The last 30,000 years of faunal history within the Grand Canyon, Arizona

The vertebrate fauna of the last 30,000 radiocarbon years in the Grand Canyon is reviewed. Faunas accompanied with 92 ¹⁴C dates have been analyzed from nine cave sites (four systematically excavated) and 50 packrat middens. Reasonably precise chronological and environmental data of late Pleistocene and Holocene age were obtained through dung studies in Rampart, Muav, and Stanton's Caves; from the numerous packrat middens; and from a ringtail refuse deposit in Vulture Cave. The desert tortoise, 8 species of lizards, 12 species of snakes, 68 species of birds, and 33 species of mammals are identified. Extinct animals include the avian carrion feeder, Teratornis merriami, and the mammalian herbivores, Oreamnos harringtoni, Camelops cf. hesternus, Equus sp., and Nothrotheriops shastense. There is no apparent abrupt end to the late Pleistocene as observed in the Grand Canyon fossil faunal or floral record. Animal and plant taxa of the Grand Canyon responded individually to the changes in climate of the last 30,000 yr. Both animal and plant fossil assemblages indicate that a pre-full glacial, a full glacial, and a late glacial woodland community with many less dominant desert taxa were slowly replaced by a Holocene desert community. All woodland taxa were absent from the lower elevations of the Grand Canyon by 8500 yr B.P.     

Terminal Pleistocene change in mammal communities in southeastern Washington State,USA

Small mammal communities in western North America experienced declines in taxonomic richness across the late Pleistocene to Holocene transition (PHT), a recent natural global warming event. One community also experienced a decline in evenness and others replaced one species with a congener. Variability in response of small mammal communities to PHT warming is apparent. At the presently arid and xeric Marmes site in the Columbia Basin of southeastern Washington State, megafauna were absent by about 13,000 cal yr BP, evenness of small mammals declined about 11,700 cal yr BP and again about 11,400 cal yr BP whereas richness declined about 11,400 cal BP. Regional faunal turnover was, however, minimal among small-bodied taxa. Local mammal communities are depauperate as a result of megafaunal extinctions and subsequent decreases in small-mammal richness and evenness. The latter chronologically corresponds with a decrease in primary productivity driven by increasing warmth and aridity. More faunas must be studied in order to fully document the range of variability in the responses of mammalian communities to PHT warming. Documentation of patterns in those responses will facilitate understanding and enhance predictive accuracy with respect to responses of mammalian communities to modern global warming.     

Late Pleistocene paleoecology of arctic ground squirrel (Urocitellus parryii) caches and nests from Interior Alaska's mammoth steppe ecosystem, USA

Botanical analyses of fossil and modern arctic ground squirrel (Urocitellus parryii) caches and nests have been used to reconstruct the past vegetation from some parts of Beringia, but such archives are understudied in Alaska. Five modern and four fossil samples from arctic ground squirrel caches and nests provide information on late Pleistocene vegetation in Eastern Beringia. Modern arctic ground squirrel caches from Alaska's arctic tundra were dominated by willow and grass leaves and grass seeds and bearberries, which were widespread in the local vegetation as confirmed by vegetation surveys. Late Pleistocene caches from Interior Alaska were primarily composed of steppe and dry tundra graminoid and herb seeds. Graminoid cuticle analysis of fossil leaves identified Calamagrostis canadensis, Koeleria sp. and Carex albonigra as being common in the fossil samples. Stable carbon isotopes analysis of these graminoid specimens indicated that plants using the C₃ photosynthetic pathways were present and functioning with medium to high water-use efficiency. Fossil plant taxa and environments from ground squirrel caches in Alaska are similar to other macrofossil assemblages from the Yukon Territory, which supports the existence of a widespread mammoth steppe ecosystem type in Eastern Beringia that persisted throughout much of the late Pleistocene.     

Palaeobotanical evidence for warm summers in the East Siberian Arctic during the last cold stage

Plant macrofossils from the “Mamontovy Khayata” permafrost sequence (71°60′N, 129°25′E) on the Bykovsky Peninsula reflect climate and plant biodiversity in west Beringia during the last cold stage. 70 AMS and 20 conventional ¹⁴C dates suggest sediment accumulation between about 60,000 and 7500 ¹⁴C yr B.P. The plant remains prove that during the last cold-stage arctic species (Minuartia arctica, Draba spp., Kobresia myosuroides) coexisted with aquatic (Potamogeton vaginatus, Callitriche hermaphroditica), littoral (Ranunculus reptans, Rumex maritimus), meadow (Hordeum brevisubulatum, Puccinellia tenuiflora) and steppe taxa (Alyssum obovatum, Silene repens, Koeleria cristata, Linum perenne). The reconstructed vegetation composition is similar to modern vegetation mosaics in central and northeast Yakutian relict steppe areas. Thus, productive meadow and steppe communities played an important role in the Siberian Arctic vegetation during the late Pleistocene and could have served as food resource for large populations of herbivores. The floristic composition reflects an extremely continental, arid climate with winters colder and summers distinctly warmer than at present. Holocene macrofossil assemblages indicate a successive paludification possibly connected with marine transgression, increased oceanic influence and atmospheric humidity. Although some steppe taxa were still present in the early Holocene, they disappeared completely before 2900 ¹⁴C yr B.P.     

Climate variability in the Aral Sea basin (Central Asia) during the late Holocene based on vegetation changes

High-resolution pollen analyses ( 50 yr) from sediment cores retrieved at Chernyshov Bay in the NW Large Aral Sea record shifts in vegetational development from subdesertic to steppe vegetation in the Aral Sea basin during the late Holocene. Using pollen data to quantify climatic parameters, we reconstruct and date for the first time significant changes in moisture conditions in Central Asia during the past 2000 yr. Cold and arid conditions prevailed between ca. AD 0 and 400, AD 900 and 1150, and AD 1500 and 1650 with the extension of xeric vegetation dominated by steppe elements. These intervals are characterized by low winter and summer mean temperatures and low mean annual precipitation (P_mm < 250 mm/yr). Conversely, the most suitable climate conditions occurred between ca. AD 400 and 900, and AD 1150 and 1450, when steppe vegetation was enriched by plants requiring moister conditions (P_mm  250–500 mm/yr) and some trees developed. Our results are fairly consistent with other late Holocene records from the eastern Mediterranean region and the Middle East, showing that regional rainfall in Central Asia is predominantly controlled by the eastern Mediterranean cyclonic system when the North Atlantic Oscillation (NAO) is in a negative phase.     

新疆阿尔泰山大罗坝盆地全新世植被与环境

阿尔泰山中部大罗坝盆地3.2m厚沉积中的孢粉化石可以划分出2个孢粉组合和8个亚组合。晚更新世晚期—早全新世属荒漠草原—草原植被,中晚全新世属森林草原植被。7000a.BP以来,盆地中针叶林一直发育较好。森林发育的最好阶段是中全新世早期,此后曾有两次衰落时期,但森林并未退出本区。从全新世阿尔泰山喀纳斯湖地区与大罗坝盆地的孢粉对比发现,森林的分布从西向东有减少趋势,温度逐渐降低,推测阿尔泰山东部的青河县气候比较干旱。     

Calibrating Late Quaternary terrestrial climate signals: radiometrically dated pollen evidence from the southern Sierra Nevada,USA

We constructed a radiometrically calibrated proxy record of Late Pleistocene and Holocene climate change exceeding 230,000 yr duration, using pollen profiles from two cores taken through age-equivalent dry lakes—one core having greater age control (via ²³⁰Th alpha mass-spectrometry) and the other having greater stratigraphic completeness. The better dated of these two serial pollen records (Searles Lake) served as a reference section for improving the effective radiometric age control in a nearby and more complete pollen record (Owens Lake) because they: (1) are situated ∼90 km apart in the same drainage system (on, and immediately leeward of, the eastern flank of the Sierra Nevada), and (2) preserved strikingly similar pollen profiles and concordant sequences of sedimentological changes. Pollen assemblages from both lakes are well preserved and diverse, and document serial changes in Late Pleistocene and Holocene plant zone distribution and composition in the westernmost Great Basin; they consist of taxa now inhabiting montane forest, woodland, steppe, and desert-scrub environments. The studied core intervals are interpreted here to be the terrestrial equivalent of marine δ¹⁸O stages 1 through 9; these pollen profiles now appear to be among the best radiometrically dated Late Pleistocene records of terrestrial climate change known.     

河套盆地东部第四纪中晚期介形类特征及其沉积环境意义

根据河套盆地东部DTLC孔介形类组合及其岩性、岩相的分析,着重地探讨了该区第四纪中、晚期介形类的生物地层特征及其沉积古环境意义。研究认为:中更新世早期（Q₂1）,未见介形类,以冲洪积沉积环境为主,兼有河湖交替演化;中更新世晚期（Q₂2）,介形类生物组合为Leucocythere plethora-Eucypris inflate-Limnocyther dubiosa,推测为封闭性的半咸水、咸水湖相沉积环境,气候偏冷;晚更新世早期（Q₃1）,介形类生物组合为Leucocythere plethora-Limnocythere dubios,反映了封闭性的半咸水深湖相沉积环境;晚更新世晚期（Q₃2）,少见淡水介形虫化石,以Ilyocypris manasensis-Candoniella albicans-Cypria luminosa为组合特征,反映了以河流冲洪积环境为主;全新世（Q₄）为河流沉积环境。通过河套盆地区域沉积环境的综合分析认为:中更新世晚期（Q₂2）至晚更新世早期（Q₃1）,河套盆地为封闭性内陆湖盆,黄河为内流河;自晚更新世晚期（Q₃2）以来,黄河逐渐地转为外流河,河套盆地为外流湖盆。      

Late Quaternary environmental studies on salt lakes in western Rajasthan,India: a summarised view

The paper reviews pollen analytical and palaeoenvironmental work carried out on saline lakes in western Rajasthan, northwest India. The saline lakes are salient geomorphological features within the arid and semi-arid landscapes to the west of the Aravalli mountain ranges. Preliminary palynological work was carried out on two profiles from a gypsum-rich lake depression around Thob (District of Barmer). The varied pollen data are indicative of a fluctuating vegetational assemblage, possibly in response to local hydrological conditions and not necessarily indicative of climatic change. Pollen of Ephedra sp. (a typical desert species) in the lower levels suggests episodes of relatively dry conditions during the early phase of lake sedimentation at the end of the Pleistocene. Other work at four lake sites, on the basis of pollen analysis has indicated shifts in climatic and vegetational belts during the early Holocene, especially during the period of Indus Valley Culture. Later in the Holocene, between 5000 and 3500 yr BP, rainfall variations have been related to the dynamics of the monsoon. Pioneering geoarchaeological studies carried out at the palaeolithic site of 16 R and the adjacent Didwana lake have illustrated palaeoclimatic fluctuations and accompanying changes in cultural stages from the lower palaeolithic to mesolithic. These studies have been extended geochemically and sedimentologically through a detailed study documenting a history of salinity from 20000 to 13000 yr BP and freshwater conditions from 9000 to 6000 yr BP. Later studies have illustrated a sequence of changes reflecting summer and winter precipitation. These have indicated steppe vegetation during the last Glacial Maximum along with hypersaline lake conditions at Didwana, inferring a weakened summer monsoon and relatively high winter precipitation. The taxa indicative of both summer and winter precipitation in the mid-Holocene declined during the late Holocene, at the same time as falling lake levels around 4000 yr BP, a time when other lakes, at Sambhar, Lunkaransar and Pachpadra, also became ephemeral. © 1997 John Wiley & Sons, Ltd.     

Small mammals from the early Pleistocene of the Granada Basin, southern Spain

The Pliocene and Pleistocene continental sedimentary records of the western sector of the Granada Basin, southern Spain, consist of alternating fluvial and lacustrine/palustrine sediments. Two Quaternary sections from this sector have been sampled: Huétor Tájar and Tojaire. They have yielded remains of rodents, insectivores and lagomorphs. The presence in the Huétor Tájar and Tojaire sections of Mimomys, Apodemus atavus, Castillomys rivas and two different species of Allophaiomys, indicates an Early Pleistocene age. These deposits, which are related to a fluvio-lacustrine system, can be differentiated from an older (Pliocene) braided fluvial system. Their dating has important repercussions on the paleogeographic reconstruction of the basin. The conditions inferred from the ecological preferences of the small mammal associations are wet and cold. These associations suggest a predominance of open herbaceous habitats, followed by forested habitats; semiaquatic habitats are the least represented.     

Late Pleistocene vegetation of Kings Canyon,Sierra Nevada,California

Seven packrat midden samples make possible a comparison between the modern and late Pleistocene vegetation in Kings Canyon on the western side of the southern Sierra Nevada. One modern sample contains macrofossils and pollen derived from the present-day oak-chaparral vegetation. Macrofossils from the six late Pleistocene samples record a mixed coniferous forest dominated by the xerophytic conifers Juniperus occidentalis, Pinus cf. ponderosa, and P. monophylla. The pollen spectra of these Pleistocene middens are dominated by Pinus sp., Taxodiaceae-Cupressaceae-Taxaceae (TCT), and Artemisia sp. Mesophytic conifers are represented by low macrofossil concentrations. Sequoiadendron giganteum is represented by a few pollen grains in the full glacial. Edaphic control and snow dispersal are the most likely causes of these mixed assemblages.The dominant macrofossils record a more xeric plant community than those that now occur on similar substrates at higher elevations or latitudes in the Sierra Nevada. These assemblages suggest that late Wisconsin climates were cold with mean annual precipitation not necessarily greater than modern values. This conclusion supports a model of low summer ablation allowing for the persistence of the glaciers at higher elevations during the late Wisconsin. The records in these middens also suggest that S. giganteum grew at lower elevations along the western side of the range and that P. monophylla was more widely distributed in cismontane California during the Pleistocene.     

Macrofossil and pollen evidence for full-glacial steppe within an ecological mosaic along the Bluefish River,eastern Beringia

Plant and insect macrofossil assemblages dating to the full-glacial (late Wisconsinan) are rare from eastern Beringia. Here we present an assemblage of fossil pollen, insect and plant macrofossils recovered from alluvium at the Bluefish Exposure, northern Yukon Territory. Nine AMS radiocarbon ages place these data between ca. 18,880–16,440 ¹⁴C yr BP (22,313–19,597 cal. yr BP). These data indicate that xeric steppe, rich in bunchgrasses Poa and Elymus, Artemisia frigida and diverse forbs was interspersed within a mosaic of local vegetation types, including mid-rich fens, mesic graminoid meadows, steppe-tundra and herb-tundra. Macrofossils and minor pollen of tundra forbs suggest steppe-tundra plant associations within midslope elevations and discontinuous herb-tundra on high elevation uplands on exposed bedrock ridges. The composition and distribution of local vegetation was dependent on available moisture, drainage, aspect and elevation. Compositional and physiognomic similarities can be made with extrazonal steppe-dominated dry slopes and high elevation steppe-tundra ecotones in central Alaska and Yukon Territory. Our paleoecological data reflect environments inhabited by the diverse late Pleistocene Bluefish Caves fauna, including woolly mammoth, horse, steppe bison, and saiga antelope.     

Paleoenvironmental changes at the northern limit of the subantarctic Nothofagus forest,lat 37°S,Argentina

Paleoenvironmental changes dating back to before 10,000 yr B.P. at the northernmost occurrences of Nothofagus forests in Argentina at about latitude 37°S permit the reconstruction of past changes in the intensity of the winter rains, related to the southern westerlies that appear to determine the forest boundary. The paleoenvironmental interpretation is based on changes in the proportions of different Nothofagus species and changes in the ratio betwen forest and steppe taxa. The relatively most diverse and dense Nothofagus forest developed only during the last 4500 yr, prior to human impact during the last 300 yr. Before the middle Holocene, climatic conditions must have been different from the modern ones, with less overall precipitation judging from the overall lower amount of tree pollen and the reduction to primarily Nothfagus pumilio. An interval dated to older than 10,000 yr B.P. is characterized by co-occurrence of Prumnopitys andina, Nothofagus pumilio, and shrub-steppe taxa. Prumnopitys andina is known today only from scattered upper montane forest sites in Chile between 36° and 43°S lat and its ecological requirements are essentially unknown. The taxa combination, however, suggests that late-glacial climate must have been drier, and probably cooler than today. This implies that the winter rains and, consequently, the seasonal shift of the westerly circulation was reduced during the late Pleistocene and did not reach modern levels before 8500 yr B.P.     

Late Pleistocene and Holocene Afromontane vegetation and headwater wetland dynamics within the Eastern Mau Forest,Kenya

The Mau Forest Complex is Kenya's largest fragment of Afromontane forest, providing critical ecosystem services, and has been subject to intense land use changes since colonial times. It forms the upper catchment of rivers that drain into major drainage networks, thus supporting the livelihoods of millions of Kenyans and providing important wildlife areas. We present the results of a sedimentological and palynological analysis of a Late Pleistocene–Holocene sediment record of Afromontane forest change from Nyabuiyabui wetland in the Eastern Mau Forest, a highland region that has received limited geological characterization and palaeoecological study. Sedimentology, pollen, charcoal, X-ray fluorescence and radiocarbon data record environmental and ecosystem change over the last ~16 000 cal a bp. The pollen record suggests Afromontane forests characterized the end of the Late Pleistocene to the Holocene with dominant taxa changing from Apodytes, Celtis, Dracaena, Hagenia and Podocarpus to Cordia, Croton, Ficus, Juniperus and Olea. The Late Holocene is characterized by a more open Afromontane forest with increased grass and herbaceous cover. Continuous Poaceae, Cyperaceae and Juncaceae vegetation currently cover the wetland and the water level has been decreasing over the recent past. Intensive agroforestry since the 1920s has reduced Afromontane forest cover as introduced taxa have increased (Pinus, Cupressus and Eucalyptus).     

Small-Mammal Data on Early and Middle Holocene Climates and Biotic Communities in the Bonneville Basin, USA

Archaeological investigations in Camels Back Cave, western Utah, recovered a series of small-mammal bone assemblages from stratified deposits dating between ca. 12,000 and 500 ¹⁴C yr B.P. The cave's early Holocene fauna includes a number of species adapted to montane or mesic habitats containing grasses and/or sagebrush (e.g., Lepus townsendii, Marmota flaviventris, Reithrodontomys megalotis, and Brachylagus idahoensis) which suggest that the region was relatively cool and moist until after 8800 ¹⁴C yr B.P. Between ca. 8600 and 8100 ¹⁴C yr B.P. these mammals became locally extinct, taxonomic diversity declined, and there was an increase in species well-adapted to xeric, low-elevation habitats, including ground squirrels, Lepus californicus and Neotoma lepida. The early small-mammal record from Camels Back Cave is similar to the 11,300–6000 ¹⁴C yr B.P. mammalian sequence from Homestead Cave, northwestern Utah, and provides corroborative data on Bonneville Basin paleoenvironments and mammalian responses to middle Holocene desertification.     

Within-taxon morphological diversity in late-Quaternary Neotoma as a paleoenvironmental indicator, Bonneville Basin, Northwestern Utah, USA

Ecological data indicate that as the amount of precipitation in an arid areas increases, so too does mammalian taxonomic richness. This correspondence has been found in two late-Quaternary mammalian faunas from Utah, one from Homestead Cave in the Bonneville Basin. We use the remains of two species of woodrat (Neotoma cinerea and Neotoma lepida) from Homestead Cave to test the hypothesis that as the amount of precipitation in an arid area increases, so too does morphological diversity within individual mammalian taxa. Morphological diversity is measured as corrected coefficients of variation and as richness of size classes of mandibular alveolar lengths. Coefficients of variation for N. cinerea are few and coincide with moisture history if temporally successive small samples are lumped together. More abundant coefficients of variation for N. lepida coincide only loosely with moisture history, likely because such coefficients measure dispersion but not necessarily other aspects of variation. Richness of size classes of N. lepida is high during the early and late Holocene when moisture was high, and lowest during the middle Holocene when climate was most arid.     

The geology and palaeohydrology of the Kow Swamp fossil hominid site,Victoria, Australia

During a lake‐full phase at the end of the Pleistocene Period, lacustrine silts and aeolian sands were deposited around the eastern margins of Kow Swamp. These sediments have yielded the remains of a large population of early Australian man, retaining certain archaic Homo erectus characteristics.

The lake‐lunette system in which the burials were made provides a geographically isolated situation for the examination of late Pleistocene and Holocene hydrological changes which have left their imprint on the sediments, soils and vegetation.     

Extinction of the woolly mammoth (Mammuthus primigenius) and woolly rhinoceros (Coelodonta antiquitatis) in Eurasia: Review of chronological and environmental issues

Kuzmin, Y. V. 2009: Extinction of the woolly mammoth (Mammuthus primigenius) and woolly rhinoceros (Coelodonta antiquitatis) in Eurasia: Review of chronological and environmental issues. Boreas, 10.1111/j.1502‐3885.2009.00122.x. ISSN 0300‐9483. The current evidence for date and environmental preferences of the extinction of two middle–late Pleistocene megafaunal species, the woolly mammoth (Mammuthus primigenius Blum.) and woolly rhinoceros (Coelodonta antiquitatis Blum.), is presented in this review. It is suggested that extinction of these large herbivores in Eurasia was closely related to landscape changes near the Pleistocene–Holocene boundary (c. 12 000–9000 uncalibrated radiocarbon years ago, yr BP), mainly involving the widespread forest formations in the temperate and arctic regions of northern Eurasia and the loss of grasslands crucial to the existence of woolly mammoth and rhinoceros. However, some woolly mammoth populations survived well into the Holocene (up to c. 3700 yr BP), showing that the process of final extinction was fairly complex, with delays in some regions of up to several millennia. The possible role of Palaeolithic humans in the extinction of Late Pleistocene megafauna is also considered.     

Constructing Seasonal Climograph Overlap Envelopes from Holocene Packrat Midden Contents, Dinosaur National Monument, Colorado

Five Neotoma spp. (packrat) middens are analyzed from Sand Canyon Alcove, Dinosaur National Monument, Colorado. Plant remains in middens dated at approximately 9870, 9050, 8460, 3000, and 0 ¹⁴C yr B.P. are used to estimate Holocene seasonal temperature and precipitation values based on modern plant tolerances published by Thompson et al. (1999a, 1999b). Early Holocene vegetation at the alcove shows a transition from a cool/mesic to a warmer, more xeric community between 9050 and 8460 ¹⁴C yr B.P. Picea pungens, Pinus flexilis, and Juniperus communis exhibit an average minimum elevational displacement of 215 m. Picea pungens and Pinus flexilis are no longer found in the monument.Estimates based on modern plant parameters (Thompson et al., 1999a) suggest that average temperatures at 9870 ¹⁴C yr B.P. may have been at least 1° to 3°C colder in January and no greater than 3° to 10°C colder in July than modern at this site. Precipitation during this time may have been at least 2 times modern in January and 2 to 3 times modern in July. Discrepancies in estimated temperature and precipitation tolerances between last occurrence and first occurrence taxa in the midden record suggest that midden assemblages may include persisting relict vegetation.     

Expansion of Magellanic Moorland during the late Pleistocene: Palynological evidence from northern Isla de Chiloé, Chile

The late Quaternary vegetation of northern Isla de Chiloé is inferred from palynological analysis of a section in the Río Negro drainage (42°03′S, 73°50′W). At ca. 30,500 yr B.P., maxima of Astelia and Donatia occurred, suggesting wetland development. From that time until ca. 27,000 yr B.P., steppe indicators such as Compositae/Gramineae dominated, suggesting drier conditions. After 27,000 yr B.P., the moorland shrub Dacrydium gradually increased, reaching a maximum by 18,000 yr B.P. At this time Astelia increased again, suggesting development of cushion bog during cold and wet conditions. The glacial-postglacial transition is characterized by a marked change from peaty sediments to clays, a decrease in the cushion bog flora, and the prevalence of Gramineae/ Compositae and swamp taxa. This vegetation prevailed until ca.7000 yr B.P. when forest taxa became dominant. The floristic pattern inferred from the pollen spectra of the Rio Negro section suggests that the late Pleistocene vegetation of Chiloé resembled modern Magellanic Moorland vegetation (52°–56°lat S). Based on climatic conditions presently associated with Magellanic Moorland, its occurrence in Chiloé at low elevations during the late Pleistocene implies a decrease in average temperature of at least 4°C and an increase in annual precipitation of at least 1500 mm.