基于灰色系统西南大西洋阿根廷滑柔鱼资源丰度预测模型的构建

阿根廷滑柔鱼（Illex argentinus）是西南大西洋鱿钓渔业的主要作业鱼种,对资源丰度进行准确的预测可指导企业合理安排渔业生产。因此,本研究根据2000-2016年我国西南大西洋阿根廷滑柔鱼的生产数据,以单位捕捞努力量的渔获量（Catch per unit effort, CPUE）为阿根廷滑柔鱼资源丰度的指标,利用灰色绝对关联分析和灰色预测建模的方法（GM（0, N））,计算2001-2015年CPUE的时间序列值与产卵期（6-8月）产卵场海表面温度（Sea surface temperature, SST）时间序列值的灰色绝对关联度,选取产卵场海域中灰色绝对关联度大于0.90的海区SST建立资源丰度预测模型,并用2016年实际CPUE进行验证。灰色绝对关联分析表明,6-8月,30°~40°S,45°~60°W海域内存在若干海区的SST与次年对数CPUE时间序列呈现较强的关联度,可作为预报因子。GM（0, N）模型结果表明,以6-8月产卵场SST作为环境因子建立的模型4能较好地拟合出阿根廷滑柔鱼资源丰度变动趋势,与2016年真实值相比,相对误差为7%,该模型可较好地作为阿根廷滑柔鱼资源丰度的预测模型。相反,包含6月和7月SST的模型1效果优于不包含6月SST的模型2或不包含7月SST的模型3,拟合得到的2016年的数据与真实值相比,相对误差分别为128%和289%,这说明6月和7月是西南大西洋阿根廷滑柔鱼的主要产卵月份。     

The impact of environmental gradients on the early life inshore migration of the short-finned squid Illex illecebrosus

Recruitment of the short-finned squid Illex illecebrosus to adult feeding grounds on the shelf off eastern Canada constitutes an important transition from warm food-limited Gulf Stream waters to cold and productive slope and coastal waters. The impact of such gradients was addressed by analysing the gladius growth of 1585 juvenile squid collected across the Gulf Stream and shelf/slope fronts during research cruises conducted between 1979 and 1989. Temperature- and size-specific growth potential, as estimated by a bioenergetics model, were compared to measured gladius growth rates and revealed that young Illex were energetically expensive and food-limited in Gulf Stream waters (their hatching environment). Growth condition improved inshore, where metabolic costs decreased and more food became available. Similar patterns were observed when size-specific growth rates of squid caught across the temperature and food gradients were directly compared. In addition, transport processes in the Gulf Stream and slope water played an important role in providing access and retention in favourable areas. Juvenile onshore migration seems to be driven by elevated food requirements and involves physiological adaptations to compensate for decreasing temperatures. The individual "success" in terms of growth and survival may depend, however, on access to concentrated patches of food which, in turn, will be determined by timing and the transport dynamics of the main water masses.     

Abundance and distribution of rhynchoteuthion larvae of Illex argentlnus (Cephalopoda: Ommastrephidae) in the south-western Atlantic

The results of ichthyoplankton surveys carried out by the Instituto Nacional de Pesca (INAPE) in the Argentine–Uruguayan Common Fishing Zone (AUCFZ) in summer 1980 and the winters of 1980–1982 are presented. In addition, the results of joint research with INIDEP, the Argentine and Japan Fisheries Agency, on larvae, juveniles and adults of the short-finned squid Illex argentinus carried out in winter 1989 are given, as are data obtained during the joint sardine and anchovy recruitment project carried out by R.V. Meteor in November/December 1989. The short-finned squid Illex argentinus is the most important cephalopod resource of the area, and an extended fishery is carried out on the Patagonian shelf and in the proximity of the Falkland Islands. The fishing season in the AUCFZ, for pre-spawning and spawning animals, extends from early autumn to winter (March–August). Abundance of rhynchoteuthion larvae (417–485 larvae·10 m^?2) was greatest in winter in surface temperatures of 12,1–16,9°C and surface salinities of 34,8 – 35,2 × 10^?3. The distribution of the larvae is related to the western boundary of the Brazil Current and the convergence front.     

基于不同权重的栖息地指数模型预报阿根廷滑柔鱼中心渔场

本文根据2003-2009年1-5月和2011年1-5月西南大西洋海域阿根廷滑柔鱼(Illex argentinus)的生产数据,结合遥感获得的海表面温度(SST)和海表面高度(SSH)数据,利用不同权重的栖息地指数模型来预报阿根廷滑柔鱼的中心渔场。采用外包络法,利用作业次数与SST、SSH建立适应性指数(SI)模型,依据作业次数比重和产量比重来比较不同权重的算术加权模型(AWM),从而筛选出最佳模型,并对最佳模型进行验证。结果显示,确定AWM(a=0.3,SST权重为0.3,SSH的权重为0.7)为最佳模型,当栖息地适应性指数(HSI)大于0.6时,作业次数的比重为93.23%,产量比重为89.28%,当HSI小于0.4时,作业次数的比重为2.12%,产量比重为3.35%。利用2011年1-5月的生产数据和环境数据对AWM(a=0.3)进行验证,结果显示,在HSI大于0.6的海域,各月作业次数比重均在91%以上,产量比重均在95%以上。研究表明,在阿根廷滑柔鱼渔场形成中SSH比SST更为重要,基于SST和SSH的AWM(a=0.3)能够较好地预测西南大西洋阿根廷滑柔鱼的中心渔场。     

利用基于环境因子的动态产量模型评估西南大西洋阿根廷滑柔鱼

西南大西洋阿根廷滑柔鱼,Illex argentinus,巴塔哥尼亚南部群体是重要的经济种类。海洋环境因子在柔鱼资源分布中起着重要的作用。本研究利用基于环境因子的动态产量模型评估2000-2010年的滑柔鱼的资源量。假设海洋环境因子（滑柔鱼产卵场最适宜海表温度占比）影响动态产量模型的参数K,DIC值表明在正态分布和均匀分布下均是基于环境因子的评估模型优于基本的动态产量模型。阿根廷滑柔鱼的最大可持续产量（MSY）在351600吨到685 100吨之间,资源生物量在1322400吨到1 803 000吨之间,其捕捞死亡系数均小于F_0.1和F_MSY,资源处在良好状态,没有遭受过度捕捞。本研究为应用环境因子在柔鱼类的资源评估与管理提中供了科学的参考方法。     

Spawning and fecundity of two flounder species in the Hauraki gulf,New Zealand

Spawning grounds and spawning times of the sand flounder, Rhombosolea plebeia (Richardson), and the yellow‐belly flounder, Rhombosolea leporina Gunther, in the Hauraki Gulf are described. The occurrence of female fish at different stages of ovarian development at different stations during the year, the distribution of eggs in the plankton, and changes in ovary weight during the year were used as indicators of spawning grounds and spawning times.

The sand flounder was found to spawn in the waters to the east of Waiheke and Ponui Islands, at the northern end of the Firth of Thames, from June to November.

Yellow‐belly flounders spawned during September, October, and November, slightly to the south of the sand flounder spawning grounds in a belt extending from Tapu, on the eastern side of the Firth of Thames, north‐westwards towards Ponui Island.

Fecundity of both species was approximately proportional to the weight of the fish, or to the length cubed, and less than proportional to the ovary weight. The mean fecundity of sand flounders in the Hauraki Gulf varied from approximately 100,000 eggs in a fish of 18 cm to 500,000 in a 30 cm fish. That of yellow‐belly flounders varied from approximately 250,000 eggs in a fish of 30 cm to 1.25 million in a 45 cm fish.     

The distribution of life cycle stages of two deep-water pleuronectids,Dover sole (Microstomus pacificus) and rex sole (Glyptocephalus zachirus), at the northern extent of their range in the Gulf of Alaska

Dover sole (Microstomus pacificus) and rex sole (Glyptocephalus zachirus) are both commercially valuable, long-lived pleuronectids that are distributed widely throughout the North Pacific. While their ecology and life cycle have been described for southern stocks, few investigations have focused on these species at higher latitudes. We synthesized historical research survey data among critical developmental stages to determine the distribution of life cycle stages for both species in the northern Gulf of Alaska (GOA). Bottom trawl survey data from 1953 to 2004 (25 519 trawls) were used to characterize adult distribution during the non-spawning and spawning seasons, ichthyoplankton data from 1972 to 2003 (10 776 tows) were used to determine the spatial and vertical distribution of eggs and larvae, and small-meshed shrimp trawl survey data from 1972 to 2004 (6536 trawls) were used to characterize areas utilized by immature stages. During the non-spawning season, adult Dover sole and rex sole were widely distributed from the inner shelf to outer slope. While both species concentrated on the continental slope to spawn, Dover sole spawning areas were more geographically specific than rex sole. Although spawned in deep water, eggs of both species were found in surface waters near spawning areas. Dover sole larvae did not appear to have an organized migration from offshore spawning grounds toward coastal nursery areas, and our data indicated facultative settling to their juvenile habitat in winter. Rex sole larvae progressively moved cross-shelf toward shore as they grew from April to September, and larvae presumably settled in coastal nursery areas in the autumn. In contrast with studies in the southern end of their range, we found no evidence in the GOA that Dover or rex sole have pelagic larval stages longer than nine months; however, more sampling for large larvae is needed in winter offshore of the continental shelf as well as sampling for newly settled larvae over the shelf to verify an abbreviated pelagic larval stage for both species at the northern end of their range.     

Oceanographic influences on the distribution and relative abundance of market squid paralarvae (Doryteuthis opalescens) off the Southern and Central California coast

Market squid (Doryteuthis opalescens) are ecologically and economically important to the California Current Ecosystem, but populations undergo dramatic fluctuations that greatly affect food web dynamics and fishing communities. These population fluctuations are broadly attributed to 5–7‐years trends that can affect the oceanography across 1,000 km areas; however, monthly patterns over kilometer scales remain elusive. To investigate the population dynamics of market squid, we analysed the density and distribution of paralarvae in coastal waters from San Diego to Half Moon Bay, California, from 2011 to 2016. Warming local ocean conditions and a strong El Niño event drove a dramatic decline in relative paralarval abundance during the study period. Paralarval abundance was high during cool and productive La Niña conditions from 2011 to 2013, and extraordinarily low during warm and eutrophic El Niño conditions from 2015 to 2016 over the traditional spawning grounds in Southern and Central California. Market squid spawned earlier in the season and shifted northward during the transition from cool to warm ocean conditions. We used a general additive model to assess the variability in paralarval density and found that sea surface temperature (SST), zooplankton displacement volume, the log of surface chlorophyll‐a, and spatial and temporal predictor variables explained >40% of the deviance (adjusted r² of .29). Greatest paralarval densities were associated with cool SST, moderate zooplankton concentrations and low chlorophyll‐a concentrations. In this paper we explore yearly and monthly trends in nearshore spawning for an economically important squid species and identify the major environmental influences that control their population variability.     

烟台多棘海盘车种群的个体大小分布及生殖周期

为了解烟台多棘海盘车种群个体大小分布及繁殖特征,自2010年5月至12月及2015年3月到5月,在烟台近海对该种群分别进行逐月采样调查。利用性腺指数(GI)和幽门盲囊指数(PCI)以及解剖组织学方法预测其产卵周期。结果表明：在所有的采样月份中,体长大于143mm的个体与体长小于42mm的个体在种群中均同时存在;在5-8月期间,种群结构特征以体长小于55mm的个体为主。同时该海域不同站点的采样月份中均有小个体存在,说明种群的产卵周期存在延后现象或者由于食物短缺造成了海星生长缓慢。腕长可以很好的用来衡量还行身体湿重。性腺在2010年5月-9月发育相对缓慢但在十月份迅速到达峰值20.95,之后明显下降,表明其产卵期从10月份持续至11月份。2015年3月-5月也发现了同样的现象,表明该种群自3月-5月存在另一个产卵期,且通过对卵的组织学分析也进一步证实该产卵期的存在。性腺指数和幽门盲囊指数呈负相关关系。由于食物来源短缺,烟台近海海星种群的繁殖特征极有可能受到当地近海扇贝养殖的影响。     

Dynamics of hypersaline coastal waters in the Great Barrier Reef

The coastal waters of the Great Barrier Reef (GBR) are hypersaline (salinity ∼37) during the dry season as a result of evaporation greatly exceeding rainfall, of shallow waters, and of the presence of numerous bays along the coast preventing rapid flushing. These hypersaline waters are not flushed out by salinity-driven baroclinic currents because these waters are vertically well-mixed. Instead these waters are transported by a longshore residual current and thus form a coastal boundary layer of hypersaline waters. As a result the hypersalinity distribution is 2-D with both cross-shelf and longshore gradients of salinity. The cross-shelf gradients are largely controlled by turbulent diffusion, while the longshore gradients are controlled by the residual currents that transport hypersaline waters longshore south ward in the central and southern regions of the GBR. Because every bay supplies hypersaline waters, the width of the coastal hypersaline layer increases southwards. Steady state is reached in about 100 days, which is the typical duration of the dry season. The dynamics of the GBR hypersaline coastal boundary layer thus differ from the classical inverse hypersaline systems, e.g. in Saloum River Estuary, Laguna San Ignacio, Mission Bay, Tomales Bay, San Diego Bay, Hervey Bay, Shark Bay, Coorong Coast Lagoon, Spencer Gulf, Gulf of California and many others where the salinity gradient is mainly 1-D with a dominant along-channel salinity gradient.     

Evaluation of the frequency of skipped spawning in Norwegian spring-spawning herring

Based upon an under-representation of second time spawners on the spawning grounds between 1935 and 1973, researchers have suggested that Norwegian spring-spawning (NSS) herring (Clupea harengus) frequently skip their second spawning event. In order to evaluate this claim with direct evidence, herring were collected over a period of three years from statutory surveys and commercial catches over a wide area covering the feeding, over-wintering and spawning grounds. The development stage of the ovaries was assessed and the intensity of atresia quantified. Only a negligible number of the analysed herring caught were considered likely to skip spawning, thus this phenomenon does not appear to be a common feature of the NSS herring stock at present. In addition, considering the reproductive strategy of herring, it seems doubtful that skipping the second spawning event has ever frequently occurred in this stock.     

中国沿海真鲷种群鉴别研究

对1990-1998年期间取样于中国沿海5个不同海区的真鲷(Pagrosomus major Temminck et Schlegel)样品的23项形态性状进行测量,并应用差异系数、均数差异显著性、方差分析和判别函数分析等数理统计学方法对性状特征的差异进行比较,结合产卵场、分布洄游习性以及海洋环境条件进行综合分析,存此基础上对上述海区的真鲷种群分群问题进行了探讨。结果表明．中国沿海真鲷可分为4个不同地方种群．即黄渤海地方种群、东海地方种群、闽南-南海地方种群和北部湾地方种群。文中还讨论了真鲷种质资源保护问题。     

环境因子对浙江沿岸春季日本鲐(Scomber japonicus)幼鱼生长、分布的影响

浙江沿岸海域是日本鲐（Scomber japonicus）的重要产卵场。当前针对日本鲐幼鱼生长发育的研究较少,而环境因子会对其生长、分布造成较大影响。根据2019年5月的科学观察员取样数据和环境数据,对其群体组成进行研究,运用Pearson相关性分析、广义加性模型（GAM）探究其分布及个体大小的影响因子,利用克里金插值法定量分析溶解氧对日本鲐幼鱼分布影响。结果表明：海水表层盐度（Sea Surface Salinity,SSS）是影响日本鲐幼鱼分布的主要环境因子,幼鱼集中分布于27.5-27.8盐度海域;浙江沿岸南部海域日本鲐幼鱼叉长、体重均值分别为37.6 mm、0.659 g,中部海域叉长、体重均值分别为64.2 mm、3.305 g,南部、中部海域间鲐幼鱼大小存在显著性差异（P<0.05）;日本鲐幼鱼个体大小主要受溶解氧和纬度影响,个体大小与溶解氧呈多波峰的非线性关系,随纬度增加呈先减小再增大趋势。研究表明：溶解氧是影响浙江沿岸日本鲐幼鱼生长、分布的关键环境因子;结合幼鱼分布特征及个体大小差异,推测浙江沿岸海域日本鲐幼鱼来自不同产卵批次。     

江苏近岸海域鮸时空分布特征研究

根据2006—2007年4个季度拖网专项调查数据,以鮸为目标种,以机轮底拖网禁渔区线内侧及毗邻海域为目标区域,对鮸资源时空分布特征进行分析,并探讨形成机制,结果表明:(1)江苏海域的鮸主要分布于34°00′N以南海域,主产卵区位于辐射沙脊群海域及周边近岸海域,春季为主要产卵期;(2)江苏省中南部海域为鮸早期幼体的过渡性索饵场所,幼鱼群体随时间推移呈逐步分散趋势;(3)鮸在江苏海域的栖息时间为春、夏、秋3个季节,冬季呈零星分布;(4)海底地形地貌、潮汐和海流类型、辐射沙脊群等特征要素是影响区域生物生态的重要因子。     

小黄鱼Larimichthys polyactis体长-体重关系幂指数与产卵群体空间分布相关性研究

鱼类产卵群体的相关研究一直是渔业领域关注的热点,对鱼类的产卵期、产卵场的研究是鱼类生态习性研究方向的重要组成部分。本文在前期研究证实鱼类体长-体重关系幂指数可以指示鱼类产卵期的基础上,利用2004年4个季节的东海大面积调查数据,尝试利用同一时间点上的各个调查点幂指数信息来分析鱼类产卵场分布特征,结果发现,成熟产卵个体仅在春季出现,而未成熟产卵个体4季均有分布,以此推测小黄鱼存在"跳过产卵"现象。另外发现,单靠幂指数大小,不能区分各个产卵成熟阶段,也不能区分产卵个体与非产卵个体;而怀卵个体出现站点的幂指数平均值偏低于匀速生长,空间上也具有类似特征,即幂指数相对低的地方一般对应怀卵个体相对集中的水域。本文依据幂指数分布,推断小黄鱼成熟产卵群体有3块相对集中水域,即舟山渔场近海、济州岛西南侧和江外与舟外渔场临近水域。     

西南大西洋阿根廷滑柔鱼资源时空分布研究

通过了解西南大西洋阿根廷滑柔鱼(Illexargentinus)资源丰度年间变化规律,从而对阿根廷滑柔鱼的资源可持续开发与管理打下基础。作者根据2012～2017年西南大西洋阿根廷滑柔鱼生产统计数据,利用灰色关联评价等数理方法对西南大西洋资源渔场的时空分布特征进行了分析。结果表明,2012～2017年间西南大西洋阿根廷滑柔鱼渔场重心主要在经度上分布于58°W～63°W海域,纬度主要集中在41°～44°S、47°～48°S两个区域,高平均网次产量(10 t/网)的海域主要在57°W～67°W、41°S～42°S和57°W～67°W、44°S～50°S海域。灰色关联度表明:6年间,以2015年滑柔鱼资源状况最好,其次是2014年,2012年最差,2013年、2016年和2017年处于中间水平。利用灰色关联分析的阿根廷滑柔鱼资源丰度变化趋势可用于其资源的可持续开发与管理。     

Feeding ecology of Japanese Spanish mackerel(Scomberomorus niphonius) along the eastern coastal waters of China

Feeding activities provide necessary nutrition and energy to support the reproduction and development of fish populations. The feeding ecology and dietary plasticity of fish are important factors determining their recruitment and population dynamics. As a top predator, Japanese Spanish mackerel(Scomberomorus niphonius) supports one of the most valuable fisheries in China. In this study, the feeding ecology and diet composition of Japanese Spanish mackerel spawning groups were analysed based on samples collected from six spawning grounds along the eastern coastal waters of China during spring(March to May) in 2016 and 2017. Both stomach contents and stable isotope analysis were conducted. Stomach content analysis showed that spawning groups of Japanese Spanish mackerel mainly fed on fish, consuming more than 40 different prey species. Diets were significantly different among sampling locations. The most important prey species were Stolephorus in Fuzhou, Japanese anchovy Engraulis japonicus in Xiangshan, Euphausia pacifica in Lüsi, sand lance Ammodytes personatus in Qingdao and Weihai, and Leptochela gracilis in Laizhou Bay. Stable isotope analysis showed that the trophic level of Japanese Spanish mackerel was relatively high and generally increased with latitude from south to north. In the 1980 s, the diet of Japanese Spanish mackerel was dominated solely by Japanese anchovies in the eastern coastal waters of China. The results in the present study showed that the importance of Japanese anchovies declined considerably, and this fish was not the most dominant diet in most of the investigated waters. Both the spatial variations in diet composition and changes in the dominant diet over the long term indicated the high adaptability of Japanese Spanish mackerel to the environment. Combining the results of stomach analysis and stable isotope analysis from different tissues provided more comprehensive and accurate dietary information on Japanese Spanish mackerel. The study provides essential information about the feeding ecology of Japanese Spanish mackerel and will benefit the management of its populations in the future.     

Habitat mapping of the Atlantic bluefin tuna derived from satellite data: Its potential as a tool for the sustainable management of pelagic fisheries

The feeding and spawning habitats of the overfished Atlantic bluefin tuna (BFT) are mapped in the Mediterranean Sea and used in the present proposal for selecting restricted fishing grounds. The feeding habitat is mainly traced by oceanic fronts of satellite-derived temperature and chlorophyll while the spawning habitat is mostly characterized by an important heating of surface waters. The proposal recommends opening the fishery in feeding areas in case the BFT stock is low (current situation). Only spawning areas at its latest stage could be opened once the stock has recovered to its optimum yield. Due to the possible concentration of fishing vessels if fishing areas are restricted (e.g. four-fold increase with a 1/16th restriction of the Mediterranean Sea) the inspection activities could be better targeted. Identified spawning grounds, opened or closed to fishing, could also be particularly monitored by control operations. Within the authorized areas, the habitat maps would guide fishermen to the favourable habitat reducing their costs. The habitat guided management could be able to adapt the spatial and temporal distribution of the effort to the requirements of both the fisheries’ control and the resource. Its implementation is likely to protect the stock (a) by apparently decreasing illegal fishing which accounts in the recent years for more than 1/3rd of total catches, (b) by protecting the spawners to ensure a suitable recruitment and (c) by distributing the effort to respect the population structure. The first species studied is the emblematic bluefin tuna which is at high risk of collapse due to overfishing. The approach is a priori transposable to other epipelagic species of commercial importance.     

Assessment of the genotoxicity of produced water discharges associated with oil and gas production using a fish embryo and larval test

Widespread discharge of produced waters (PW) into brackish and marine waters of the northwest Gulf of Mexico, which serve as spawning grounds for many commercially important species of fish, has raised concern over the potential impact that these complex mixtures may have on aquatic life. In the present study, the results of investigations which assess the clastogenic effects of production water discharged as the result of oil and gas production activity at two coastal sites are reported. In order to assess the impact of the PW discharges on spawning populations of aquatic organisms, chromosome aberration analysis was performed on Cyprinodon variegatus embryos 4–7 days old, which were exposed directly to varying dilutions of PW collected at Pass Fourchon and Timbalier Island for a period of 5 days. Metaphase chromosomes were scored (percentage aberration per 100 metaphases) for aberrations at a magnification of 1000 ×. When compared to the control treatment, higher levels of chromosomal aberrations were observed for treatments (2 and 4% PW) in which embryos were exposed to PW from Pass Fourchon or Timbalier Island. A maximum frequency of 13% was found to occur at the highest test concentration (4% PW), with most of these expressed as breaks (chromatid and chromosome). In contrast, exposure to water from the enclosed canal at Fourchon resulted in the formation of chromosome breaks and changes in ploidy.     

基于个体模型的东海鲐鱼渔场形成机制研究

鲐鱼Scomber japonicus资源丰富,在我国近海渔业中占有重要地位。其渔场的形成受海洋环境的制约,本文确定鲐鱼运动和物理环境之间的响应关系,建立起了基于个体的东海鲐鱼生长洄游模型。结果显示,鲐鱼集群分布与捕捞生产渔场基本吻合,鲐鱼聚集主要受台湾暖流、大陆沿岸水、黑潮影响,往往集群在一定温度范围内并在冷暖交汇区温盐梯度大偏暖水一侧。在台湾暖流和沿岸水交汇的锋面附近、台湾暖流暖水舌前端、黑潮与中国大陆沿岸水形成的潮境区域均有大量的鲐鱼聚集,并形成渔场。产卵位置的变动使偏西产卵位置的鲐鱼由于受台湾暖流影响较大,鲐鱼会呈长带状大量聚集在台湾暖流和沿岸水的锋面附近,并使在台湾暖流暖水舌前端的聚集数量增多,而偏东的产卵的鲐鱼受黑潮影响较大,聚集分布范围较大,会使黑潮形成的锋面附近聚集数量增多,而使台湾暖水舌的前端的聚集量减少。正常产卵位置在生存率方面是最佳产卵位置。研究表明鲐鱼所处空间位置不同,会影响其集群的位置,用数值模型验证了物理环境会对鲐鱼的洄游和渔场的形成产生影响。