cooling of the Bering Sea region at that time.     

Arctic Siberia: refuge of the Mammoth fauna in the Holocene

Global climate change at the end of Pleistocene led to extinction in the huge territories of Northern Eurasia of the typical representatives of the Mammoth fauna: mammoth, woolly rhinoceros, wild horse, bison, musk-ox, and cave lion. Undoubtedly the Mammoth fauna underwent pressure from Upper Paleolithic humans, whose hunting activity could also have played a role in decreasing the number of mammoths and other representatives of megafauna. Formerly it was supposed that the megafauna of the “Mammoth complex” had become extinct by the beginning of the Holocene. Nevertheless the latest data indicate that extinction of the Mammoth fauna was significantly delayed in the north of Eastern Siberia. In the 1990s some radiocarbon dates established that mammoths existed in the Holocene on Wrangel Island—from 7700 until 3700 yBP. Radiocarbon data show that wild horses inhabited the north of Eastern Siberia 4600–2000 yBP. Muskoxen lived here about 3000 yBP. Some bison remains from Eastern Siberia belong to the Holocene. The following circumstances could promote the survival of representatives of Mammoth fauna. Cool and dry climate in this region promotes the maintenance of steppe associations—the habitats of those mammals. Late Paleolithic and Mesolithic settlements are not found in the Arctic zone of Eastern Siberia from Taimyr Peninsula to the lower Yana River; they are very rare in basins of the Indigirka and Kolyma Rivers. The small number of Stone Age hunting tribes in the northern part of Eastern Siberia was probably another factor that contributed to the survival of some Mammoth fauna representatives.     

Extinction of the woolly mammoth (Mammuthus primigenius) and woolly rhinoceros (Coelodonta antiquitatis) in Eurasia: Review of chronological and environmental issues

Kuzmin, Y. V. 2009: Extinction of the woolly mammoth (Mammuthus primigenius) and woolly rhinoceros (Coelodonta antiquitatis) in Eurasia: Review of chronological and environmental issues. Boreas, 10.1111/j.1502‐3885.2009.00122.x. ISSN 0300‐9483. The current evidence for date and environmental preferences of the extinction of two middle–late Pleistocene megafaunal species, the woolly mammoth (Mammuthus primigenius Blum.) and woolly rhinoceros (Coelodonta antiquitatis Blum.), is presented in this review. It is suggested that extinction of these large herbivores in Eurasia was closely related to landscape changes near the Pleistocene–Holocene boundary (c. 12 000–9000 uncalibrated radiocarbon years ago, yr BP), mainly involving the widespread forest formations in the temperate and arctic regions of northern Eurasia and the loss of grasslands crucial to the existence of woolly mammoth and rhinoceros. However, some woolly mammoth populations survived well into the Holocene (up to c. 3700 yr BP), showing that the process of final extinction was fairly complex, with delays in some regions of up to several millennia. The possible role of Palaeolithic humans in the extinction of Late Pleistocene megafauna is also considered.     

Population-level genotyping of coat colour polymorphism in woolly mammoth (Mammuthus primigenius)

Patterns in the spatial or temporal distribution of genotypes may be indicative of natural selection. Previous work on the woolly mammoth melanocortin-1 receptor (Mc1r) gene identified three polymorphic positions that suggest Pleistocene populations may have harboured both light- and dark-haired mammoths (Rompler et al., 2006, 313: 62). Here, we extend this work and present the first population-level analysis of a functional gene in an extinct species. We genotyped the Mc1r gene in 47 woolly mammoth samples excavated from sites across the central portion of the woolly mammoths’ former range to examine the extent of variation of this polymorphism through time and across space. Only one individual was found to be heterozygous, indicating that the frequency of the ‘light’ mutant allele was very low. We conclude that light-coloured woolly mammoths would have been very rare, and may even have been non-existent if the ‘light’ mutant allele was strongly selected against in its homozygotic form. With the increasing availability of large-scale sequencing technologies, population-level datasets capable of identifying local adaptation will become increasingly attainable.     

The Sevsk woolly mammoth (Mammuthus primigenius) site in Russia: Taphonomic,biological and behavioral interpretations

Excavations at Sevsk, Bryansk Region, Russia, by the Paleontological Institute of the Russian Academy of Sciences in 1988–1991 recovered 3800 bones of woolly mammoth (Mammuthus primigenius Blum.) representing a minimum of 33 individuals. The locality is one of the largest naturally occurring deposits of mammoth remains in Europe and is inferred to be a catastrophic death assemblage. The material includes five skeletons of juvenile mammoths, from 1 month to 6 or 7 years of age, as well as partial skeletons and isolated bones of adult individuals. A femur and humerus of an approximately 10–12-month-old fetus are also among the finds. Morphological features suggest that the Sevsk mammoths belonged to one family group; the age structure and sexual composition of the assemblage do not differ significantly from that of a family group of Modern African elephants. In contrast to other localities in Siberia and central Russia, relatively more (about 45% of individuals) prepubertal animals are preserved at Sevsk. Radiocarbon dates indicate that the mammoths died about 14,000 years ago. Data from diatoms, pollen and rodents, as well as archeological evidence, corroborate this age, and provide the basis for a paleoenvironmental reconstruction at the end of the Valdaian Glaciation in western Russia.     

Pleistocene vertebrates of the Yukon Territory

Unglaciated parts of the Yukon constitute one of the most important areas in North America for yielding Pleistocene vertebrate fossils. Nearly 30 vertebrate faunal localities are reviewed spanning a period of about 1.6 Ma (million years ago) to the close of the Pleistocene some 10 000 BP (radiocarbon years before present, taken as 1950). The vertebrate fossils represent at least 8 species of fishes, 1 amphibian, 41 species of birds and 83 species of mammals. Dominant among the large mammals are: steppe bison (Bison priscus), horse (Equus sp.), woolly mammoth (Mammuthus primigenius), and caribou (Rangifer tarandus) – signature species of the Mammoth Steppe fauna (Fig. 1), which was widespread from the British Isles, through northern Europe, and Siberia to Alaska, Yukon and adjacent Northwest Territories. The Yukon faunas extend from Herschel Island in the north to Revenue Creek in the south and from the Alaskan border in the west to Ketza River in the east. The Yukon holds evidence of the earliest-known people in North America. Artifacts made from bison, mammoth and caribou bones from Bluefish Caves, Old Crow Basin and Dawson City areas show that people had a substantial knowledge of making and using bone tools at least by 25 000 BP, and possibly as early as 40 000 BP. A suggested chronological sequence of Yukon Pleistocene vertebrates (Table 1) facilitates comparison of selected faunas and indicates the known duration of various taxa.     

Revised radiocarbon ages on woolly rhinoceros (Coelodonta antiquitatis) from western central Scotland: significance for timing the extinction of woolly rhinoceros in Britain and the onset of the LGM in central Scotland

Woolly rhinoceros bones, from a number of sites in Britain, have been AMS radiocarbon dated following ultrafiltration pre-treatment. These determinations give a coherent set of ages between >50 and c. 35 cal ka BP. The youngest (35,864–34,765 cal BP) come from the area around Bishopbriggs in western central Scotland and are derived from glaciofluvial sand and gravel overlain by till, both deposited during the Last Glacial Maximum (LGM) glaciation. A previous radiocarbon date from the site suggested that woolly rhinoceros lived c. 27 ¹⁴C ka BP and the region was ice-free at the time. This date has had significant influence on the timing of extinction of woolly rhinoceros and the onset of glaciation over Britain during the LGM. The new dates revise this earlier determination and confirm that woolly rhinoceros became extinct in Britain after c. 35 cal ka BP, that central Scotland was ice-free at this time, and glaciation extended across this region sometime after 35 cal ka BP.     

Woolly mammoth (Mammuthus primigenius Blum.) and its environment in northern Europe during the last glaciation

Woolly mammoths were large, herbivorous, cold-adapted mammals of the Late Pleistocene. The diet and habitat requirements of the species set certain constraints on the palaeoenvironments it could occupy. The relationship between the mammoth’s shifting range and changing environments can be explored using independent data on ice sheet configuration, temperature, and vegetation, provided the locality and age of the fossil remains can be validated. Here we present a comprehensive record of occurrence of the woolly mammoth in the circum-Baltic region of northern Europe during the last glaciation, based on a compilation of radiocarbon-dated remains. The record shows that the mammoth was widespread in northern and north-eastern Europe during Marine Isotope Stage 3 (MIS 3), at 50,000–30,000 calibrated years ago (50–30 ka). The presence of the species up to 65°N latitude supports the restriction of the Scandinavian Ice Sheet (SIS) during MIS 3. The widest distribution range round 30 ka was followed by a decline that led to the disappearance of mammoths from the area during the maximum extent of the SIS, from 22 to 18 ka. The woolly mammoth re-colonized the Baltic region and southern Scandinavia after the onset of the late-glacial deglaciation at 17 ka. The late-glacial record suggests a markedly fluctuating population changing its range in tune with the rapid environmental changes. The last appearance of mammoth in our study region was in Estonia during the Younger Dryas (Greenland Stadial 1; GS1) at about 12 ka. The two major periods of occurrence during MIS 3 and the late-glacial stadial suggest that mammoth had a wide tolerance of open to semi-open tundra and steppe-tundra habitats with intermediately cold climate, whereas the 22–18 ka disappearance suggests a major southward and/or eastward retreat in response to extremely cold, glacial conditions near the SIS margin. The final regional extinction correlates with the re-forestation during the rapid warming at the Younger Dryas–Holocene boundary.     

Mammal fauna during the Late Pleistocene and Holocene in the far northeast of Europe

The paper summarises materials on the mammal remains in northeastern Europe, dated by radiocarbon. Altogether, 23 local faunas of small mammals and 47 local faunas of large mammals were analysed. Multidimensional statistical analysis shows a strong correlation between changes in small mammal fauna composition and climate changes throughout time. The correlations with the spatial gradients, however, are less pronounced. The faunas are classified into three groups: (1) faunas of Holocene age; (2) Late Pleistocene ‘stadial’ assemblages; and (3) Late Pleistocene ‘interstadial’ assemblages. In some cases, changes in species abundance are better understood in terms of biotic interrelations rather than of climatic effects. The most pronounced change in small mammal fauna composition and structure occurred at the Preboreal/Boreal boundary, and a less conspicuous alteration took place at the LGM/Lateglacial transition. The most noticeable transformation in the large mammal fauna composition is dated to the early Holocene. Less significant changes are observed at the Middle Weichselian/LGM transition and at the LGM/Lateglacial transition. It is safely concluded that variations in the faunas of small and large mammals recorded in NE Europe during the last 35 000 years occurred synchronously and unidirectionally.     

AMS 14C chronology of the world's southernmost woolly mammoth (Mammuthus primigenius Blum.)

The world's definite southernmost woolly mammoth record is a molar from Ji’nan (around 36°N), Shandong Province, China. AMS ¹⁴C dating of the specimen, gave a conventional ¹⁴C age of 33,150±250 BP. The period of 40–30 ka BP corresponds to the later phase of the Marine Oxygen Isotope Stage 3 (MIS 3a), recognized as the global interstadial of the last glacial period. However, it is known that the winter monsoon strengthened in Asia during the period 35–33 ka BP, and the age of the woolly mammoth specimen from Ji’nan corresponds to that age. The specimen suggests that this area became cold and dry in 33 ka BP, and grassland or open forest, suitable habitat for woolly mammoths, developed during this short time span. This age is similar to the age of the southernmost woolly mammoth record in Europe, therefore supporting a hypothesis by Porter and An [1995. Correlation between climate events in the North Atlantic and China during the last glaciation. Nature 375, 305–308] that an important component of Chinese palaeoclimates may be linked to changes in North Atlantic oceanic conditions.     

The taphonomy of mammoth localities in southeastern Wisconsin (USA)

Four southeastern Wisconsin mammoth localities located within a glacial landscape had well-preserved remains found in inter-morainal depressions filled with lacustrine clays covered with peat. Numerous radiocarbon ages dated the mammoths to the late Pleistocene. The taphonomic analysis focused on determining the agency or agencies involved in site formation and the agency or agencies involved in modification to the bones. Statistical approaches to bone orientation data underscored that water transport was not a factor in the formation of the bone beds nor was water movement a disturbance factor. The bone modification profile of the overall assemblage was dominated by chemical weathering, followed by other chemical and microbiological processes. Root etching was the most common biological modification, and the most frequent modifications are involved with the microenvironments of the surface to burial substrate. Potentially cultural modification accounted for only a small portion of the cortical damage to the bones. This general profile indicated an assemblage far more influenced by the immediate environment than by passing animals or people.     

Taphonomy of two last glacial maximum mammoth sites in the central Great Plains of North America: A preliminary report on La Sena and Lovewell

Two mammoth sites from the central Great Plains of North America, each containing one adult Columbian mammoth (Mammuthus columbi), were excavated from Last Glacial Maximum (LGM) loess and fine-grained alluvial deposits, respectively. Taphonomic data from both sites indicate that the mammoth skeletons exhibit numerous spirally fractured limb elements. Dynamic loading points are present on midshafts of large limb bones. Bone flakes produced from the partial thickness of thick cortical bone are also present. Hypotheses of carnivore activity, mammoth trampling, and human-induced fracturing are evaluated as possible causes of the fractured limb bone. Testing the hypotheses using modern data from actualistic taphonomic studies of elephant skeletons, paleontological data from two proboscidean natural death sites, experimental data from elephant bone fracturing, and archaeologically derived data concerning late Pleistocene human modification of mammoth limb bone indicates that the first two hypotheses can be rejected, while the third hypothesis is supported.     

Stable isotopes reveal ecological differences amongst now‐extinct proboscideans from the Cincinnati region,USA

The mobility and dietary preferences of now‐extinct proboscideans have not been comprehensively examined in the central USA. We used stable carbon (δ¹³C), oxygen (δ¹⁸O) and strontium (⁸⁷Sr/⁸⁶Sr) isotopic signatures in molar enamel to investigate the foraging ecology of four mastodons (Mammut americanum) and eight mammoths (Mammuthus spp.) from southwestern Ohio and northwestern Kentucky. We tested two hypotheses: (i) these individuals were nomadic migrants that were passing through the region when they died; and (ii) mammoths and mastodons foraged in different environments. Unexpectedly, our results suggest that 11 of the 12 sampled individuals were regional residents. With the exception of one mastodon, ⁸⁷Sr/⁸⁶Sr ratios for proboscideans and regional water samples were statistically indistinguishable; slightly lower ratios for waters suggest glacial loess has an impact on modern samples. Amongst the individuals identified as residents, ⁸⁷Sr/⁸⁶Sr ratios indicate that mammoths and mastodons foraged in discrete geographical areas, and δ¹³C values imply dietary differences between the genera, which is consistent with our expectations. Oxygen isotope values may be able to distinguish animals that lived during the Last Glacial Maximum (LGM) from those that lived more recently. Three mammoths and one mastodon yielded δ¹⁸O values that are similar to modern regional precipitation and surface water, but too high for estimated drinking water during the LGM. We propose that these individuals lived during a relatively warm period following the LGM. Compellingly, the mammoth with the highest δ¹⁸O value also has the lowest δ¹³C value, suggesting that this individual was alive after regional vegetation shifted from open parkland to deciduous forest dominated by C₃ species. Our results demonstrate that a wealth of information can be gleaned from fossil museum specimens and lay a foundation for future work on the foraging ecology of proboscideans and other extinct megafauna from the Midwest USA.     

The ecological implications of a Yakutian mammoth's last meal

Part of a large male woolly mammoth (Mammuthus primigenius) was preserved in permafrost in northern Yakutia. It was radiocarbon dated to ca. 18,500 ¹⁴C yr BP (ca. 22,500 cal yr BP). Dung from the lower intestine was subjected to a multiproxy array of microscopic, chemical, and molecular techniques to reconstruct the diet, the season of death, and the paleoenvironment. Pollen and plant macro-remains showed that grasses and sedges were the main food, with considerable amounts of dwarf willow twigs and a variety of herbs and mosses. Analyses of 110-bp fragments of the plastid rbcL gene amplified from DNA and of organic compounds supplemented the microscopic identifications. Fruit-bodies of dung-inhabiting Ascomycete fungi which develop after at least one week of exposure to air were found inside the intestine. Therefore the mammoth had eaten dung. It was probably mammoth dung as no bile acids were detected among the fecal biomarkers analysed. The plant assemblage and the presence of the first spring vessels of terminal tree-rings of dwarf willows indicated that the animal died in early spring. The mammoth lived in extensive cold treeless grassland vegetation interspersed with wetter, more productive meadows. The study demonstrated the paleoecological potential of several biochemical analytical techniques.     

Animals and humans in the European Russian Arctic towards the end of the last Ice Age and during the mid‐Holocene time

Here, we present and discuss results from geo‐archaeological and palaeo‐zoological investigations at the Palaeolithic site Pymva Shor, in the Russian Arctic. As many as 3324 vertebrate fauna remains were recovered during two excavations. This includes bones of mammals, birds and fish. Radiocarbon dates were obtained from 26 specimens. The results show ages in the range 30–3 cal. ka BP. Hare and reindeer are the best represented amongst the identified mammalian species, whilst ptarmigan and various wader species dominate the avian bones. The Pleistocene assemblage includes herbivorous herd animals such as horse, bison and musk ox. These species are typical of the treeless tundra‐steppe landscape that existed during the Lateglacial. Of particular interest is a cave lion specimen that has been radiocarbon dated to approximately 15.5 cal. ka BP. According to our knowledge, this is one of the latest dated examples of this species in Eurasia. The faunal composition in the Holocene assemblage is strikingly different and includes distinct forest taxa such as beaver and pine marten. The avifauna also supports a forested environment with the presence of black grouse. A few stone artefacts were found within the strata, and have been radiocarbon dated to 16–15 cal. ka BP, suggesting that there were humans in the Pymva Shore area at that time. We identified impact notches and cut marks on some radiocarbon‐dated reindeer and bison bones, showing that humans were present twice during the Younger Dryas period. A fourth occupation phase is identified during the mid‐Holocene (6–5 cal. ka BP). We also investigated river terraces and obtained a series of luminescence dates. These have been used to reconstruct the geological history and the relationship to the find‐bearing strata.     

Pleistocene raised marine deposits on Wrangel Island, northeast Siberia and implications for the presence of an East Siberian ice sheet

Two previously undocumented Pleistocene marine transgressions on Wrangel Island, northeastern Siberia, question the presence of an East Siberian or Beringian ice sheet during the last glacial maximum (LGM). The Tundrovayan Transgression (459,000–780,000 yr B.P.) is represented by raised marine deposits and landforms 15–41 m asl located up to 18 km inland. The presence of high sea level 64,000–73,000 yr ago (the Krasny Flagian Transgression) is preserved in deposits and landforms 4–7 m asl in the Krasny Flag valley. These deposits and landforms were mapped, dated, and described using amino acid geochronology, radiocarbon, optically stimulated luminescence, electron spin resonance, oxygen isotopes, micropaleontology, paleomagnetism, and grain sizes. The marine deposits are eustatic and not isostatic in origin. All marine deposits on Wrangel Island predate the LGM, indicating that neither Wrangel Island nor the East Siberian or Chukchi Seas experienced extensive glaciation over the last 64,000 yr.