The calculation of estuarine turnover times using freshwater fraction and tidal prism models: A critical evaluation

Freshwater fraction and tidal prism models are simple methods for estimating the turnover time of estuarine water. The freshwater fraction method prominently features flushing by freshwater inflow and has sometimes been criticized because it appears not to include flushing by seawater, but this is accounted for implicitly because the average estuary salinity used in the calculation reflects all the processes that bring seawater into the estuary, including gravitational circulation and tidal processes. The model relies on measurable salinity differences among water masses and so must be used for estuaries with substantial freshwater inflow. Tidal prism models are based on flushing by flood tide inflow and ignore seawater inflow due to gravitational circulation. These models should only be applied to estuaries with weak or nonexistent gravitational circulation, which are generally those with little freshwater inflow. Using a framework that is less ambioguous and more directly applicable to the estimation of turnover times than those used previously, this paper critically examines the application of tidal prism models in well-mixed estuaries with complete tidal exchange, partial ebb return, or incomplete flood mixing and in partially mixed estuaries. Problems with self-consistency in earlier versions of these models also apply to the budgeting procedure used by the LOICZ (Land-Ocean Interactions in the Coastal Zone) program. Although freshwater fraction and tidal prism models are different approaches to estimating turnover times in systems with very different characteristics, consistent derivation shows that these models have much in common with each other and that they yield equivalent values that can be used to make comparisons across systems.     

A percent-of-flow approach for managing reductions of freshwater inflows from unimpounded rivers to Southwest Florida estuaries

The Southwest Florida Water Management District has implemented a management approach for unimpounded rivers that limits withdrawals to a percentage of streamflow at the time of withdrawal. The natural flow regime of the contributing river is considered to be the baseline for assessing the effects of withdrawals. Development of the percent-of-flow approach has emphasized the interaction of freshwater inflow with the overlap of stationary and dynamic habitat components in tidal river zones of larger estuarine systems. Since the responses of key estuarine characteristics (e.g., isohaline locations, residence times) to freshwater inflow are frequently nonlinear, the approach is designed to prevent impacts to estuarine resources during sensitive low-inflow periods and to allow water supplies to become gradually more uvailable as inflow increases. A high sensitivity to variation at low inflow extends to many invertebrates and fishes that move upstream and downstream in synchrony with inflow. Total numbers of estuarine-resident and estuarine-dependent organisms have been found to decrease during low-inflow periods, including mysids, grass shrimp, and juveniles of the bay anchovy and sand seatrout. The interaction of freshwater inflow with seasonal processes, such as phytoplankton production and the recruitment of fishes to the tidal-river nursery, indicates that withdrawal percentages during the springtime should be most restrictive. Ongoing efforts are oriented toward refining percentage withdrawal limits among seasons and flow ranges to account for shifts in the responsiveness of estuarine processes to reductions in freshwater inflow.     

A conceptual model of estuarine freshwater inflow management

As humans continue to influence the quantity, timing, and quality of freshwater input to estuaries, it is becoming increasingly common for policies to be enacted that mandate the establishment of freshwater inflow criteria that will serve to preserve and protect estuarine ecosystems. This paper reviews the scientific literature describing how changes in freshwater inflow affect estuaries, proposes a conceptual model that explores the roles of scientists, citizens, politicians, and managers in the management of freshwater inflow to estuaries, and uses the model to explore the ways in which freshwater inflow is managed in a variety of estuaries. The scientific review is organized to provide an overview of the connections between freshwater inflow (in terms of the quantity, quality, and timing of water delivery), estuarine conditions (such as salinity and concentrations of dissolved and particulate material), and estuarine resources (such as the distribution and abundance of organisms), and to highlight our understanding of the causative mechanisms that underlie the relationships among these variables. The premise of the conceptual model is that the goal of estuarine freshwater inflow policy is to protect those resources and functions that we as a society value in estuaries, and that management measures use scientific information about the relationships among inflow, conditions, and resources to establish inflow standards that can meet this goal. The management approach can be inflow-based (flow is kept within some prescribed bounds under the assumption that taking too much away is bad for the resources), condition-based (inflow standards are set in order to maintain specified conditions in the estuary), or resource-based (inflow standards are set based on the requirements of specific resources), but each of these is carried out by regulating inflow. This model is used as a framework to describe the development of freshwater inflow criteria for estuaries in Texas, Florida, and California.     

A resource-based framework for establishing freshwater inflow requirements for the Suwannee River estuary

The availability of methods for establishing freshwater inflow requirements for estuaries lags behind those for establishing flow requirements in riverine ecosystems. Some of the basic principles and approaches for establishing riverine flow requirements may be applicable to estuaries. An emerging approach for establishing freshwater inflow needs for the Suwannee River estuary involves maintaining a natural inflow regime (in terms of magnitude, frequency, duration, and timing of freshwater flows) and identifying important habitat targets to be protected. The salinity-river flow conditions needed to sustain the habitat targets in their existing condition are then identified. A variety of tools are employed, such as salinity metrics, biological metrics, limits of distribution of communities or habitats, and landscape-scale characteristics to define the salinity and corresponding flow ranges needed to protect and maintain the resource targets. With this information, combined with use of models to evaluate flow-salinity relationships and various withdrawal scenarios, river flow criteria can be set which address the freshwater inflow requirements to maintain these ranges. Subsequentmonitoring and research is undertaken to evaluate the effectiveness of the river flow criteria in protecting the estuarine resource targets. This information can be used to subsequently confirm, refine, or modity the flow criteria.     

Of manatees, mangroves, and the Mississippi River: Is there an estuarine signature for the Gulf of Mexico?

Important parameters of estuarine variability include morphology, flushing times, nutrient loading rates, and wetland: water ratios. This variability both reflects and disguises underlying relationships between the physics and biology of estuaries, which this comparative analysis seeks to reveal, using the Gulf of Mexico (GOM) estuaries as a starting point. A question used to focus this analysis is: are the GOM estuaries unique? The GOM receives the Mississippi River, a uniquely large, world-class river, which dominates the freshwater and nutrient inflows to the GOM continental shelf, whose margins include 35 major estuarine systems. These GOM estuaries have 28% and 41% of the U.S. estuarine wetlands and open water, respectively. Within the GOM, estuarine nitrogen, phosphorus, and suspended matter loading varies over 2 orders of magnitude. Anoxic estuarine events tend to occur in estuaries with relatively slow freshwater turnover and high nitrogen loading. Compared to estuaries from other regions in the U.S., the average GOM estuary is distinguished by shallower depths, faster freshwater flushing time, a higher wetland area:open water area ratio, greater fisheries yield per area wetland, lower tidal range, and higher sediment accumulation rates. The average GOM estuary often, but not always, has a flora and fauna not usually found in most other U.S. estuaries (e.g., manatees and mangroves). Coastal wetland loss in the GOM is extraordinarily high compared to other regions and is causally linked to cultural influences. Variations in nutrient loading and population density are very large among and within estuarine regions. This variation is large enough to demonstrate that there are insufficient systematic differences among these estuarine regions that precludes cross-system analyses. There are no abrupt discontinuities among regions in the fisheries yields per wetland area, tidal amplitude and vegetation range, salt marsh vertical accretion rates and organic accumulations, nitrogen retention, or wetland restoration rates. These results suggest that a comparative analysis emphasizing forcing functions, rather than geographic uniqueness, will lead to significant progress in understanding how all estuaries function, are perturbed, and even how they can be restored.     

Habitat partitioning of fishes in an urban,estuarine Bayou

The physiochemical variability inherent to estuaries makes it difficult to identify those variables of importance to habitat selection by fishes. We used factor analysis to investigate habitat partitioning of fishes in Contraband Bayou, an oligohaline creek within the City of Lake Charles, in southwestern Louisiana. This bayou receives runoff and organic wastes from the City of Lake Charles and adjacent farmland (adding levels of complexity to an analysis of fish habitat preferences). Fishes were sampled from locations within the Contraband drainage in winter and summer of 1983 and 1984. At each location, 14 environmental variables were measured. Distribution of each common species in a season relative to these variables was characterized by calculating means of individuals for each environmental variable. A species' mean for a variable (the state of the variable where individuals of the species are most likely to be found) was interpreted as the species' “preference” for the variable. Factor analysis was used to identify trends in habitat partitioning among species, based on their preferences for all environmental variables. In all seasons, habitat partitioning separated typically freshwater forms from estuarine fishes, and separated forms more tolerant of organic pollution from less tolerant species. Some species entered the bayou only in summer, and in this season, additional factors were related to the preference of some of these species for areas with little cover that were normally shunned by resident species.     

The effect of freshwater inflow on meiofaunal and macrofaunal populations in the Guadalupe and Nueces Estuaries,Texas

Two estuaries with very different inflow characteristics were compared to test the hypothesis that benthic standing crops are enhanced by freshwater inflow. Assuming predation pressure is similar in both estuaries, this would imply that freshwater inflow enhances secondary production. The Guadalupe Estuary had 79 times more freshwater inflow than the Nueces Estuary, and a third of the salinity. The Guadalupe had higher macrofaunal densities and biomass than the Nueces, and both parameters increased with decreasing salinity within the Guadalupe Estuary. Macrofauna density increased with increasing salinity in the Nueces Estuary, due to invasion by marine species. However, meiofauna population size responds differently than macrofauna. Meiofaunal densities were higher in the low-inflow Nueces Estuary, and increased with increasing salinity in both estuaries. Macrofauna diversity increased with salinity, both within and between estuaries. The macrofauna response supports the hypothesis that increased freshwater inflow stimulates secondary production. A review of past benthic studies in these estuaries and the historical climatic patterns indicate that wet years with high inflow result in increased macrofaunal productivity. Since, macrofaunal diversity decreased with lower salinity both within and between the estuaries, the enhanced productivity is due to increases by freshwater and estuarine species that can tolerate low salinities. Increased macrofaunal densities are associated with decreasing meiofaunal densities. The latter result could be due to either increased macrofaunal competition with or predation on meiofauna, or a lack of low-salinity tolerance by meiofauna.     

Freshwater inflow and estuarine variability in the Gamtoos Estuary, South Africa

The Gamtoos is a shallow flood-tidal estuary located on the south coast of South Africa. Even though it has an extensive catchment area, dams limit runoff and mean freshwater inflow is estimated at less than 1 m³ s^?1, and the flood tidal deltas constrict and at times even close the mouth. The results presented here derive from an intensive measurement program carried out over a 3-wk period at the end of 1992, immediately after good rains in the Gamtoos catchment region. Freshwater inflow increased to more than 10 m³ s^?1, driving the salt wedge downstream and resulting in intense haloclines in the mid-estuary region. The program monitored the return to more average estuarine structures, and even though tidal exchange was restricted, marked differences occurred in stratification at neap and spring tides; tidal exchanges provided the dominant mixing forces. It is found that the shallower upper reaches of the estuary are flushed with relatively small increases in freshwater inflow, though a balance exists with the tidal exchanges through the constricted mouth. The variation in the position of the salt wedge and in the salinity stratification can have substantial implications for biota.     

Testate amoebae as estuarine water‐level indicators: modern distribution and the development of a transfer function from a freshwater tidal marsh (Scheldt estuary,Belgium)

Little is known about the century‐scale response of water levels in inland estuaries to sea‐level change and human modifications to estuarine morphology. This study explored the ability of using testate amoebae (Protozoa, Rhizopoda) from sediments of a freshwater tidal marsh as indicators of water level in an inland estuary. The hypothesis was that modern testate amoeba assemblages change with surface elevation (approximately the duration of tidal flooding) within a freshwater tidal marsh. Variation in testate amoeba assemblages in relation to multiple environmental variables and sediment characteristics was studied through redundancy analysis. This demonstrated that a significant part of the variation in modern testate amoeba assemblages could be explained by flooding frequency, surface elevation, organic content and particle size of the soil. Transfer functions, partial least squares and weighted average regressions were made to show that testate amoebae can be used for reconstruction of water level (with an accuracy of 0.05 Normalized Elevation). A preliminary test of application of the transfer function to palaeo testate amoeba assemblages showed promising results. Testate amoebae from a freshwater tidal marsh provide a potentially powerful new tool for estuarine water‐level reconstructions. Copyright © 2011 John Wiley & Sons, Ltd.     

Spatial and temporal variability and drivers of net ecosystem metabolism in western Gulf of Mexico estuaries

Net ecosystem metabolism (NEM) is becoming a commonly used ecological indicator of estuarine ecosystem metabolic rates. Estuarine ecosystem processes are spatially and temporally variable, but the corresponding variability in NEM has not been properly assessed. Spatial and temporal variability in NEM was assessed in four western Gulf of Mexico shallow water estuaries. NEM was calculated from high-frequency dissolved oxygen measurements. Interbay, intrabay, and water column spatial scales were assessed for NEM, gross primary production (GPP), and respiration (R) rate variability. Seasonal, monthly, and daily temporal scales in NEM, GPP, and R were also assessed. Environmental conditions were then compared to NEM to determine which factors were correlated with each temporal and spatial scale. There was significant NEM spatial variability on interbay, intrabay, and water column spatial scales. Significant spatial variability was ephemeral, so it was difficult to ascertain which environmental conditions were most influential at each spatial scale. Significant temporal variability in NEM on seasonal, monthly, and daily scales was found and it was correlated to temperature, salinity, and freshwater inflow, respectively. NEM correlated strongly with dissolved oxygen, temperature, and salinity, but the relationships where different in each bay. The dynamics of NEM on daily scales indicate that freshwater inflow events may be the main driver of NEM in the semiarid estuaries studied. The variable nature of NEM found here is further evidence that it is not valid to use single station monitoring deployments for assessment of whole estuarine ecosystem metabolic rates in large ecosystems. The relationship between NEM and temperature, salinity, and freshwater inflow events could drive predictive models assessing the potential influence of projected climate change and watershed development scenarios on estuarine metabolic rates.     

Estuarine Habitat and Demographic Factors Affect Juvenile Chinook (<Emphasis Type="Italic">Oncorhynchus tshawytscha</Emphasis>) Growth Variability in a Large Freshwater Tidal Estuary

Estuarine rearing has been shown to enhance within watershed biocomplexity and support growth and survival for juvenile salmon (Oncorhynchus sp.). However, less is known about how growth varies across different types of wetland habitats and what explains this variability in growth. We focused on the estuarine habitat use of Columbia River Chinook salmon (Oncorhynchus tshawytscha), which are listed under the Endangered Species Act. We employed a generalized linear model (GLM) to test three hypotheses: (1) juvenile Chinook growth was best explained by temporal factors, (2) habitat, or (3) demographic characteristics, such as stock of origin. This study examined estuarine growth rate, incorporating otolith microstructure, individual assignment to stock of origin, GIS habitat mapping, and diet composition along ~130 km of the upper Columbia River estuary. Juvenile Chinook grew on average 0.23 mm/day in the freshwater tidal estuary. When compared to other studies in the basin our growth estimates from the freshwater tidal estuary were similar to estimates in the brackish estuary, but ~4 times slower than those in the plume and upstream reservoirs. However, previous survival studies elucidated a possible tradeoff between growth and survival in the Columbia River basin. Our GLM analysis found that variation in growth was best explained by habitat and an interaction between fork length and month of capture. Juvenile Chinook salmon captured in backwater channel habitats and later in the summer (mid-summer and late summer/fall subyearlings) grew faster than salmon from other habitats and time periods. These findings present a unique example of the complexity of understanding the influences of the many processes that generate variation in growth rate for juvenile anadromous fish inhabiting estuaries.     

Is the Response of Estuarine Nekton to Freshwater Flow in the San Francisco Estuary Explained by Variation in Habitat Volume?

Abundance of estuarine biota can vary with freshwater inflow through several mechanisms. One proposed mechanism is that the extent of physical habitat for an estuarine species increases with flow. We estimated the contribution of variation in habitat volume to the responses of eight species of estuarine nekton to changes in freshwater flow in the San Francisco Estuary. Resource selection functions for salinity and depth were developed for each species (and for five additional species) using five monitoring data sets. The TRIM3D hydrodynamic model was run for five steady flow scenarios to determine volume by salinity and depth, and resource selection functions were used as a weighting factor to calculate an index of total habitat for each species at each flow. The slopes of these habitat indices vs. flow were consistent with slopes of abundance vs. flow for only two of the species examined. Therefore, other mechanisms must underlie responses of abundance to flow for most species.     

Fish-Habitat Relationships Along the Estuarine Gradient of the Sacramento-San Joaquin Delta,California: Implications for Habitat Restoration

Estuaries are highly variable environments where fish are subjected to a diverse suite of habitat features (e.g., water quality gradients, physical structure) that filter local assemblages from a broader, regional species pool. Tidal, climatological, and oceanographic phenomena drive water quality gradients and, ultimately, expose individuals to other habitat features (e.g., stationary physical or biological elements, such as bathymetry or vegetation). Relationships between fish abundances, water quality gradients, and other habitat features in the Sacramento-San Joaquin Delta were examined as a case example to learn how habitat features serve as filters to structure local assemblages in large river-dominated estuaries. Fish communities were sampled in four tidal lakes along the estuarine gradient during summer-fall 2010 and 2011 and relationships with habitat features explored using ordination and generalized linear mixed models (GLMMs). Based on ordination results, landscape-level gradients in salinity, turbidity, and elevation were associated with distinct fish assemblages among tidal lakes. Native fishes were associated with increased salinity and turbidity, and decreased elevation. Within tidal lakes, GLMM results demonstrated that submersed aquatic vegetation density was the dominant driver of individual fish species densities. Both native and non-native species were associated with submersed aquatic vegetation, although native and non-native fish populations only minimally overlapped. These results help to provide a framework for predicting fish species assemblages in novel or changing habitats as they indicate that species assemblages are driven by a combination of location within the estuarine gradient and site-specific habitat features.     

Suspended-Sediment Flux and Retention in a Backwater Tidal Slough Complex near the Landward Boundary of an Estuary

Backwater tidal sloughs are commonly found at the landward boundary of estuaries. The Cache Slough complex is a backwater tidal region within the Upper Sacramento–San Joaquin Delta that includes two features that are relevant for resource managers: (1) relatively high abundance of the endangered fish, delta smelt (Hypomesus transpacificus), which prefers turbid water and (2) a recently flooded shallow island, Liberty Island, that is a prototype for habitat restoration. We characterized the turbidity around Liberty Island by measuring suspended-sediment flux at four locations from July 2008 through December 2010. An estuarine turbidity maximum in the backwater Cache Slough complex is created by tidal asymmetry, a limited tidal excursion, and wind-wave resuspension. During the study, there was a net export of sediment, though sediment accumulates within the region from landward tidal transport during the dry season. Sediment is continually resuspended by both wind waves and flood tide currents. The suspended-sediment mass oscillates within the region until winter freshwater flow pulses flush it seaward. The hydrodynamic characteristics within the backwater region such as low freshwater flow during the dry season, flood tide dominance, and a limited tidal excursion favor sediment retention.     

Nekton community change along estuarine salinity gradients: Can salinity zones be defined?

Organisms tend to inhabit predictable portions of estuaries along salinity gradients between the ocean inlets (salinity > 35 psu) and the freshwater tributaries (salinity = 0). Previous studies have suggested that the continuous change in biological community structure along this gradient is relatively rapid at certain salinities. This is the basis for estuarine salinity zonation schemes similar to the classic Venice System (i.e., 0–0.5, 0.5–5, 5–18, 18–30, 30–40, > 40). An extensive database (n > 16,000 samples) of frequency of occurrence of nekton was used to assess evidence for estuarine salinity zones in two southwest Florida estuaries: Tampa Bay and Charlotte Harbor. Rapid change in nekton community structure occurred at each end of the estuarine salinity gradient, with comparatively slow (but steady) change in between. There was little strong evidence for estuarine salinity zones at anything other than low salinities (0.1–1). As previously suggested by other authors, estuaries may be regarded as ecoclines, because they form areas of relatively slow but progressive ecological change. The ends of the estuarine salinity gradient appear to be ecotones (areas of rapid change) at the interfaces with adjacent freshwater and marine habitats. This study highlights the rapid change that occurs in nekton community structure at low salinities, which is of relevance to those managing freshwater inflow to estuaries.     

Ecological Signatures of Anthropogenically Altered Tidal Exchange in Estuarine Ecosystems

One of the most conspicuous anthropogenic disturbances to estuaries worldwide has been the alteration of freshwater and tidal influence through the construction of water control structures (dikes, tide gates, culverts). Few studies have rigorously compared the responses of differing groups of organisms that serve as contrasting conservation targets to such anthropogenic disturbances in estuarine ecosystems. Elkhorn Slough in central California includes a spectrum of tidally restricted habitats behind water control structures and habitats experiencing full tidal exchange. To assess community composition for several different taxa in habitats with varying tidal exchange, we employed a variety of field approaches and synthesized results from several different studies. Overall, we found that communities at sites with moderately restricted tidal exchange were fairly similar to those with full tidal exchange, but those with extremely restricted tidal exchange were markedly different from other categories. These differences in community composition are likely the result of several factors, including restricted movement due to physical barriers, differences in water quality characteristics, and differences in habitat structure. Indeed, in this study, we found that water quality characteristics strongly vary with tidal restriction and may strongly influence patterns of species presence or absence. We also found that different conservation targets showed contrasting responses to variation in tidal exchange. Full exchange appears to favor native oysters, commercially valuable flatfish, migratory shorebirds, and site-level biodiversity. Minimal tidal exchange due to water control structures supports a suite of estuarine endemics (including the tidewater goby and California brackish snail) not represented elsewhere and minimizes invasions by non-native marine species. Altogether, our results suggest that total estuary-wide biodiversity may be enhanced with a mosaic of tidal exchange regimes.     

ENSO impacts on salinity in Tampa Bay, Florida

Estuarine salinity distributions reflect a dynamic balance between the processes that control estuarine circulation. At seasonal and longer time scales, freshwater inputs into estuaries represent the primary control on salinity distribution and estuarine circulation. El Niño-Southern Oscillation (ENSO) conditions influence seasonal rainfall and stream discharge patterns in the Tampa Bay, Florida region. The resulting variability in freshwater input to Tampa Bay influences its seasonal salinity distribution. During El Niño events, ENSO sea surface temperature anomalies (SSTAs) are significantly and inversely correlated with salinity in the bay during winter and spring. These patterns reflect the elevated rainfall over the drainage basin and the resulting elevated stream discharge and runoff, which depress salinity levels. Spatially, the correlations are strongest at the head of the bay, especially in bay sections with long residence times. During La Niña conditions, significant inverse correlations between ENSO SSTAs and salinity occur during spring. Dry conditions and depressed stream discharge characterize La Niña winters and springs, and the higher salinity levels during La Niña springs reflect the lower freshwater input levels.     

Origin and significance of mud-filled incised valleys (Upper Cretaceous) in southern Alberta, Canada

Seven mud-filled incised valleys (MFIVs) in the paralic facies of the Dinosaur Park and Horseshoe Canyon formations (Upper Cretaceous) of southern Alberta were studied to better understand their morphology, geometry and depositional histories in an estuarine context. Two preservational geometries occur: simple, U-shaped forms; and internally complex forms. Both types of MFIV record deposition in the central zone of low energy (turbidity) in an estuarine setting. Simple, U-shaped MFIVs have sharp basal erosional surfaces and consist of mudstone-dominated heterolithic fills of channel-wide, concave-up laminae. Associated fossil assemblages are marine to brackish. Each simple MFIV records a cut-and-fill history associated with a cycle of relative sea-level drop and rise. Low-energy depositional settings, loss of channel form during infilling, and associated shoreface deposits, as well as the absence of clear tidal indicators suggest a coastal plain estuarine setting, along a wave-dominated, barred coastline. Complex MFIVs are rarer, and consist of imbricated, wedge-shaped sets of inclined-to-horizontal heterolithic strata. Tidal deposits and/or nonmarine-to-marine macrofossils occur locally. Complex MFIVs were infilled in meandering reaches of the central zone of low energy in tide-dominated estuaries. Their rarity compared to simple MFIVs and their freshwater palaeontological content suggest that they were contiguous landward with extensive fluvial channels. A complex MFIV near Onefour comprises three in-channel depositional cycles. Each cycle consists of an erosional surface overlain by lateral accretion bedding and a conformable transition to vertically aggraded strata. Each cycle reflects a cut-and-fill event under the control of changes in relative sea-level that culminated in overbank flooding. All MFIVs formed in low-gradient settings (≤0.03%) where estuarine zones were stretched out over many tens of kilometres. Tide-dominated estuaries apparently exhibited simple, straight-to-meandering upstream transitions and extensive landward penetration (≥200 km) of tidal backwater effects. Few modern estuaries serve as adequate modern analogues to these ancient, tide-dominated estuaries. Radiometric data indicate that MFIV cut-and-fill cycles were 100 000-400 000 years in maximum duration and thus, equivalent to 4th order sea-level cycles. However, negative evidence tentatively suggests that these cycles took place over time intervals 1-2 orders of magnitude smaller (5th order or higher sea-level cycles).     

Tidal intrusion fronts

A tidal intrusion front forms as a dense seawater inflow plunges (subducts) beneath ambient estuarine water during flood tide. The associated foam lines and color changes have been observed on many smaller estuaries with constricted mouths. Internal hydraulic theory and laboratory experiments are reviewed and expressions are obtained for the position of plunging and the amount of associated mixing. The existence of a tidal intrusion front and its structure are discussed in terms of densimetric Froude numbers. These fronts are particularly important in smaller estuaries in which the intrusion process may dominate wind and tidal mixing and thus determine the overall stratification of the estuary. Three classes of three-dimensional plunging flow are identified and discussed. In particular, it is suggested that the peculiar, cursive V-shape plunge line is characteristic of strongly plunging flow.     

Comparison of Estuarine Salinity Gradients and Associated Nekton Community Change in the Lower St. Johns River Estuary

Salinity is an important determinant of estuarine faunal composition; previous studies, however, have indicated conflicting accounts of continuous vs. relatively rapid change in community structure at certain salinities from geographically distinct estuaries. This study uses a large fisheries monitoring database (n?>?5,000 samples) to explore evidence for estuarine salinity zonation by nekton in the lower St. Johns River estuary (LSJR). There was little evidence to support the presence of estuarine salinity zones except at the extremes of the salinity gradient (i.e., 0.1?C1.0 and 34?C39). The LSJR estuarine nekton community exhibits progressively slow ecological change throughout most of the salinity gradient with rapid change at the interfaces with fresh and marine waters??an ecoline bounded by ecotones. This study affirms the rapid change that occurs at the extremes of the salinity spectrum in certain estuaries and is relevant to efforts to manage surface water resources and estuarine ecosystems. Given the disparity in the results of the studies examining biological salinity zones in estuaries, it would be wise to have, at minimum, a regional understanding of how communities are structured along the gradient from freshwater to marine.