New data from the Holsteinian interglacial site Öje, central Sweden

The stratigraphy at Öje, central Sweden, has been reinvestigated. Till-covered organic sediments were encountered in two out of three excavations, while in one excavation only till occurred. One sediment succession was analysed for pollen and macrofossils. The vegetation represented by both pollen and macrofossil flora indicates a correlation with (the latter) part of the Holsteinian Interglacial, also in accordance with the predominantly acidophilous diatom flora.     

A uranium-thorium dated late Eemian or early midlandian organic deposit from near Kilfenora between Spa and Fenit,Co. Kerry,Ireland

This paper discusses the dating of stratigraphically important Quaternary sequences from a site near Fenit, Co. Kerry, which have been the subject of debate since they were first described by Mitchell in 1970. The overall stratigraphy of the Quaternary deposits have been investigated and detailed analyses of the organic material carried out. Pollen from biogenic sediments have been analysed and samples of peat dated using the uranium-thorium disequilibrium method. The pollen assemblages match no others previously recorded in Ireland and appear to represent a cool temperate phase following the last interglacial. The uranium-thorium dates of between 114000 and 123 000 yr BP indicate that the deposit dates from Oxygen Isotope Stage 5, possibly post-dating the last temperate stage (the Eemian Stage interglacial; Oxygen Isotope Substage 5e). The dating of this deposit and the realisation that it is not penultimate temperate stage (Gortian) in age invalidates much recent speculation on the age of the Gortian interglacial.     

A new Middle Pleistocene interglacial record from Denmark: Chronostratigraphic correlation,palaeovegetation and fire dynamics

Previously only three terrestrial interglacial periods were known from southern Scandinavia, all of which could be relatively easily correlated within the central European stratigraphical framework. Here, we present a new interglacial–interstadial pollen, plant macrofossil and charcoal record from Trelde Klint, Denmark, and analyse its biostratigraphy, correlation with other European records, vegetation development, fire dynamics and absolute dating. Except for a slight truncation of the early part of the record, the pollen stratigraphy exhibits a full interglacial succession, including temperate trees (Quercus, Ulmus and Tilia) during its mesocratic stage. Macrofossil analysis allowed identification to species level for Quercus robur, Picea abies and two mosses. Conifers (Pinus and Picea) dominate the pollen record of the interglacial sequence, and the occurrence of Larix pollen in the top part of the interglacial record as well as in the interstadial sediments is especially indicative of this interglacial. The overall diversity of tree genera is rather low. These biostratigraphical features suggest that Trelde Klint is unique among Danish records, but it is similar to records from northern Germany. Numerical analyses (REVEALS and DCA) indicate that forests during the temperate stage were dense and that vegetation openness increased only towards the end of the interglacial, accompanied by increased fire occurrence. A short interstadial sequence with a dominance of Pinus and Betula and the presence of Larix is present above the interglacial deposit. We argue that lack of attention to differences in fire regimes may hamper understanding of between‐site correlations of interglacial pollen records. OSL dating, using a novel feldspar technique, yields an average age of 350±20 ka for the sandy sediments above the interglacial layers at Trelde Klint, suggesting that the whole interglacial–interstadial succession belongs to Marine Isotope Stage 11.     

Middle Pleistocene temperate deposits at Dingé, Ille-et-Vilaine,northwest France: pollen,plant and insect macrofossil analysis

Nine cores were taken from a damp depression at Dingé, Ille-et-Vilaine, northwest France. Analyses of the pollen, plant macrofossil and Coleoptera remains preserved in the same organic samples of two profiles suggest a temperate vegetation characterised by a mixed deciduous forest with mesophilous taxa (Carpinus, Fagus, Quercus) followed by a coniferous forest with Pinus and Picea. The determination of plant taxa to species was made either directly through the identification of plant macrofossil remains and pollen or indirectly through the identification of phytophagous Coleoptera specifically related to certain plants. Stratigraphical information derived from pollen, plant macrofossil and insect data indicates that this sequence may be correlated with a temperate episode older than the Eemian and younger than the Holsteinian, possibly the Bouchet 2 (Oxygen Isotope Stage 7c) or Bouchet 3 (Oxygen Isotope Stage 7a) temperate periods or the Landos Interglacial (Oxygen Isotope Stage 9 pro parte). © 1997 John Wiley & Sons, Ltd.     

Upper Pleistocene stratigraphy at the Medininkai site, eastern Lithuania: a continuous record of the Eemian-Weichselian sequence

Eemian—Weichselian sequences, located outside the maximum limit of the Late Weichselian ice sheet, provide excellent opportunities for the discovery of continuous sedimentary records encompassing the whole Last Interglacial/Glacial cycle. Such a sequence is recorded in a borehole (117P) through the succession in a small kettlehole lake located at Medininkai, eastern Lithuania. The succession consists of peat, gyttja and silt deposited on top of a Saalian till. Pollen and plant macrofossil analysis, lithological analysis, U/Th dating and mineral magnetic measurements on the sediments have allowed 19 lithostratigraphic units and 16 local pollen assemblage zones (LPAZ) to be identified. The palaeocarpological record reveals a clear transition from the Saalian Glacial to the Weichselian stadial and interstadial phases. The mineral magnetic parameters suggest a good correlation between the concentration of magnetic minerals and stadial and interstadial periods. The Merkine (Eemian) Interglacial and two Early Weichselian Interstadials, Jonionys 1 (Brörup) and Jonionys 2 (Odderade), separated by cryomers, are identified. Intervals interpreted as analogous to the Middle Weichselian Denekamp and Hengelo interstadials are also recognized on the basis of pollen assemblages. The results show alternating periglacial and interstadial palaeoenvironments in Lithuania during the Early and Middle Weichselian and are of importance for Late Pleistocene palaeoenvironmental reconstruction of the Baltic area as a whole.     

Early and Middle Pleistocene vegetation history of the Médoc region,southwest France

Pleistocene deposits, together with their pollen, plant macrofossil, foraminiferal, dinoflagellate and coleopteran remains, from five sites along the Atlantic coast of the Médoc Peninsula are described and discussed. Sediments making up the Négade Formation are shown to have been laid down under either estuarine or lagoonal conditions when closed Quercus–Pinus‐Tsuga canadensis regional woodland existed. Comparison with plant records from The Netherlands indicates that these deposits are most likely attributable to either the Early Pleistocene Bavel Interglacial (marine oxygen isotope stage (MIS) 31), or an interglacial of the Waalian (MIS 37–49) or Tiglian (MIS 63–79). In addition, clays assigned to the Argiles du Gurp sensu stricto, were similarly deposited in either an estuary or lagoon, which subsequently was cut off from the sea. A freshwater lake with vegetation dominated by Azolla filiculoides then developed. This was succeeded by reedswamp and an organic mud (termed Lignite in the corresponding French stratigraphical records) formed. Regional Quercus–Abies woodland was replaced by one with Pinus dominant and Pterocarya a minor component. Comparison with plant records from France and other parts of Europe suggest that the clays and organic mud might be correlated with the Holsteinian (Praclaux) Interglacial (MIS 11c). Copyright © 2003 John Wiley & Sons, Ltd.     

Geology,stratigraphy and palaeoenvironmental evolution of the Stephanorhinus kirchbergensis-bearing Quaternary palaeolake(s) of Gorzów Wielkopolski (NW Poland,Central Europe)

The sedimentary succession exposed in the Gorzów Wielkopolski area includes Eemian Interglacial (MIS 5e) or Early Weichselian (MIS 5d–e) deposits. The sedimentary sequence has been the object of intense interdisciplinary study, which has resulted in the identification of at least two palaeolake horizons. Both yielded fossil remains of large mammals, alongside pollen and plant macrofossils. All these proxies have been used to reconstruct the environmental conditions prevailing at the time of deposition, as well as to define the geological context and the biochronological position of the fauna. Optically stimulated luminescence dating of the glaciofluvial layers of the GS3 succession to 123.6 ± 10.1 (below the lower palaeolake) and 72.0 ± 5.2 ka (above the upper palaeolake) indicate that the site formed during the Middle–Late Pleistocene (MIS 6 – MIS 5). Radiocarbon-dating of the lacustrine organic matter revealed a tight cluster of Middle Pleniglacial Period (MIS 3) ages in the range of ~41–32 ka cal bp (Hengelo – Denekamp Interstadials). Holocene organic layers have also been found, with ¹⁴C ages within a range of 4330–4280 cal bp (Neolithic). Pollen and plant macrofossil records, together with sedimentological and geochemical data, confirm the dating to the Eemian Interglacial.     

Environmental and climatic conditions at a potential Glacial refugial site of tree species near the Southern Alpine glaciers. New insights from multiproxy sedimentary studies at Lago della Costa (Euganean Hills, Northeastern Italy)

It has been hypothesized that refugia of thermophilous tree species were located in Northern Italy very close to the Alps, though, this hypothesis has yet to be tested thoroughly. In contrast to Central and Southern Italy with its relative wealth of data, only a few fragmentary records are currently available from Northern Italy for the last Glacial (Würm, Weichselian). Our new study site Lago della Costa lies adjacent to the catchment of the megafans of the Alpine forelands and the braided rivers of the Northeastern Po Plain that have so far inhibited the recovery of continuous Glacial and Late-Glacial records. We analyze pollen, plant macrofossils, charcoal and ostracods to reconstruct the vegetation, fire and lake history for the period 33,000–16,000 cal. BP. We compare our data with Glacial records from Southern Europe to discuss similarities and dissimilarities between these potential refugial areas. A comparison with independent paleoclimatic proxies allows to assess potential linkages between environmental and climatic variability. New macrofossil and pollen data at Lago della Costa unambiguously document the local persistence of boreal tree taxa such as Larix decidua and Betula tree species around the study site during the last Glacial. The regular occurrence of pollen of temperate trees in the organic lake sediments (fine-detritus calcareous gyttja) suggests that temperate taxa such as Corylus avellana, Quercus deciduous, Tilia, Ulmus, Fraxinus excelsior, Carpinus, Abies alba and Fagus sylvatica, most likely survived the Last Glacial Maximum (LGM) at favorable sites in the Euganean Hills. The percentage values of temperate trees are comparable with those from Southern Europe (e.g. Monticchio in Southern Italy). We conclude that the Euganean Hills were one of the northernmost refugial areas of temperate taxa in Europe. However, the relative and absolute abundances of pollen of temperate trees are highly variable. Pollen-inferred declines of temperate tree communities (e.g. Quercetum mixtum) and low ostracod-inferred water levels at Lago della Costa correspond to the cold Heinrich events H-2 (LGM; 23,000–19,000 cal. BP) and H-3 (around 28,000 cal. BP), as recorded in the marine sediments of the North Atlantic. Similar patterns of significant temperate tree population collapses during cold Heinrich events are recorded at southern Mediterranean sites (e.g. Monticchio and the Alboran Sea). These findings suggest close linkages between Northern Atlantic and South-Central European climates during the past Glacial.     

Pollen analytical biostratigraphy of the last five climatic cycles from a long continental sequence from the Velay region (Massif Central,France)

In the volcanic region of Velay (Massif Central, France), lake sediment sequences derived from maar craters situated close to one another (Ribains, Praclaux and Lac du Bouchet) have been correlated on the basis of tephrostratigraphical analyses. This has enabled the construction of a sequence (2980 pollen spectra) that begins during the glaciation preceding the Holsteinian and ends in the present. This sequence covers the period from 450 ka, i.e. from the end of marine oxygen isotope stage 12, to the present time, and includes five climatic cycles. The study of a second core from the Praclaux site completed the long Velay sequence. The biostratigraphy of the whole sequence is described in detail and illustrated by a synthetic and simplified pollen diagram. Thirteen temperate forest episodes (interglacials or interstadials) and as many cold periods (glacials or stadials) are defined. Each temperate episode is characterised by particular forest development and vegetation dynamics, and all of them, with the exception of the Holocene, begin and end with a Pinus forest. The vegetation dynamics observed during the Holocene resemble more closely those recorded during the interstadials than during the interglacials. Copyright © 2000 John Wiley & Sons, Ltd.     

Evidence from coleopteran assemblages for a short but intense cold interlude during the latter part of the MIS11 Interglacial from Quinton, West Midlands, UK

A sequence of fossiliferous organic sediments spanning the whole of an interglacial has been investigated from Quinton, near Birmingham, United Kingdom This interglacial can be correlated on palynological grounds with the Hoxnian Interglacial of England; equivalent to the Holsteinian Interglacial of central Europe and to Marine Isotope Stage 11. A pollen diagram and coleopteran spectrum were obtained from the same series of samples. By using the mutual climatic range method on the coleopteran assemblages, it has been possible to quantify the thermal climatic conditions at this time. They show clear evidence for a sudden and intense climatic oscillation during the late temperate stage of the interglacial—named the Quinton Cold Interlude. At this time mean temperatures of the warmest month (July) declined suddenly by at least 5 °C and mean temperatures of the coldest months (January and February) by more than 10 °C. Subsequently cool-temperate conditions returned. The pollen diagram through this interval shows little or no response to this climatic event. It is too early yet to generalise about the regional significance of the Quinton Cold Interlude but a similar climatic oscillation also occurs towards the closing stage of the MIS11 Interglacial at Hoxne, Norfolk, UK, suggesting that the cold interlude was at least regional in extent. A short climatic oscillation of similar intensity has also been interpreted from coleopteran evidence towards the end of the MIS13 Interglacial at Waverley Wood Farm Pit, Warwickshire, UK, indicating that such events may characterise the later stages of other interglacials. These severe climatic oscillations towards the end of interglacials may have relevance to our understanding of the closing phase of the present interglacial.     

Differences in pollen and macrophytic remains in sediments from various depths in a small kettle-hole lake in southern Finland

Stratigraphical analyses from four locations within the same lake revealed differential dispersal of pollen in sediment, and horizontal movement and focusing of sediment, but revealed only slight differences in pollen and macrofossil assemblages. Between 9,000 and 8,000 B.P., organic sediments focused into the central depth of the basin and no sediment accumulated in the littoral sites in water < 7.5 m deep. From around 5,000 to 4,500 B.P., deposition took place more or less continuously at all four sites. After 1,600 B.P., the marked change in lithology is due to a heavy rise in organic and minerogenic input from the unforested watershed. The littoral cores show that not only does the water depth cause rcdeposition and focusing of sediment, but also the morphometry of the basin. The comparison between macrofossils and pollen, both produced primarily by the same vegetation around the lake, confirmed the reconstruction of local environmental short-term changes which occurred after the clearance of forest vegetation by man.     

Late and Middle Pleistocene deposits at Somersham,Cambridgeshire, U.K.: a model for reconstructing fluvial/estuarine depositional environments

Gravel quarries at Somersham, Cambridgeshire, have yielded evidence for a Pleistocene channel of the R. Great Ouse, containing temperate stage sediments between cold stage sediments. In the earlier cold stage, fluviatile gravels and floodplain loessic sediment accumulated. In the later cold stage a further series of gravel units and floodplain sediments were deposited, together with lake sediments. The lake sediments are associated with Lake Sparks, dammed by Late Devensian ice in the Wash at ca. 18.5 ka BP. The lake sediments overlie gravels with a radiocarbon date from an organic horizon indicating a Middle Devensian age. Clast lithological analyses from the earlier and later gravels suggest that reworking of gravels has occurred within a relatively stable catchment. The petrography of the earlier cold stage loessic sediment and temperate stage fine sediment indicates an Anglian affinity, which conflicts with the biostratigraphic interpretation. Pollen and macroscopic plant remains from sediments of both cold stages and from the temperate stage indicate, respectively, assemblages with a typical full-glacial aspect with a rich flora of shrubs and open ground herbs(including an assemblage at ca. 18 ka), and temperate freshwater and marine-influenced organic sediments. On the basis of pollen analysis these are ascribed to substages Ip II and III of the Ipswichian Stage(O.I.S. 5e), with a Pinus-Quercus-Corylus biozone in the former and a biozone with Carpinus in the latter. Marine-influenced sediments, at −3.7 to −0.3 m OD, indicate transgression in Ip II and regression in Ip III.Molluscan assemblages from the temperate stage and the later cold stage are described; two are from the Late Devensian, at a time near the maximum extension of ice into the Wash. Foraminifer and ostracod faunas are described from post-Ipswichian sediments and may be reworked. Radiocarbon dates confirm the age of the later gravel suite as Devensian and a calibration of the measurements is given. Amino acid ratios from Corbicula fluminalis valves from temperate stage sediments are reported, with measurements from different parts of the valve; the results tend to support an Ipswichian age. TL measurements of the earlier cold stage loessic sediment and associated sand indicate a pre-Ipswichian age for the sediments. The earlier cold stage is correlated with the pre-Ipswichian cold stage, the Wolstonian of Mitchell et al.(1973); problems with this correlation are discussed.Various periglacial phenomena, including thermal contraction networks and cracks, diapirs, involutions and coversand are associated with the Devensian sequence. The complex environmental history, based on stratigraphy and palaeontology, is described, and related to other nearby sites in southern Fenland.     

Sedimentology,palaeoecology and geochronology of Last Interglacial deposits from Deeping St James,Lincolnshire, England

Pollen, plant macrofossil, molluscan and coleopteran data from organic muds below the low terrace of the River Welland at Deeping St James, Lincolnshire indicate deposition in the mixed oak forest phase of a Late Pleistocene interglacial. Coleopteran and molluscan data suggest summer temperatures up to 4°C warmer than at present in eastern England, and plant macrofossil material suggests a climate more continental than that of Britain in the Holocene. No direct analogue of this biota, however, exists currently in Europe. Biostratigraphical indications from the pollen coleoptera and Mollusca suggest an age in the Ipswichian Interglacial. Thermoluminescence dates between 120 ka and 75 ka and amino-acid ratios with a mean of 0.11 show that deposition of the sediments took place during Oxygen Isotope Stage 5. This accurate dating of a partial Ipswichian succession allows discussion of the ages of a number of other interglacial sites in eastern England of assumed Ipswichian age. Copyright © 1999 John Wiley & Sons, Ltd.     

Palynology of organic beds below Devensian glacigenic sediments at Pen-y-bryn,Gwynedd, North Wales

Pollen diagrams from organic facies overlain by glacigenic sediments at Pen-y-bryn, North Wales (53°7′N, 4°16′W), suggest that it is the first locality in the British Isles to provide evidence for several Devensian interstadials prior to Devensian stadial glaciation(s). The evidence is not, however, unequivocal. At least two main episodes of organic sedimentation are indicated, separated by an interval of uncertain duration. One eposide records a Pinus-Picea-Betula forest. A second, possibly later, event, with tree pollen less than 10% of total land pollen (TLP), suggests a cool, largely unforested environment. A third episode, with tree pollen up to 25% of TLP, may be related to the latter or be of intermediate age, whereas a fourth and later episode of organic accumulation may be of reworked material. Radiocarbon dating of organic deposits and of a wood macrofossil enclosed in basal till is inconclusive, as is preliminary uranium-series dating. The principal biogenic episodes may correlate with Oxygen Isotope Substages 5c and 5a, but as the pollen records reflect facies floras, the correlation remains tentative; it is also possible that other temperate periods are recorded. Further resolution of the chronostratigraphy is required to help clarify the problematic correlation of interstadial and stadial events in northwest Europe and to correlate more accurately the biogenic and glacigenic facies at Pen-y-bryn with their Pleistocene equivalents elsewhere.     

Re‐interpretation of the Meikirch pollen record,Swiss Alpine Foreland,and implications for Middle Pleistocene chronostratigraphy

The Meikirch drilling site in the Swiss Midlands north of Bern is re‐interpreted using a combination of sedimentological logging, pollen analyses and luminescence dating. The sedimentary sequence comprises about 70 m of lacustrine deposits, overlain by about 39 m of coarse glacial outwash interpreted to represent at least two independent ice advances. Pollen analyses of the apparently complete limnic sequence reveal a basal late glacial period followed by three warm phases that are interrupted by two stadial periods (Meikirch complex). The warm periods were previously correlated with the Holsteinian and Eemian Interglacials. According to luminescence dating, and with consideration of evidence for Middle Pleistocene climate patterns at other central European sites, a correlation of the Meikirch complex with marine isotope stage (MIS) 7 is now proposed. If this correlation is correct, it implies the presence of three intervals with interglacial character during MIS 7. However, the late Middle Pleistocene vegetational features of the Meikirch complex show significant differences when compared with the pollen record from the Velay region, central France. Possible explanations for this discrepancy are distinct Middle Pleistocene patterns of atmospheric circulation over central Europe and a different distribution of vegetation refugia compared to the Eemian Interglacial and the Holocene. Copyright © 2005 John Wiley & Sons, Ltd.     

Coherent slumping of early postglacial lake sediments at Hall Lake, Ontario, Canada

Cores of organic postglacial sediment from Hall Lake, southern Ontario, Canada, are described. The laminated, calcareous sediments deposited between 9,000 and 8,000 B.P. have coherently slumped into the deep part of the lake, causing overturning and replication of parts of the sequence. This process may be considered as a mechanism for sediment focusing, since it involves reworking of marginal sediments into deeper areas. Unlike previously described mechanisms, it may be a source of error in the interpretation of diagrams of pollen percentages as well as accumulation rates. The slumping has only been recognized through the distortion of the laminae. It might not be possible to recognize slumping in unstratified sequences.     

Evolution of the Thames estuary during MIS 9: insights from the Shoeburyness area, Essex

Multidisciplinary, litho-, bio- and amino-stratigraphical investigations of the infills of buried channels on the coast of eastern Essex have a direct bearing on the differentiation of MIS 11 and MIS 9 in continental records. New data are presented from Shoeburyness, where a deeply incised channel filled with interglacial sediment can be directly related to the terrace stratigraphy of the River Thames. Fossil assemblages confirm that the interglacial beds began accumulating in a freshwater environment, which became transformed into a dynamic estuary as relative sea-levels rose. Pollen data confirm that this occurred early in the interglacial when mixed oak forest was becoming established.The geological context of the sediments indicates that they post-date the Anglian glaciation, yet pre-date the Barling Gravel terrace aggradation, which has been ascribed to MIS 8. Amino acid racemisation data based on Bithynia opercula further constrain the age to the Hoxnian (=MIS 11) or to MIS 9. An MIS 9 attribution is favoured because (i) AAR data suggest that the sequence post-dates the interglacial channel-fill at Clacton, which is widely ascribed to the Hoxnian; (ii) the bivalve Corbicula occurred early within the interglacial (unlike its late appearance during the Hoxnian); and (iii) the sequence includes evidence for a marine transgression that occurred earlier in the interglacial cycle than it did at local Hoxnian sites.Plant macrofossil remains suggest that the early part of the Shoeburyness interglacial was associated with warmer-than-present summer temperatures. This is in keeping with inferences from sites at Barling, Cudmore Grove and Purfleet, which are also attributed to MIS 9. All three sites are similar in terms of their palaeo-vegetation and inferred relative sea-level histories and provide an emerging picture of this temperate episode in southern Britain.     

Vegetation dynamics during the Younger Dryas-Holocene transition in the extreme northern taiga zone, northeastern European Russia

Vegetation dynamics during the Younger Dryas-Holocene transition in the extreme northern taiga zone of the Usa basin, northeastern European Russia, were reconstructed using plant macrofossil and pollen evidence from a sediment core from Lake Llet-Ti. The pollen stratigraphy during the Younger Dryas (about 12 500-11 500 cal. yr BP) is characterized by pollen types indicative of treeless arctic vegetation, whereas the macrofossil evidence shows the occurrence of scattered spruce and birch trees around the lake. The Younger Dryas-early Holocene transition is characterized by a rapid increase in vegetation density, including an increase in the birch population, followed by the expansion of the spruce population at about 10 000 cal. yr BP. Dense spruce-birch forest dominated until 5000 cal. yr BP. Our results contribute to the debate about the Lateglacial environments in northern Russia, and illustrate the importance of plant macrofossil records in Lateglacial vegetation reconstructions.     

Macrofossil evidence of alder (Alnus sp.) in Britain early in the Late Glacial Interstadial: implications for the northern cryptic refugia debate

Wood macrofossil remains of alder and willow/poplar have been recovered from a sediment sequence in the valley of the Turker Beck in the Vale of Mowbray, North Yorkshire. These remains have yielded radiocarbon dates early in the Devensian Late Glacial (14.7–14k cal a bp ), equivalent to the early part of the Greenland Interstadial (GI-1e) of the GRIP ice-core record. These are the earliest dates recorded for the presence of alder in the Late Glacial in the British Isles. Associated biological remains have provided a palaeoenvironmental record for this early part of the Greenland Interstadial, generally indicative of open environments dominated by herbaceous taxa on both the wetland and dryland surfaces. However, stands of alder, birch and willow woodland were also present, and indicate the possibility that such tree species survived in cryptic refugia in Britain as elsewhere in northern Europe during the Last Glacial Maximum. The absence of alder pollen at Turker Beck, in a sequence in which its macrofossil remains are relatively abundant, lends support to the view that pollen can be a poor indicator of the presence of tree species in Late Glacial sequences in northern and western Europe.