A late Early Pleistocene pollen record from Fontana Ranuccio (central Italy)

A pollen record from a sediment core drilled underneath the Acheulian site of Fontana Ranuccio provides new data on the vegetation characters in central Italy during the late Early Pleistocene, in a forest phase possibly corresponding to a substage of Marine Isotope Stage 21. A number of tree taxa nowadays extinct in Europe (Cathaya, Tsuga, Taxodium type, Carya and Pterocarya) are present at Fontana Ranuccio in a temperate phase characterised by dense forest conditions, dominated by Alnus and Pterocarya. The comparison of the Fontana Ranuccio record with other dated sites distributed along the Italian peninsula indicates that the floristic and vegetational characters of the Mid Pleistocene revolution present a rather distinct succession of biostratigraphic events. Compared to other Early Pleistocene pollen records from southern Europe, the Italian Peninsula shows greater floristic similarities with the Eastern Mediterranean regions than with the Iberian Peninsula. Copyright © 2011 John Wiley & Sons, Ltd.     

Early Pleistocene climate changes in the central Mediterranean region as inferred from integrated pollen and planktonic foraminiferal stable isotope analyses

Vegetation inherited from a Pliocene subtropical climate evolved through obliquity oscillations and global cooling leading to modern conditions. An integrated, highly time-resolved record of pollen and stable isotopes (δ¹⁸O and δ¹³C of Globigerina bulloides) was obtained to understand vegetation responses to Early Pleistocene climate changes. Continental and marine responses are compared in the Central Mediterranean region with a particular consideration of environmental changes during anoxic events.Pollen data illustrate vegetation dynamics as follows: [1] development of mesothermic elements (warm and humid conditions); [2] expansion of mid- and high-altitude elements (cooler but still humid conditions); and [3] strengthening of steppe and herb elements (cooler and dry conditions). These successions correlate with precession. δ¹⁸O variations recorded by Globigerina bulloides define two cycles (MIS 43-40) related to obliquity. At northern low- to mid-latitudes, the pollen signal records temperature and wetness changes related to precession even during global climate changes induced by obliquity. This may result in unexpected increasing wetness during glacial periods, which has to be considered specific to the Central and Eastern Mediterranean region. Lastly, an analysis of anoxic events reveals that enhanced runoff is indicated by increasing frequency of the riparian trees Liquidambar and Zelkova.     

Pollen record from Colle Curti and Cesi: Early and Middle Pleistocene mammal sites in the Umbro–Marchean Apennine Mountains (central Italy)

The palynological record from the Colle Curti and Cesi continental deposits has been examined in order to identify the main palaeofloristic and vegetational changes between 0.99 and 0.6–0.7 Ma. These data show a progressive increase in aridity, as well as a progressive decrease in temperature, which are associated with the transition in dominance from the 41 to 100 ka cyclicity in the Milankovitch orbital record during the Middle Pleistocene. The disappearance of Tsuga, recorded during the lower part of the Brunhes Chron, also has been related to a shift in global aridity. During the successive open vegetational phases (glacials), Chenopodiaceae and Artemisia progressively increase, whereas Cyperaceae decrease. Forest phases (interglacials) are successively dominated by Tsuga, Abies with Picea and, finally, Pinus; but all lack significant expansion of broad‐leaved deciduous taxa. Palynological and sedimentological data, in addition to taphonomic interpretations, demonstrate the occurrence of several hiatuses in the lower parts of the interglacials. These hiatuses are considered to represent the palaeoenvironmental response to climatic changes affecting local sedimentological and geomorphological conditions. Copyright © 2000 John Wiley & Sons, Ltd.     

Postglacial vegetation and climate dynamics in the Seymour‐Belize Inlet Complex,central coastal British Columbia,Canada: palynological evidence from Tiny Lake

A pollen‐based study from Tiny Lake in the Seymour‐Belize Inlet Complex of central coastal British Columbia, Canada, permits an evaluation of the dynamic response of coastal temperate rainforests to postglacial climate change. Open Pinus parklands grew at the site during the early Lateglacial when the climate was cool and dry, but more humid conditions in the later phases of the Lateglacial permitted mesophytic conifers to colonise the region. Early Holocene conditions were warmer than present and a successional mosaic of Tsuga heterophylla and Alnus occurred at Tiny Lake. Climate cooling and moistening at 8740 ± 70 ¹⁴C a BP initiated the development of closed, late successional T. heterophylla–Cupressaceae forests, which achieved modern character after 6860 ± 50 ¹⁴C a BP, when a temperate and very wet climate became established. The onset of early Holocene climate cooling and moistening at Tiny Lake may have preceded change at more southern locations, including within the Seymour‐Belize Inlet Complex, on a meso‐ to synoptic scale. This would suggest that an early Holocene intensification of the Aleutian Low pressure system was an important influence on forest dynamics in the Seymour‐Belize Inlet Complex and that the study region was located near the southern extent of immediate influence of this semi‐permanent air mass. Copyright © 2008 John Wiley & Sons, Ltd.     

Cesi,an early Middle Pleistocene site in the Colfiorito Basin (Umbro-Marchean Apennine), central Italy

Near the village of Cesi, at the head of the Chienti River, in the Colfiorito Basin (Umbro-Marchean Apennines, central Italy), fluvio-lacustrine deposits have yielded mammal fossil remains. The results of a multidisciplinary investigation indicate that the vertebrate-bearing sediments date about 700 ka and accordingly provide a fossil assemblage for the Middle–Late Galerian. Palynological investigations carried out from sediments underlying the fossiliferous level suggest predominantly cold and dry conditions, whereas the fauna suggests a slight climatic amelioration towards cool and moist conditions in the uppermost part of the sequence. © 1997 John Wiley & Sons, Ltd.     

An experimental approach to the palynology of cave deposits

Surface sediment, speleothems, and bat guano from two adjacent, topographically different cave sites in eastern Spain have been studied palynologically to elucidate the potential of cave sediments for palaeoenvironmental reconstruction. A cave opening with a large entrance and constant width presented far fewer problems of alteration in the pollen assemblages than a sac‐like cave opening with a constricted entrance. Pollen concentration is linked primarily to the amount of pollen input rather than to the result of post‐depositional alteration. Sampling should be undertaken away from parietal and rear areas and avoiding moisture zones. Lateral differences in the pollen spectra indicate that sampling should be on the basis of a multiple‐profile approach and selection of dry rather than wet sediments. If these procedures are followed, within‐cave sediments can realistically reflect not only local but also regional vegetation of the site. Copyright © 2000 John Wiley & Sons, Ltd.     

A coastal reservoir of terrestrial resources for neanderthal populations in north‐eastern Iberia: palaeoenvironmental data inferred from the small‐vertebrate assemblage of Cova del Gegant,Sitges, Barcelona

Coastal areas as reservoirs of resources for hominid groups have been widely studied in recent years. These areas combine marine with terrestrial and wetland resources and would have been optimum sites for hominids, including Neanderthals. This is the case with the Cova del Gegant, a cave that today opens directly onto the Mediterranean Sea and is located in the north‐eastern Iberian Peninsula. The geomorphological evolution of the Massis del Garraf has provided evidence that during the late Pleistocene there was a littoral platform between 8 and 13 km wide in front of the Cova del Gegant. Within this framework, the data derived from analysis of the small vertebrates and large mammals recovered from Cova del Gegant, including taxa currently absent from the Massis del Garraf, suggest that the landscape surrounding the cave provided a richer terrestrial ecosystem for Neanderthals than is available in this zone today. Analysis of the small‐vertebrate association from the cave reveals that the landscape surrounding the cave was dominated by woodland‐edge and open environments and that the climate was Mediterranean. The results have been compared with the only Iberian site with similar characteristics to the Cova del Gegant, Gorham's cave (southern Iberia, Gibraltar), revealing differences and similarities in the landscape and climate on the basis of the small‐mammal assemblages as well as the differences in the accessibility to terrestrial mammalian resources for the Neanderthal groups. The landscape and the climate were reasonably similar at the two sites, but the differences in the accessibility of resources for the Neanderthals are directly related to the location of the sites and the coastal position. Whereas the Cova del Gegant was on a route of mammal migration (between the Ebro Valley and France) suited for securing terrestrial resources, Gorham's cave is located on a small peninsula with a lower abundance of terrestrial mammal resources. This is probably why the Neanderthal groups at Gorham's cave exploited marine resources, whereas there is no evidence of marine resources having been exploited at Cova del Gegant, even though the seashore was nearby. Copyright © 2011 John Wiley & Sons, Ltd.     

Small mammals from the early Pleistocene of the Granada Basin, southern Spain

The Pliocene and Pleistocene continental sedimentary records of the western sector of the Granada Basin, southern Spain, consist of alternating fluvial and lacustrine/palustrine sediments. Two Quaternary sections from this sector have been sampled: Huétor Tájar and Tojaire. They have yielded remains of rodents, insectivores and lagomorphs. The presence in the Huétor Tájar and Tojaire sections of Mimomys, Apodemus atavus, Castillomys rivas and two different species of Allophaiomys, indicates an Early Pleistocene age. These deposits, which are related to a fluvio-lacustrine system, can be differentiated from an older (Pliocene) braided fluvial system. Their dating has important repercussions on the paleogeographic reconstruction of the basin. The conditions inferred from the ecological preferences of the small mammal associations are wet and cold. These associations suggest a predominance of open herbaceous habitats, followed by forested habitats; semiaquatic habitats are the least represented.     

Pleistocene forest on Tenerife's southern coast: a case study of Montaña Negra

Montaña Negra is a 121 m cinder cone in the Bandas del Sur region of southern Tenerife. Formed in the Middle Pleistocene, it comprises alternating phonolitic pumice deposits and scoria layers; the latter are extremely fossiliferous with good taphonomical fidelity. ⁴⁰Ar/³⁹Ar age determination provides new dates of 302 ± 7.6 ka and 299.9 ± 11.4 ka for the Lower and Upper Aldea Blanca pumice fall deposits, respectively. This chronological constraint allows comparison of the palaeo‐habitat with the global climate at the time of pyroclastic activity. Abundant terrestrial gastropod species and rare disarticulated Coleoptera fragments are to be found. The occurrence of the endemic semi‐slug genus Plutonia (Family Vitrinidae) is significant in indicating a woodland habitat in the region during the Middle Pleistocene. We suggest that this may have been forest, possibly dominated by laurel, which is in stark contrast to the present‐day semi‐desert. Copyright © 2011 John Wiley & Sons, Ltd.     

Fossil hyrax dung and evidence of Late Pleistocene and Holocene vegetation types in the Namib Desert

Pollen was derived from fossil dung of herbivorous hyraxes, deposited in a rock shelter on the highest mountain in Namibia, Dâures or Brandberg, situated on the Namib Desert margin. Radiocarbon dating ranging in age between modern times and 30 000 yr BP showed it represents the first empirical pollen evidence of continental palaeovegetation during the Late Pleistocene along the western escarpment of southern Africa. The initial results indicate Last Glacial Maximum vegetation differed totally from the current pattern as vegetation types were dominated by small Asteraceae shrubs, in contrast to those of the Holocene and modern times which show more succulents, grass and woody elements (arboreal pollen). The results suggest that Cape floral communities did not reach into the tropics along the western escarpment of Africa, despite such pollen types occurring in marine cores. Copyright © 2004 John Wiley & Sons, Ltd.     

Late Quaternary vegetation,climate and fire dynamics inferred from the El Tiro record in the southeastern Ecuadorian Andes

In order to study the stability and dynamics of mountain rainforest and paramo ecosystems, including the biodiversity of these ecosystems, the Holocene and late Pleistocene climate and fire variability, and human impact in the southeastern Ecuadorian Andes, we present a high‐resolution pollen record from El Tiro Pass (2810 m elevation), Podocarpus National Park. Palaeoenvironmental changes, investigated by pollen, spores and charcoal analysis, inferred from a 127 cm long core spanning the last ca. 21 000 cal. yr BP, indicate that grass‐paramo was the main vegetation type at the El Tiro Pass during the late Pleistocene period. The grass‐paramo was rich in Poaceae, Plantago rigida and Plantago australis, reflecting cold and moist climatic conditions. During the early Holocene, from 11 200 to 8900 cal. yr BP, subparamo and upper mountain rainforest vegetation expanded slightly, indicating a slow warming of climatic conditions during this period. From 8900 to 3300 cal. yr BP an upper mountain rainforest developed at the study site, indicated by an increase in Hedyosmun, Podocarpaceae, Myrsine and Ilex. This suggests a warmer climate than the present day at this elevation. The modern subparamo vegetation became established since 3300 cal. yr BP at El Tiro Pass. Fires, probably anthropogenic origin, were very rare during the late Pleistocene but became frequent after 8000 cal. yr BP. Copyright © 2007 John Wiley & Sons, Ltd.     

Anthropogenic changes in the landscape of west Java (Indonesia) during historic times,inferred from a sediment and pollen record from Teluk Banten

Palynological and charcoal analyses of shallow marine core 98‐28 from the northern coastal area of West Java provide a regional vegetation history during the last few centuries. Reliable chronostratigraphical control is provided by ²¹⁰Pb analyses and the occurrence of the 1883 Krakatau ash/tsunami layer as a time marker. The results permit the distinction of four successive stages, reflecting increased disturbance and land clearance, with some evidence for the presence of deciduous lowland forests in the Banten area during the early Holocene. The establishment of coconut and pine plantations and the severe loss of biodiversity in the last few decennia are also echoed in the pollen record. The effect of the Krakatau eruption was insignificant compared with human impact on vegetation in the Banten area. Copyright © 2004 John Wiley & Sons, Ltd.     

The Würm in NW Iberia, a pollen record from Area Longa (Galicia)

The pollen record at Area Longa is the westernmost sequence available for investigation of the last glaciation in continental Europe. It is located in a region, NW Iberia, for which data from times earlier than the late glacial period are scarce. It comprises a series of exposed limnetic levels that lie above an Eemian (Oxygen Isotope Stage [OIS] 5e) beach and are separated by inorganic layers. The oldest limnetic level (Level I), attributed to the early glacial period (OIS 5a to OIS 5d), shows a dominance of woodland with high proportions of Fagus pollen and is tentatively identified with St. Germain I. The lower pleniglacial (OIS 4) Level II records a stadial landscape of grassland and shrub. Level III, from the pleniglacial interstade (OIS 3), reflects a complex period in which three warmer woodland phases alternated with periods of more open vegetation. This cyclical behavior correlates with the ice core isotope record and with the general tendencies observed in other Würmian pollen records, but the composition of our pollen profiles differs from those observed in these other records. In NW Iberia, the dominant trees were deciduous taxa, not conifers. Of particular note is the presence of lowland Fagus woodlands during the pre-Würm, and the occurrence of Carpinus considerably farther west than the boundary of its current distribution in the Iberian Peninsula.     

Iberian Plio‐Pleistocene biochronology: micromammalian evidence for MNs and ELMAs calibration in southwestern Europe

A multivariate biochronological analysis of the Plio‐Pleistocene Iberian rodent fossil record is presented. Among more than 300 fossil sites, 49 sites have been selected. The ordination of the fossil sites selected from the Iberian Peninsula has been achieved by the appearance event ordination method. After calibration of this ordination with geochronological dates at some of the localities, numerical dates are proposed for all the localities studied. Hence, mammal fossil sites non‐datable by other means (palaeomagnetism, radiometric dating, etc.) have been dated with a relatively high degree of confidence. Final results allow dating of boundaries between the Plio‐Pleistocene European Land Mammal Ages (ELMAS) or the Mammal Neogene (MN) units. These boundaries agree with previous studies using magnetostratigraphical calibration of these scales. Copyright © 2004 John Wiley & Sons, Ltd.     

Middle and Late Pleistocene fluvial systems in central Poland

This reconstruction of the fluvial palaeogeography of central Poland is based on an exhaustive and critical review of the published and archival data for the Middle and Late Pleistocene sediments of the area. The Warsaw Basin in central Poland was a major confluence area during the Middle and Late Pleistocene. The fluvial watersheds have been only slightly modified since that time. The past river systems resembled the contemporary one, therefore indicating rebuilding during successive interglacials, at least since the Holsteinian when the sea undoubtedly occupied the southern Baltic Basin. The Weichselian fluvial system was strongly influenced by the Scandinavian ice sheet, especially by meltwater runoff in the extraglacial area and ice-damming in the Warsaw Basin where a large proglacial lake developed. The Weichselian fluvial sediments form up to three terraces in the valleys of the Vistula and its tributaries. The most contentious issue is the mutual relation of the ice-dammed lake and ice marginal spillways in the Warsaw Basin, both being important fragments of a widespread drainage network in the Central European Lowland.     

Pollen analysis of hyena coprolites and sediments from Equus Cave,Taung, southern Kalahari (South Africa)

Equus Cave, in Quaternary tufa near Taung in the semiarid woodland of the southern Kalahari, yielded 2.5 m of sediment in which a rich assemblage of bones and coprolites was preserved. The fossils were accumulated mainly by hyenas during the late Pleistocene and Holocene. Pollen from coprolites reflects diet as well as vegetation over relatively large areas visited by hyenas, while pollen from sediments represents more local sources. The pollen sequence derived from coprolites and sediments demonstrates how the vegetation evolved from open grassland with small shrubs and occasional trees during the late Pleistocene, to open savanna with more small shurbs, then, during the last 7500 yr, to modern savanna. Temperatures were not more than 4°C cooler and it was slightly moister than today during the late Pleistocene phase; it became gradually warmer but relatively dry before optimal temperature and moisture conditions developed around 7500 yr B.P. Climatic conditions slightly less favorable for woodland occurred during part of the late Holocene.     

Dinosaur coprolites from the Late Cretaceous (Maastrichtian) Lameta Formation of India: isotopic and other markers suggesting a C3plant diet

A single locality of the Late Cretaceous (Maastrichtian) Lameta Formation at Pisdura in Central India has yielded a large number of coprolites attributed to titanosaurian dinosaurs. Internally the coprolites are dark grey and contain abundant plant tissues and other organic materials. The plant tissues are mostly of gymnospermous origin. In addition, remains of bacterial colonies, fungal spores and algae are seen in the macerated fraction under scanning electron microscope. The dark grey appearance is probably attributable to fine-grained organic matter within voids in tracheids or xylem. The average ¹³C/¹²C ratio of the organic matter in the coprolites is -24‰ (relative to PDB) suggesting that plants of C₃type were the main diet of their producers. A comparison of δ¹⁵N value (about 4‰ w.r.t. air) of the coprolites with that of faecal matter of modern herbivores and carnivores suggests that gut fermentation may not have been an active mechanism in the digestion process of titanosaurs.     

银川盆地PL02钻孔孢粉记录的晚上新世-早更新世时期的古气候变化周期

晚上新世-早更新世时期是全球环境发生转折的重要过渡时期。银川盆地位于中国西北内陆地区,同时又处于干旱与半干旱区域的交界地带,在盆地内部蕴藏着丰富的新生代沉积,是研究古环境演化的理想区域,深度为720.77 m的PL02钻孔位于盆地内部的沉降中心附近,属于典型沉降盆地的洪泛沉积环境。本文选取银川盆地PL02钻孔下部(深度705.7~247.9 m)晚上新世-早更新世时期(古地磁年龄3.3~1.5 Ma)的孢粉、磁化率及粒度等指标,通过孢粉分析、频谱分析和小波变换等方法,恢复了银川盆地晚上新世-早更新世时期的古环境演化特征,并对该时期的古气候变化周期进行分析。研究结果显示：银川盆地在晚上新世时期(约2.8 Ma)气候已经开始逐渐变冷变干;进入早更新世时期(约2.6 Ma),研究区气候变得更加干旱并且冷暖波动较大;同时,孢粉、磁化率和粒度等指标均反映出银川盆地晚上新世-早更新世时期古气候变化具有较显著的约20 ka的岁差周期与约40 ka的地轴倾角周期,但约100 ka的地球轨道偏心率周期仅在个别指标中有所表现。另外,本研究结果与中国黄土高原和深海氧同位素记录所揭示的晚上新世-早更新世时期气候变化周期较为一致。因此,认为银川盆地在该时期的气候波动可能受到了地球轨道因素的影响,且与全球环境变化具有相似性。

     

Tephrochronology and tephrostratigraphy of two Pleistocene continental fossiliferous successions from central Italy

The stratigraphical context of two Middle Pleistocene fossiliferous palaeosols from Central Italy (Abruzzo and Tuscany) have been studied. Small mammals and molluscs occur in both palaeosols, which are covered by tephra layers that were analysed using an interdisciplinary approach. Application of fission‐track dating to apatites separated from the Case Picconetto tephra (Pescara, Abruzzo), yielded an age of 0.48 ± 0.04 Ma, indistinguishable from those previously determined for the Campani Quarry (Lower Valdarno, Tuscany) (0.46 ± 0.05 Ma and 0.48 ± 0.05 Ma). Geochemical and petrographic investigations indicate that these tephra originated from different volcanoes, the Alban Hills Volcanic Complex and the Vico Volcano (Latium) respectively. Small mammal and mollusc assemblages indicate different palaeoclimatic and palaeoenvironmental conditions for the Case Picconetto and Campani Quarry palaeosols. Warm and humid conditions can be inferred for the Campani Quarry site, whereas open and cold conditions can be inferred for Case Picconetto. On the basis of faunal data, fission‐track dates and attribution of tephra to specific volcanic eruptions, we suggest a correlation of these faunas with marine oxygen isotope stage 14 (Case Picconetto) and with marine oxygen isotope stage 11 (Campani Quarry), respectively. Copyright © 2003 John Wiley & Sons, Ltd.