The first record of Ichneumonidae (Insecta: Hymenoptera) from the Upper Cretaceous of Myanmar

A new subfamily of Ichneumonidae, Novichneumoninae subfam. nov., is established based on two new genera with two new species: Novichneumon longus gen. et sp. nov. and Caloichneumon perrarus gen. et sp. nov. These two new species are the first ichneumonids described from the Upper Cretaceous Myanmar (Burmese) amber. A list of all described Mesozoic ichneumonid fossil species with their respective localities and ages is summarized. The distribution of Ichneumonoidea during the Cretaceous indicates that Cretaceous ichneumonids were documented from localities at high latitudes while braconids were distributed worldwide, a pattern consistent with the distribution of extant ichneumonids and braconids.     

Archegocimicidae (Insecta: Heteroptera) from the Purbeck Limestone Group (Lower Cretaceous: Berriasian) of southern England

Four new genera and five new species of Archegocimicidae are described from the Lower Cretaceous of England: namely Mortalia martini gen. et sp. nov., Tyrion lannister gen. et sp. nov., T. cersei sp. nov., Stannis baratheon gen et sp. nov., Daenerys khaleesi gen. et sp. nov. A transitional position of the complex between Asian Late Jurassic and Early Cretaceous entomofaunas is indicated by this new material.     

Two new genera of belytine wasps (Hymenoptera: Diapriidae) from Cretaceous Burmese amber

Christophus myanmarensis gen. et sp. nov., Christophus ohmkuhnlei gen. et sp. nov., and Mintara parva gen. et sp. nov., three new species and two new genera of belytine wasps (Diapriidae: Belytinae) are described and figured from males preserved in mid-Cretaceous Kachin (Burmese) amber. These new fossils enrich the fossil record of Cretaceous diapriids, summarized here, demonstrating the diversity of the family before the K/Pg extinction and suggesting an older origin for the family.     

Early Cretaceous Aleyrodidae (Hemiptera: Sternorrhyncha) from the Lebanese amber

Four new, monotypic genera of Aleyrodidae from the Lower Cretaceous Lebanese amber are described: Shapashe aithiopa gen. sp. nov.; Milqartis azari gen. sp. nov.; Aretsaya therina gen. sp. nov.; and Yamis libanotos gen. sp. nov. All new taxa represent subfamily Aleurodicinae. The key for identification of genera of Aleurodicinae from the Lebanese amber is provided. The morphological features of newly established taxa are discussed. The importance of the fossils for phylogenetic, evolutionary and ecological studies is presented.     

Non-marine gastropods from the Cretaceous–Paleogene transition in the Pingyi Basin,eastern China

Our results present a taxonomic and palaeoecological study on non-marine gastropods from the latest Cretaceous to Paleocene deposits of the Pingyi Basin, Shandong Province, eastern China. These gastropods are systematically described: three species belonging to three genera including Physa dongtaiensis Gu, 1989, Hydrobia datangensis Yü, 1977, and the newly established species Cyathopoma pingyiensis sp. nov.; two indeterminata genera and species including Truncatelloidea gen. et sp. indet., and Pomatiopsidae gen. et sp. indet. Among them, Truncatelloidea gen. et sp. indet. and Cyathopoma pingyiensis sp. nov. are the dominant species with the longest record. Cyathopoma pingyiensis, sp. nov. is the earliest representative of this widely distributed Asian extant genus. Sedimentological facies analysis of the gastropod-bearing beds suggested that Truncatelloidea gen. et sp. indet. lived in a small pond with a river inlet and a shallow lake, while Ph. dongtaiensis, ?Pomatiopsidae gen. et sp. indet, and ?H. datangensis only thrived in the shallow lake. Cyathopoma pingyiensis sp. nov. inhabited the land area around the shallow lake. Our results showed that no significant species change of the gastropod fauna across the K/Pg (Cretaceous/Paleogene) boundary was observed in the Pingyi Basin.     

New lacewings (Insecta: Neuroptera) from the Lower Cretaceous Wealden supergroup of Southern England

Eight new genera and thirteen new species of lacewings (Neuroptera) are described from the Lower Cretaceous Wealden Supergroup, Weald Sub-basin: Principiala rudgwickensis sp. nov. (Ithonidae), Stenomylina medialis gen. et sp. nov., Protosmylina bifasciata gen. et sp. nov. (Osmylidae), Cretapsychops corami gen. et sp. nov., Micropsychops parallelus gen. et sp. nov., Valdipsychops logunovi gen. et sp. nov., V. brigidae gen. et sp. nov., V. maculosus gen. et sp. nov., V. proudlovei gen. et sp. nov., V. minimus gen. et sp. nov., Psychopsites rolandi gen. et sp. nov. (Psychopsidae), Actinophlebioides valdensis gen. et sp. nov. and Protohemerobius perexiguus gen. et sp. nov. (Neuroptera familia incertae sedis). Psychopsidae are shown to dominate the neuropteran fauna, while the presence of Brongniartiellidae has not been confirmed in the Wealden, due to previously thought representatives of this family being in fact psychopsids. This is the first fossil record of Ithonidae and Osmylidae in Britain. The range of Kalligrammatidae has been extended in the Cretaceous of Britain from the mid-Valanginian to early Barremian.     

New fossil mesosciophilids (Diptera: Nematocera) from the Yixian Formation (Lower Cretaceous) of western Liaoning,China

A new genus with two new species, Orentalphila gravia gen. et sp. nov. and O. caloa sp. nov., and three new species in two known genera, Sinosciophila angustia sp. nov., Sinosciophila seboa sp. nov. and Similsciophila undulata sp. nov., are described from the Lower Cretaceous of Yixian Formation at Huangbanjigou, Beipiao City, western Liaoning, China. These new taxa, representing the first records of mesosciophilids from this locality, provide morphological information and diversity for Mesosciophilidae in the Early Cretaceous. An updated key to known genera of Mesosciophilidae is provided. In addition, according the generic diagnosis revised by Zhang, 2007, Mesoplecia antiqua Hao and Ren, 2009 should be transferred to Mesosciophila Rohdendorf, 1946.     

New Early Cretaceous sharks (Chondrichthyes,Elasmobranchii) from deep-water deposits of Austria

Early Cretaceous elasmobranchs still are very insufficiently known despite all progress that has been accomplished in recent years. Here, a small elasmobranch assemblage is presented from the Valanginian of Austria that contributes significantly to a better understanding of early Cretaceous elasmobranch diversity. The new assemblage comprises two new carcharhiniform sharks, Altusmirus triquetrus gen. et sp. nov. and Fornicatus austriacus gen. et sp. nov., a new orectolobiform shark, Similiteroscyllium iniquus gen. et sp. nov., and a galeomorph shark tooth of uncertain affinities. The recent identification of Similiteroscyllium gen. nov. has shown that it has strong similarities with Ornatoscyllium rugasimulatum from the Lower Cretaceous of France. Significant differences of the tooth morphology of O. rugasimulatum and the type specimen O. freemani justify full reconsideration of the systematic position of O. rugasimulatum and require it to be reassigned to Similiteroscyllium gen. nov. described in this paper. The new assemblage described here, and those from the Valanginian of France and Poland comprising 30 additional taxa, indicates that Early Cretaceous elasmobranch diversity was significantly higher than previously assumed. Consequently, the supposed diversity decline of elasmobranchs across the Jurassic-Cretaceous boundary represents a collecting bias rather than a real pattern.     

New pelecinid wasps (Hymenoptera: Pelecinidae) from Upper Cretaceous Myanmar amber

A new genus with a new species, Brachypelecinus euthyntus gen. et sp. nov., and two new species, Abropelecinus tytthus sp. nov. and Zoropelecinus periosus sp. nov., are described and figured from three exquisitely preserved pelecinid wasps in the Upper Cretaceous Myanmar (Burma) amber. These taxa, owing to their well-preserved characters in amber, provide a better understanding of morphological changes and relationships among the constituent groups, while further highlighting the diversity of Pelecinidae in the Mesozoic.     

First record of the beetle family Lasiosynidae (Insecta: Coleoptera) from the Lower Cretaceous of China

A new genus, Cretasyne gen. nov., in the family Lasiosynidae, with two species C. lata sp. nov. and C. longa sp. nov., is described from the Lower Cretaceous Yixian Formation of Inner Mongolia, China. These fossils are the first record of the family from the Yixian Formation and provide significant information on lasiosynid morphology and diversity. The fossil record of Lasiosynidae is also reviewed.     

内蒙古赤峰早白垩世昆虫化石的新发现

本文首次描述了内蒙古自治区赤峰市早白垩世九佛堂组的鞘翅目和异翅目昆虫化石４属５种，其中３新属５新种。这些昆虫化石通过与国内、外相关类群的对比，为该区的地层划分、对比和地质时代的确定，提供了最新的依据。由古生态方面分析，该区为湖沼环境，水体充沛，为古黑龙江流域的一部分。文中里阿龙虱科、沼梭科和中生五节蝽科的材料在国内是首次报道。     

Lacewings (Insecta: Neuroptera) from the Lower Cretaceous Purbeck Limestone Group of southern England

Eight new genera and 12 new species are described from the Lower Cretaceous Purbeck Limestone Group, southern England. Sophogramma wimbledoni sp. nov. (Kalligrammatidae) is described from the Purbeck of Wiltshire. Pterinoblattina peverilensis sp. nov., Pterinoblattina fasciata sp. nov., Purbepsychopsis parallela gen. et sp. nov. (Psychopsidae), Ovalorobius edmondsi gen. et sp. nov. (Prohemerobiidae), Mesosmylidus vulgaris gen. et sp. nov., Osmylochrysa anomala gen. et sp. nov., Osmylochrysa fragilis gen. et sp. nov., Stenochrysa gradata gen. et sp. nov. (Osmylidae), Mesypochrysa minuta sp. nov. (Chrysopidae), Purbemerobius medialis gen. et sp. nov. (?Hemerobiidae), Epimesoberotha parva gen. et sp. nov. (Berothidae) and Pseudocorydasialis alleni (Neuroptera familia incertae sedis) are described from Durlston Bay, Dorset. The species Pterinoblattina penna Scudder, Pterinoblattina pluma (Giebel) (Psychopsidae) Sialium sipylus (Nymphidae) and Osmylopsis duplicata (Osmylidae) are re-examined, described and figured. The genus Valdipsychops Jepson et al. has been synonymized with Pterinoblattina.     

The Albian fern flora of Alexander Island,Antarctica

The Albian Alexander Island macrofossil flora from the Antarctic Peninsula preserves a diverse community of liverworts (Marchantiophyta), ferns (Polypodiopsida), Lycopodiales, Equisetales, Cycadales, Ginkgoales, seed-ferns (Bennettitales and Pentoxylales), Coniferales, and the first representatives of angiospermous leaves in Antarctica. Despite the presence of angiosperms in this assemblage, ferns are the most diverse element of the flora and are also ecologically dominant, while angiosperms contribute a smaller component to floristic diversity and have low abundance. Here we describe 11 fern taxa from this assemblage. The fossils are assigned to Cladophlebis, Sphenopteris and two newly created genera. The new genera and species are described under Adiantitophyllum serratum gen. et. sp. nov. and Nunatakia alexanderensis gen. et. sp. nov., and the new species are recognized as Cladophlebis dissecta sp. nov., Cladophlebis drinnanii sp. nov., Cladophlebis macloughlinii sp. nov. and Sphenopteris sinuosa sp. nov. In total, there are 24 fern species known from Alexander Island. In comparison to older floras (Jurassic) there is a greater diversity of ferns, while latest Cretaceous floras preserve significantly fewer fern species and more angiosperms. Possible factors that might account for such high fern diversity are high rainfall or generally humid conditions, regular disturbances by flooding and occasionally fire, and the preservation of a diverse range of fern communities that represent several palaeoenvironments.     

New Stigmaphronidae and Megaspilidae (Hymenoptera: Ceraphronoidea) from Canadian Cretaceous amber

Three new species within the stigmaphronid genus Tagsmiphron Engel and Grimaldi, 2009, and one new species within the megaspilid genus Conostigmus Dahlbom, 1858 are described from Upper Cretaceous (Campanian) amber originating at the Grassy Lake locality in Alberta, Canada. New taxa include Tagsmiphron spiculum sp. nov., Tagsmiphron leucki sp. nov., Tagsmiphron exitorum sp. nov., and Conostigmus cavannus sp. nov. The new Conostigmus species is a rare discovery. It is the third megaspilid species to be found in Cretaceous amber, with the two specimens described herein effectively doubling the number of known Mesozoic exemplars for the family. We provide the first comprehensive report of known Ceraphronoidea within Canadian amber, and contrast this against other Cretaceous amber assemblages, discussing the potential palaeobiogeographic and palaeoenvironmental implications of the Canadian amber assemblage.     

New Cretaceous Elcanidae from China and Myanmar (Insecta,Orthoptera)

A new species of Elcanidae (Orthoptera: Elcanoidea), Panorpidium yixianensis sp. nov., is described based on two new specimens from the Lower Cretaceous Yixian Formation of northeastern China. It differs from other species in forewing characters and spines on the hind tibiae. In addition, a new specimen Burmelcana sp., is described and figured based on an amber inclusion from the mid-Cretaceous Burmese (Myanmar) amber. P. yixianensis sp. nov. represents the first definite record of Elcanidae in the Early Cretaceous Jehol biota, and Panorpidium is the only genus of Ensifera to be found in the Early Cretaceous faunas of England, Russia and China.     

New fossil helorid wasps (Insecta,Hymenoptera, Proctotrupoidea) from the Jehol Biota,China

A new genus with a new species, Sinohelorus elegans gen. et sp. nov., is described and assigned to the subfamily Mesohelorinae, Heloridae. In addition, two new species, Gurvanhelorus beipiaoensis sp. nov. and Spherogaster saltatrix sp. nov., are also described. The diagnoses of Gurvanhelorus Rasnitsyn, 1986 and Spherogaster Zhang and Zhang, 2001 are emended based on new materials and findings. These well-preserved specimens were collected from the Jehol Biota, Early Cretaceous Yixian Formation of Huangbanjigou Village in Liaoning, China. These new species broaden the diversity of this family and enhance our understanding of evolutionary trend of helorid’s metasomal structure, antenna, and forewing venation from the Middle Jurassic to extant.     

Fossil caddisflies (Insecta: Trichoptera) from the Early Cretaceous of southern England II

The following new fossil trichopteran insects are described or revised from the Purbeck Limestone Group and Wealden Supergroup of southern England: Pteromixanum inviolatum gen. et sp. nov., P. ruderatum gen. et sp. nov., P. purbeckianum (Handlirsch) gen. et comb. nov. and P. poxwellense gen. et sp. nov. (Necrotauliidae);Purbimodus minor gen. et sp. nov., P. medius gen. et sp. nov., P. rasnitsyni gen. et sp. nov. and P. saxosus gen. et sp. nov. (Vitimotauliidae);Palaeoludus popovi gen. et sp. nov. (Dysoneuridae);Palaeotarsus desertus gen. et sp. nov. (Plectrotarsidae);Eucrunoecia ridicula gen. et sp. nov. (Lepidostomatidae);Palaeocentropus placidus gen. et sp. nov. (Calamoceratidae) and Helicophidae gen. et sp. incertae sedis. The general composition of the fauna is compared with Early Cretaceous faunas of Asia. The early appearance of several living families is noted.     

Belemnites in the Jurassic-Cretaceous boundary interval of the Mauryn’ya and Yatriya River sections, Western Siberia: Biostratigraphic significance and dynamics of taxonomic diversity

The stratigraphic distribution and taxonomic diversity of belemnites from the Volgian and Ryazanian deposits exposed in the eastern foothills of the North and Subpolar Urals, are studied and analysed. Studies of new collections from the Mauryn’ya and Yatriya rivers have revealed within the Laugeites groenlandicus-basal Surites analogus ammonite zones, the East Siberian Lagonibelus napaensis, Cylindroteuthis knoxvillensis, and Liobelus russiensis belemnite zones (the latter in the beds’ rank), the Lagonibelus gustomesovi and Arctoteuthis porrectiformis Beds, and local beds with Boreioteuthis explorata and Simobelus compactus. Since in the boundary beds of the Volgian and Ryazanian stages a great number of belemnite species known from the Tordenskjoldberget Member, Kong Karls Land, Svalbard, were found, it is inferred that the accumulation of the member started not in the Valanginian as many researchers believe but immediately at the beginning of the Cretaceous. The dynamics of belemnite species diversity in the northwestern margin of the West Siberian marine basin correlates well with climatic events. An increase of species number in the terminal Volgian-beginning of the Ryazanian corresponds to a temperature elevation in the Siberian paleoseas, and the subsequent reduction of species diversity in the second half of the Ryazanian is correlated with a gradual cooling. The peak of belemnite taxonomic diversity falls at the beginning of the Cretaceous when up to 15 species belonging to eight Cylindroteuthididae genera occurred concurrently. The new species Cylindroteuthis ornata sp. nov., Acroteuthis pseudoconoides sp. nov., Pachyteuthis eximia sp. nov., and Simobelus compactus sp. nov. are described.