A new genus of Tanyderidae (Insecta: Diptera) from Myanmar amber,Upper Cretaceous

A new genus and species, Similinannotanyderus lii gen. et sp. nov., is described from Myanmar amber. It differs from other genera of Tanyderidae based on special characters of wing venation and male genitalia. This new taxon broadens the diversity of Tanyderidae in the Cretaceous, and its morphological characters enhance our understanding of the development and evolution of the primitive crane flies.     

A new genus of alderflies (Megaloptera: Sialidae) in Upper Cretaceous Burmese amber

A new genus and species of Mesozoic alderfly is described as Haplosialodes liui gen. et sp. nov., and from an adult male preserved in Cretaceous Burmese amber. The new genus is closely related to the genera Haplosialis Navás (Recent fauna of Madagascar), Indosialis Lestage (Recent fauna of Southeast Asia), and Eosialis Nel et al. (Eocene of France), suggesting a possible Early Cretaceous age for the clade that comprises these groups.     

Aphrastomedes anthocoroides,a remarkable new cimicomorphan genus and species (Hemiptera: Heteroptera) from Upper Cretaceous Burmese amber

A new genus and species of cimicomorphan bug, Aphrastomedes anthocoroides Yamada and Yamamoto, gen. and sp. nov., is described and illustrated based on two specimens preserved in Upper Cretaceous (Cenomanian) amber from the Hukawng Valley of northern Myanmar (Burma). This remarkable cimicomorphan species is tentatively assigned to the family Velocipedidae primarily on the basis of the structure of the labium, forewings, and genitalia. Aphrastomedes is considered best placed in the Cimiciformes as a member of the Velocipedidae, and it also has a close relationship to the Medocostidae.     

A new genus and species of polypore fungus beetle in Upper Cretaceous Burmese amber (Coleoptera,Tetratomidae, Eustrophinae)

A new polypore fungus beetle is described and figured from an individual preserved in Upper Cretaceous (Cenomanian, ca. 99 Ma) amber from northern Myanmar. Cretosynstrophus archaicus gen. et sp. nov. is confidently placed in the extant subfamily Eustrophinae based on its elongate oval body and pronotum with two basal, sublinear impressions. It shares several characters belonging to two Recent tribes (Eustrophini and Holostrophini), but it cannot be attributed to either of them. Together with other tetratomid genera from the Cretaceous, the new discovery implies that the Recent small family Tetratomidae is much more diverse and more widespread than previously documented. In addition, a morphological similarity between Cretosynstrophus and extant Synstrophus suggests a similar fungi feeding habit for Cretosynstrophus, highlighting an ancient association between tetratomid beetles and fungi in the Mesozoic.     

A new genus and species of soldier beetle from Upper Cretaceous Burmese amber (Coleoptera,Cantharidae, Malthininae)

A new Upper Cretaceous genus and species of soldier beetles, Archaeomalthodes rosetta gen. et sp. nov., is described and illustrated from an individual preserved in Upper Cretaceous (Cenomanian, ca. 99 Ma) amber from northern Myanmar. It is undoubtedly placed in extant subfamily Malthininae based on its small-sized body, somewhat abbreviated elytra and fusiform terminal maxillary palpomere, representing the oldest documented occurrence of Malthininae. It suggests that this subfamily is an ancient group, which originated at least in the earliest Late Cretaceous. Our discovery sheds light on the palaeodiversity of Cantharidae in the Late Mesozoic. Together with other previously reported fossil cantharids, it is likely that Malthininae has been fairly diverse during the early evolution of Cantharidae. On the other hand, a morphological similarity between Archaeomalthodes and Recent malthinines and the occurrence of flowering plants in the Burmese amber implies a potential flower-visiting behaviour of this fossil species.     

The Green Castle Sandstone Formation: A new name for the Cwmffrwd Formation (Old Red Sandstone) preoccupied by an Ordovician Member

The Geological Survey’s Framework Report on the Old Red Sandstone of the Anglo-Welsh region includes a proposal for a new formational name, the Cwmffrwd Formation, for the basal formation of the Daugleddau Group in Carmarthenshire, S. Wales.The name is preoccupied by the Early Ordovician Cwmffrwd Member of the Carmarthen Formation and a new name, the Green Castle Sandstone Formation is proposed for the Cwmffrwd Formation.     

A new genus and species of Praeaulacidae (Hymenoptera: Evanioidea) from Upper Cretaceous Myanmar amber

A new fossil praeaulacid wasp, Habraulacus zhaoi gen. et sp. nov., is described and figured from Upper Cretaceous Myanmar amber. The new genus is established based on the following characters: mesosoma stout, metasoma elongated ellipse with the first metasomal segment very thin and tubular (like a petiole), forewing with pterostigma long and narrow, and both veins Rs+M and 1m-cu very long, forming a long cell 1mcu. Species of Praeaulacidae demonstrate a distinct evolutionary trend, which consists of miniaturization, venational reduction and enlargement of pterostigma, and reduction of number of the antennal segments from the Jurassic to Cretaceous. However, except for the miniaturization, this hardly concerns Habraulacus.     

A new genus of Brochocoleini beetle in Upper Cretaceous Burmese amber (Coleoptera: Archostemata: Ommatidae)

Ommatidae is a small, but ancient, basally nested family of the hyper-diverse beetles (Coleoptera). Here, a remarkable new taxon, Paraodontomma burmitica gen. et sp. nov., is described based on a well-preserved specimen in Upper Cretaceous Burmese amber. Based on a wider epipleural rim on each elytron, the new genus belongs to the extinct tribe Brochocoleini. This finding further reinforces the idea that the Burmese paleofauna of ommatid beetles was much more diverse and abundant than previously thought.     

The occurrence of the genus Tanousia Servain (Mollusca, Gastropoda, Hydrobiidae) in the Middle Pleistocene Piànico-Sèllere Basin (Bergamo, Northern Italy)

The occurrence of the freshwater genus Tanousia Servain 1881 (Hydrobiidae) in the Middle Pleistocene lacustrine succession of the Piànico-Sèllere Basin (Bergamo, Northern Italy) represents a significant biostratigraphical discovery. The Tanousia were recovered from the basal part of the carbonate varved bed (BVC), in siltysandy slump sediments, and from the lower and middle parts of the La Palazzina Member (MLP) of the Piànico formation. The BVC unit, in which a dated tephra layer occurs, accumulated during an interglacial phase, as shown by pollen analyses. The MLP Tanousia-bearing layers have been laid down during cool–temperate climate oscillation. The Piànico population of Tanousia displays great morphological variability but is similar to T. runtoniana (Sandberger, 1880), T. stenostoma (Nordmann, 1901) and T. cf. T. stenostoma (Nordmann, 1901), from the Bavelian and Cromerian interglacial deposits of late Early and early Middle Pleistocene age in Northern Europe. As for these species, the Tanousia population from Piànico appears to indicate fluvial conditions under a fully temperate climate.     

A new genus of Sycoracinae (Diptera: Psychodidae) from Upper Cretaceous amber of New Jersey

We describe herein a new genus and species of Sycoracinae (Xenosycorax engeli). This new taxon is characterized, described, illustrated and its taxonomic position is discussed. This discovery is very interesting for the understanding of the evolution of this group.     

A new genus and species of pygidicranid earwigs from the Upper Cretaceous of southern Asia (Dermaptera: Pygidicranidae)

A third genus and species of fossil pygidicranid earwigs from Burmese amber is described and figured as Stonychopygia leptocerca Engel, Huang, Thomas, and Cai, gen. et sp. nov. (type species of Stonychopygia Engel and Huang, gen. nov.; Stonychopygiinae Engel and Huang, subfam. nov.). Stonychopygia have features similar to the subfamily Echinosomatinae such as the combination of shorter, subequal second and third flagellomeres and femora that are neither compressed or keeled. However, the new fossil species differs from echinosomatines in many details, particularly the slender form, head longer than wide, slender scape, slender and elongate forceps, and absence of stout, short bristles over the integument. The new fossil is compared with its contemporaneous and modern relatives, and comments are provided regarding the classification of Pygidicranoidea, with the genus Haplodiplatys Hincks removed from Diplatyidae to Haplodiplatyidae Engel, fam. nov.     

The oldest tachyporine rove beetle in amber (Coleoptera,Staphylinidae): A new genus and species from Upper Cretaceous Burmese amber

A new genus and species of extinct rove beetle, Procileoporus burmiticus gen. and sp. n., is described from Upper Cretaceous Burmese amber (Cenomanian; ca. 99 Ma). Due to the condition of the fossil, parts of the body could not be examined, but sufficient characteristics were visible to allow the specimen to be placed within the subfamily Tachyporinae (tribe Tachyporini). Procileoporus is most similar to the extant genus Cileoporus Campbell in having a slender abdomen, but it differs in having much shorter abdominal setae, prominent striated microreticulation on the elytra, non-lobed tergite VIII, and in other details of the male genital segments. This is the first tachyporine rove beetle from Burmese amber and the oldest record of the subfamily in amber. The evolution of sexual dimorphism in the genital segments is briefly discussed.     

A new Lyrodiscus (Mollusca, Gastropoda) assemblage from Saint-Acheul (Somme Valley): a reappraisal of MIS 11 malacofaunas from northern France

Recent studies undertaken at Saint-Acheul in the Somme Valley (France) have provided an opportunity to recover malacological assemblages from a tufa deposit located at the top of a Middle Pleistocene fluvial sequence. Molluscan communities are rich and, although dominated by open grassland species, contain up to 23 forest taxa. This high diversity of thermophilous molluscs allows the recognition of fully temperate climatic conditions. Moreover, the occurrence of an extinct zonitid belonging to the genus Retinella (Lyrodiscus), along with several species beyond their modern range (Platyla polita, Ena montana, Ruthenica filograna Clausilia pumila, Clausilia dubia, Macrogastra ventricosa, Perforatella bidentata, Monachoides incarnatus, Belgrandia marginata, Hygromia limbata) allows correlation of the Saint-Acheul assemblage with malacofaunas recovered in other MIS 11 tufa deposits from the Somme and Seine valleys. This age attribution is reinforced at Saint-Acheul by stratigraphy and an ESR date on quartz of 403±73 kyr from the underlying fluvial deposits. In addition, old collections containing R. (Lyrodiscus) specimens have been rediscovered, allowing taxonomic reassessment of the species. This shows that Retinella (Lyrodiscus) skertchlyi Kerney, 1976 is a junior synonym of Retinella (Lyrodiscus) elephantium (Bourguignat, 1869) and that at least two extinct species of R. (Lyrodiscus) occurred in western Europe during the Quaternary. Finally, reappraisal of these French molluscan assemblages shows that they are similar to British malacofaunas of Hoxnian age. These new results strengthen the uniqueness and biostratigraphical value of the 'Lyrodiscus assemblage'.     

A new titanosaur (Dinosauria,Sauropoda) from the Upper Cretaceous of Lo Hueco (Cuenca,Spain)

The upper Campanian-lower Maastrichtian site of Lo Hueco (Cuenca, Spain) has provided a set of well-preserved partial skeletons in anatomical connection or with a low dispersion of their skeletal elements. One partial skeleton is herein described and a new titanosaurian sauropod is established, Lohuecotitan pandafilandi. This titanosaur is diagnosed by eight autapomorphic features: dorsally and ventrally widened or bifurcated posterior centrodiapophyseal lamina in anterior and middle dorsal vertebrae; short postspinal lamina with a transversely expanded distal end represented by smooth scars in the dorsal vertebrae; anteriormost caudals with the medial spinoprezygapophyseal and medial spinopostzygapophyseal laminae ventrally connected with the prespinal and postspinal laminae, respectively; anterior caudal neural spines with a dorsal projection of the prespinal and postspinal laminae; anterior caudal neural spines bears a “greek-cross”-like cross-section; middle caudal centra having two round and rough structures in the dorsal edge of the posterior articulation, which extends to the dorsal surface of the centrum; the articular ends of the rami of the haemal arches are divided in two articular surfaces; and tuberosity between the anterior and the lateral trochanter of the fibula. The herein performed phylogenetic analysis considered L. pandafilandi as a member of Lithostrotia more derived than Malawisaurus. The known palaeodiversity of the Late Cretaceous Ibero-Armorican titanosaurs is increasing, and further analyses focused on this group will be necessary to better understand the evolutionary history of European titanosaurs and to clarify their relationships within Titanosauria.     

Cretoleptochromus archaicus gen. et sp. nov., a new genus of ant-like stone beetles in Upper Cretaceous Burmese amber (Coleoptera,Staphylinidae, Scydmaeninae)

A new genus and species of the ant-like stone beetle supertribe Mastigitae, Cretoleptochromus archaicus gen. et sp. nov., is described and figured based on an exceptionally well-preserved individual in the Upper Cretaceous (lowermost Cenomanian, ca. 99 Ma) amber from Myanmar. Cretoleptochromus is undoubtedly placed in Mastigitae based on its distinctly geniculate antennae with strongly elongate antennal scape, and pointed labial palpomere III. Cretoleptochromus is most likely the sister taxon to the extinct Palaeoleptochromus O'Keefe from the Late Cretaceous Canadian amber. Cretoleptochromus shares with Clidicini the distinct posterior collar of pronotum; with Leptomastacini the presence of mesocoxal lateroventral bristles. The new discovery may help to elucidate the relationships among the Recent three tribes, particularly for Leptomastacini and Clidicini.     

A new species of Platanus from the Cenomanian(Upper Cretaceous) in eastern Heilongjiang,China

The stratigraphic division and sequence of the Upper Cretaceous sediments in eastern Heilongjiang Province,China,have been ambiguous and controversial,mainly due to a lack of biostratigraphically useful fossils and related radiometric dating.A new species of angiospermous fossil plant.Platanus heilongjiangensis sp.nov.,from Qitaihe in eastern Heilongjiang has been found in sediments conformably above which zircons from a rhyolitic tuff has been dated by U-Pb radiometric methods as 96.2± 1.7 Ma.indicating that the Upper Houshigou Formation is of Cenomanian age.This discovery not only provides new data to improve our stratigraphic understanding of the Houshigou Formation,but also shows that Platanus flourished in the early Late Cretaceous floras of the region.This new study also indicates active volcanism taking place in the eastern Heilongjiang region during the Cenomanian of the Late Cretaceous.     

Hebeigramma nom. nov., a new name for Mesogramma Hong,1984 (Caloneurodea) from the Lower Cretaceous of Hebei Province, China

1984年,笔者发表河北早白垩世华脉目(Caloneurodea)的中华脉科、属:MesogrammatidaeHong,1984和Meso-grammaHong,1984。属名与1850年Stephens采用过的鳞翅目Mesogramma属和1865,1872,Loew采用过的双翅目Meso-grapta属是异物同名,必须废除笔者后来建立的科、属学名,重新命名,以维护《法规》中命名优先权的原则。故原MesogrammaHong,1984属名重新命名为Hebeigrammanom.nov.,原MesogrammatidaeHong,1984科名随属名的更改而重新命名为Hebeigrammidaenom.nov.,在此申明。     

First record of a clupeomorph fish in the Neuquén Group (Portezuelo Formation), Upper Cretaceous of Patagonia, Argentina

A new genus and species of clupeomorph fish, Leufuichthys minimus, is described from the fluvial deposits of the Portezuelo Formation, Upper Cretaceous (Turonian-Coniacian) of the Neuquén Group, Patagonia, Argentina. It is a small-sized fish with an estimated body length up to 46 mm. Among other characters, the new species shows the following: abdominal scutes; abdomen moderately convex; anal fin elongate-based; three uroneurals; two epurals; caudal fin bearing very elongate rays; and cycloid scales. Leufuichthys minimus gen. et sp. nov. shows a greater similarity with Kwangoclupea dartevellei, a clupeomorph described from a marine Cenomanian deposit of the Democratic Republic of Congo (Africa), mainly due to the presence of an elongate-based anal fin, bearing more than 20 fin-rays, differing from it by the presence of a not hypertrophied abdomen. As far as known, L. minimus gen. et sp. nov. is the first clupeomorph described in the Upper Cretaceous of Patagonia and represents one fortuitous preservation of an articulated fish in fluvial deposits.     

The first Oriental protorhyssaline wasp (Hymenoptera: Braconidae): A new genus and species in Upper Cretaceous amber from Myanmar

The exclusively Cretaceous braconid wasp subfamily Protorhyssalinae is reported from the Upper Cretaceous deposits of northern Myanmar. Archaeorhyssalus subsolanus Engel, gen. et sp. nov., is represented by a well-preserved female in Burmese amber and is similar to other protorhyssalines, but differs in numerous details of forewing venation and flagellomere number. This species is the first definitive Oriental occurrence for protorhyssalines, expanding their palaeogeographic distribution from the Lower Cretaceous of Spain, and the Upper Cretaceous of France and New Jersey, USA. A key to the known genera of protorhyssalines is provided.     

A new extinct genus of Glandulariini with two species from Upper Cretaceous Burmese amber (Coleoptera: Staphylinidae: Scydmaeninae)

The Glandulariini (=Cyrtoscydmini) is today the largest, most species-rich and most diverse tribe of Scydmaeninae comprising over 70% of the extant species of this subfamily. Named genera and species of Glandulariini are known mostly from Miocene to Eocene ambers, with only one, recently described Mesozoic taxon. Here we report the second genus of Glandulariini from Upper Cretaceous Burmese amber, Cenomaniola Jałoszyński and Yamamoto, gen. nov., with two species, C. carinata Jałoszyński and Yamamoto, sp. nov. and C. macrophthalma Jałoszyński and Yamamoto, sp. nov. Cenomaniola shows the general body plan typical of the ‘Euconnus complex’ within Glandulariini, with thick bristles on the sides of head and pronotum, a character conserved for nearly a hundred million years. With the previous discovery of Scydmobisetia Jałoszyński and Yamamoto, two major body forms typical of the extant Glandulariini are already proved to have differentiated in or before the Late Cretaceous. This demonstrates early origins and a long conservation of ‘Euconnus-like’ and ‘Sciacharis/Horaeomorphus-like’ body plan in the currently largest group among Scydmaeninae.