Futher reflections on the Isimila Acheulian

Uranium series dates on bone from the Isimila Prehistoric Site, Tanzania, indicate that this East African Acheulian site may be older than originally estimated. They raise the possibility that the Middle Pleistocene sediments containing Acheulian occurrences span a longer range of time than had been supposed. This could add to the plausibility of a contention by Hansen and Keller that differences between artifacts in the uppermost Acheulian horizon and in lower-lying beds are primarily a reflection of change through time rather than a result of different human activities.A summary of the artifact content and associations of the several archeological stratigraphic units is given, and, although a sort of “developmental sequence” can be seen, the existence of discrepant occurrences suggests that an interpretation of directional change as a “function of time” does not, in fact, well accommodate the evidence.Quantitative changes in artifact class proportions in aggregates seem just as likely to be related to local exploitation potentials for early man which were produced during the process of basin in-filling, while observed qualitative differences in the lower-lying horizons may be due to samples biased because of limited exposures which are restricted to a single sedimentary environment.     

Paleorivers and geoarchaeology in the southern Egyptian Sahara

Interpretation of Shuttle Imaging Radar (SIR) images acquired during the November 1981 flight of Columbia led to the idea that a paleodrainage system of regional and perhaps transcontinental proportions crossed southern Egypt and northern Sudan prior to the onset of Quaternary aridity. Three seasons of field investigations in two locales (Wadi Arid and Wadi Safsaf, some 70-80 km apart) near the Sudan border of southern Egypt confirmed initial interpretations of the radar images and produced geologic evidence of fluvial deposition in broad “radar river” valleys (designated RR-1) whose presence is now obscured by eolian sand sheets. Development of an integrated, regional river system probably occurred in the late Paleogene or early Neogene. the river courses eventually were disrupted by tectonism, volcanism, and stream piracy. Their broad valleys were almost fully aggraded long before the middle Pleistocene appearance of man in the area. Some of the large paleovalleys on the radar images contain narrow (0.05-2.0 km wide), braided inset channels (designated RR-2). the RR-2 channels, we believe, represent the last episodes of running water in the valleys during the Quaternary pluvials. In Wadi Arid (the type area for the RR-1 valleys) 36 backhoe trenches and manual excavations yielded Acheulian handaxes, flakes, and cores from nearshore alluvial sediments and surface and shallow subsurface locales. These assemblages are typologically Middle to Late Acheulian and date from ca. 0.15th 0.5 million years ago. the unabraded, unrolled, buried artifacts and their geological contexts indicate that human groups were widely present in a subhumid riparian environment along the edges of the ancient valleys, while these localities were undergoing episodic, local aggradation. Scarce Middle Paleolithic and abundant Neolithic assemblages are widespread along the edges of Wadi Arid and bordering interfluves. In Wadi Safsaf, another broad RR-1 valley, geologic evidence from 20 backhoe excavations demonstrates that an RR-2 channel complex is inset into the alluvial fill in the central part of the valley. No artifacts were found in the alluvium excavated there, but Late Acheulian, Middle Paleolithic, and Neolithic surface assemblages are present nearby. Stratigraphic relations indicate the RR-2 channels are late Pleistocene in age; they were already fully aggraded and their surfaces deflated before an eolian sand sheet of Holocene age was deposited on them. Bir Safsaf, on the northern edge of Wadi Safsaf, was a magnet for Late Acheulian populations, and the paleovalley just south of the bir is bordered with extensive alignments of artifacts, mainly handaxes. Some of these handaxes were found in situ, embedded in alluvial sediments, while others appear to have been exposed on the surface by deflation of the valley fill. The accumulated data support a provisional archaeo-geochronological framework for the last half-million years in the Wadi Arid-Wadi Safsaf area. This framework provides a new geomorphic explanation for the widespread distribution of buried and exposed Middle Acheulian and Late Acheulian assemblages. the interpretation emphasizes the importance of exploring the non-oasis zones for their archaeological and geochronological potential. Although our research has literally only scratched the surface, it demonstrates the potential of radar imaging for defining ancient drainage patterns in arid regions and their associated habitats for human occupation.     

Continual fire-making by Hominins at Gesher Benot Ya‘aqov, Israel

This paper presents the culmination of an extensive study of fire-use at the Early and early Middle Pleistocene site of Gesher Benot Ya‘aqov. Using software available in the GIS package, we have examined the spatial distribution of burned and unburned flint microartifacts from eight Acheulian archaeological horizons. The results of this study demonstrate that the burned microartifacts are never evenly distributed and dense concentrations are observed. The circumstances that introduced these burned flint artifacts to the archaeological layers are examined, suggesting that anthropogenic rather than natural fires are responsible for the observed patterns. As the evidence for the use of fire is recorded throughout the long stratigraphic sequence, it seems that fire was continually used by the Acheulian hominins of the site. This repetitive use of fire indicates that the hominins of Gesher Benot Ya‘aqov had a profound knowledge of fire-making, enabling them to make fire at will.     

The lower paleolithic landscape of 'Ayoun Qedim,al‐Jafr Basin,Jordan

The 'Ayoun Qedim locality includes seven deflated Acheulian sites within and around a limestone box canyon, Juwit el Ghuweir, in the northern al‐Jafr Basin, Jordan. Survey of these sites identified dense scatters of Acheulian bifaces ( _ 1650) along with debris, cores, and reduction material from their production and maintenance. Although there is a clear Levallois component in the assemblages, reflecting a Late Acheulian occupation, strictly Middle and Upper Paleolithic sites are uncommon in this locality. Analysis of the geology and geomorphology of the northern al‐Jafr Basin indicates the area is strongly influenced by shallow subsurface groundwater flow and discharge, and Juwit el Ghuweir displays classic features of a canyon formed by groundwater seepage and sapping, except for the lack of a perennial spring within the modern canyon. The authors suggest that the extreme density of Lower Paleolithic material at 'Ayoun Qedim is associated with perennial groundwater discharge at Juwit el Ghuweir during the Middle Pleistocene and that the dearth of later Paleolithic material is attributable to the cessation of perennial spring discharge. The cause of the spring's cessation is unknown, but may have been related to local geomorphic factors, such as head migration of adjacent streams, or to more regional geomorphic factors, such as the lowering of theregional water table in the area of al‐Jafr Basin. © 2007 Wiley Periodicals, Inc.     

Early and Middle Pleistocene Faunal and hominins dispersals through Southwestern Asia

This review summarizes the paleoecology of the Early and Middle Pleistocene of southwestern Asia, based on both flora and fauna, retrieved from a series of ‘windows’ provided by the excavated sites. The incomplete chrono-stratigraphy of this vast region does not allow to accept the direct chronological correlation between the available sites and events of faunal and hominin dispersals from Africa. It also demonstrates that hominins survived in a mixed landscape of open parkland with forested surrounding hills. In addition, the prevailing environmental conditions are not sufficient to explain successful adaptations to new ecological niches away from the African savanna of the bearers of ‘core and flake’ and the Acheulian industries, The differences in knapping and secondary shaping of stone artifacts probably reflect the learned traditions of different groups of hominins. The current distribution of lithic industries across Eurasia is undoubtedly incomplete due to lack of cultural continuities as well as paucity of field research in several sub-regions. This observation supports the contention that what we view as a constant stream of migrants was actually interrupted many times. The continuous occupation of southwestern Asia by the makers of the Acheulian is in contrast with neighboring regions such as the Iranian plateau and Eastern Europe. A more complex model is required to explain the recorded Eurasian archaeological–cultural mosaic.     

A demographic model for Palaeolithic technological evolution: The case of East Asia and the Movius Line

The Pleistocene record of East Asia continues to pose controversial questions for palaeoanthropology, especially with regard to Palaeolithic technological patterns. In recent years, an increased understanding of the effect of demography on cultural transmission has improved our understanding of the incidence, proliferation, and elaboration of technological traditions. Here, we present a generalised null model of Lower–Middle Palaeolithic technological evolution, which expressly links cultural transmission theory and demographic factors (i.e. population size, density, and social interconnectedness). Consistent with our model, Africa exhibits evidence of major technological innovations during the Early to Middle Pleistocene, due to a constant source of population and growth due to accumulation through time. In comparison, Pleistocene East Asian assemblages are dominated by Mode 1-type technologies, and only a few localized occurrences of bifacial technology are currently known. We detail evidence suggesting that during much of the Pleistocene a combination of biogeographical, topographical, and dispersal factors are likely to have resulted in relatively lower effective population sizes in East Asian hominins compared with western portions of the Old World, particularly Africa. Thus, the Movius Line – as is the case with its namesake ‘Wallace's line’ – must be examined in terms of its biogeographical context, if the divergent evolutionary trajectories of entities either side of it are to be understood. Most parsimoniously, the Movius Line sensu lato is thus a ‘line’ which represents the crossing of a demographic threshold. Under the parameters of our (testable) null model, geographically and temporally sporadic occurrences of bifacial technology in East Asia are the product of short-lived instances of technological convergence. As a consequence, the in situ evolution of Levallois (Mode 3) was inhibited in East Asia due to the constraints of relatively smaller effective population sizes.     

Tephrostratigraphy of the Bedded Tuff Member (Kapthurin Formation, Kenya) and the nature of archaeological change in the later middle Pleistocene

Correlation of volcaniclastic deposits of the Bedded Tuff Member (K4) of the Kapthurin Formation (Kenya) provides the means to assess the nature of archaeological change during the later middle Pleistocene, a time period critical to human evolution but poorly represented at other African localities. Field stratigraphic evidence, and petrographic and electron microprobe geochemical analyses of volcanic glass and phenocrysts, define eight subdivisions of K4 tephra. These include a succession of deposits from a local volcanic source that erupted intermittently, as well as other tuffs likely from different sources outside the Baringo basin. Upper portions of the Bedded Tuff Member date to 235,000 yr. The Bedded Tuff Member is underlain by sediments that include the Grey Tuff, dated to 509,000 ± 9000 yr. The tephrostratigraphic framework defined here is used to place Acheulian and Middle Stone Age (MSA) archaeological sites in chronological order. Results show the persistence of Acheulian large cutting tool manufacture after the advent of points, considered an MSA artifact type. Two assemblages from the site of Koimilot record the appearance at 200,000–250,000 yr of a variety of Levallois flake production methods, an integral if incompletely understood feature of the MSA, here likely derived from local technological antecedents. Combined evidence from the tools and flake production methods suggest an incremental and mosaic pattern of change in hominin adaptive strategies during the Acheulian–MSA transition.     

Structural position of the Pleistocene Gesher Benot Ya'aqov site in the Dead Sea Rift zone

Newly discovered outcrops of the middle Pleistocene Benot Ya'aqov Formation are strongly disturbed due to recent tectonic activity along the Dead Sea Rift. The lacustrine-fluviatile sediments of this formation comprise the littoral facies of a paleo-lake that occupied the adjacent Hula Basin. Acheulian artifacts, found embedded in the formation, have typical African characteristics. The geographical position of the site (the northern extension of the East African Red Sea Rift System) is important for understanding hominid diffusion from Africa to Eurasia.     

Early Human Evolution in the Western Palaearctic: Ecological Scenarios

This review presents the themes of a special issue dealing with environmental scenarios of human evolution during the Early Pleistocene (2.6–0.78 Ma; MIS 103-MIS 19) and early Middle Pleistocene (0.78–0.47 Ma; MIS 19-base of MIS 12) within the western Palaearctic. This period is one of dramatic changes in the climates and the distribution of Palaearctic biota. These changes have played their role in generating adaptive and phyletic patterns within the human ancestry, involving several species such as Homo habilis, “Homo georgicus”, Homo erectus, Homo antecessor and Homo heidelbergensis. In the archaeological record, these species include the Oldowan (Mode 1) and Acheulian (Mode 2) lithic technologies. Taphonomic considerations of palaeoecological research in hominin-bearing sites are provided and evaluated. Syntheses are provided for north Africa, western Asia, the Mediterranean Basin, Britain, and continental Europe. Palaeoenvironmental reconstructions based on multidisciplinary data are given for Ain Boucherit, Ain Hanech and El-Kherba in Algeria, Dmanisi in Georgia, Atapuerca, Cueva Negra, and the Orce Basin in Spain, Monte Poggiolo and Pirro Nord in Italy, Pont-de-Lavaud in France, and Mauer in Germany. The state of the art with the Out of Africa 1 dispersal model is reviewed. A source-sink dynamics model for Palaeolithic Europe is described to explain the morphological disparity of H. heidelbergensis (we will sometimes use the informal name “Heidelbergs”) and early Neanderthals. Other aspects debated here are the selective value of habitat mosaics including reconstructions based on mammal and avian databases, and the role of geological instability combined with topographic complexity. This review is completed by addressing the question of whether the appearance of evolutionary trends within hominins is concentrated in regions of highest worldwide biological diversity (biodiversity hotspots). It is concluded that the keys for the activation of evolutionary change in hominins may have been geological instabilities, and a shifting physiographical heterogeneity combined with high biodiversity and ecological interaction.     

Early and Middle Pleistocene landscapes of eastern England

This paper reviews the pattern of climate and environmental change in eastern England over the period of the Early and Middle Pleistocene, focussing especially upon northern East Anglia. Particular attention is given to the climate and tectonics that have brought about these changes and the distinctive geology, topography and biology that has developed. Throughout, an attempt is made to describe the new models that have been proposed for the Early and Middle Pleistocene of eastern England, and explain the reasons for these changes. The Early Pleistocene experienced relatively high insulation and relatively low magnitude climatic change and is represented primarily by non-climatically forced processes in the form of tidal current- and wave-activity which formed shallow marine deposits. It is possible to recognise a tectonic control in the distribution of deposits of this age because the surface processes do not have the power to remove this signature. The early Middle Pleistocene was dominated by higher magnitude climatic change involving, occasionally, climatic extremes that ranged from permafrost to mediterranean. The landscape at this time was dominated by the behaviour of major rivers (Thames, Bytham, Ancaster) and extensive coastal activity. In the latter part of the early Middle Pleistocene and the Late Middle Pleistocene the climate experienced major changes which resulted in periods of lowland glaciation and short intervals when the climate was warmer than the present. Details of tectonic activity are difficult to identify because they are removed by powerful surface processes, but it is possible to infer uplift focussed on the major interfluves of central England and subsidence in the North Seas basin. In the areas of glaciation the landscape changed radically from an organised terrain dominated by large rivers and extensive shallow coastal zones to complex, with small valleys, disrupted drainage and often discontinuous river, slope and coastal deposits. Likewise the switching off of the North Sea Delta and the opening of the Strait of Dover, separating Britain from continental Europe can be attributed to the onset of lowland glaciation. The case is made that eastern England was glaciated four times during the Middle Pleistocene: during MIS 16, 12, 10 and 6, and attention is given to recent evidence contradicting this model. Over the period of the Middle Pleistocene there is evidence for high biomass production occurring over short intervals coinciding with the climatic optima of MIS 19, 17, 15, 13, 11, and 7c, 7a and during most of these warmer periods, extending back to c. 750 ka (MIS 19/17), there is evidence in the region for the brief appearance of humans.     

Continuous record of environmental changes in Chukotka during the last 350 thousand years

The comprehensive study of the upper 1283 cm of sediment from Lake El’gygytgyn, which formed nearly 4 Ma ago following a meteorite impact in northern Chukotka, yielded the first continuous record of the extreme changes in the Beringian climate and vegetation from the middle Middle Pleistocene to recent time (equivalent of marine isotope stages of 1–7 and the upper part of isotope stage 8). During this period, the climate was warmer than at present between 8600 and 10 7000 ¹⁴C years and during the Late Pleistocene (isotope substage 5e, 116–128 ka ago). In 2003, the German-Russian-USA expedition continued studying sediments of Lake El’gygytgyn to obtain new evidence of the change in the vegetation cover in the Middle Pleistocene and the first information on the Middle Pleistocene interglacial (isotope stage 9; 297–347 ka ago). Pollen spectra characterizing the Middle Pleistocene interglacial are similar to spectra of the early stage of the Early Pleistocene interglacial and the climatic optimum in the Pleistocene to Holocene transitional period. The climatic history of Lake El’gygytgyn is basic for stratigraphic interpretations and correlations in the eastern sector of the Arctic. These data also expand our understanding of climatic changes that are studied within the framework of the “Pole-Equator-Pole Paleoclimate,” “Past Global Changes,” and other international projects.     

旧石器时代东亚、西亚之间的关系

通过实地考察和对收藏标本的细致研究,以获取第一手资料的方式,文章客观地评价了中国古人类学研究者的工作业绩和历史局限,认为在中国发现的东亚早期人类的新证据为探讨人类起源的单极和多极理论提供了新的可能;东亚人的古老性是目前科研工作的关键问题,必须利用各种手段,对此予以尽可能准确的界定。与中国学者合作研究人字洞遗址、龙骨坡遗址和东谷坨遗址的结果说明中国的考古遗址具有多样性,各遗址的石器相互间差异性很大,与同一时期非洲的模式不同,在这里可以找到更加有趣的材料来认识早期人类的方方面面,因此,应以最严谨的态度和最高的注意力对待中国的考古发现。此外,结合中国古人类、旧石器遗址和古气候学的研究资料,文章从宏观地理学的角度阐述了旧石器时代阿舍利和勒瓦娄哇两个典型的文化现象及其发展脉络,探讨了旧石器时代东亚和西亚之间的关系,对中国旧石器文化的发展模式及其与西亚乃至全球的关系提出了个人的独到见解;着眼于研究早期工业技术的发现地、创新地及其传播地点等问题认为,中国同西欧一样,在地理环境和人类生存方面都处于孤立无援的境地,其技术的发展亲历了在本土发明、从境外引进和向外输出等不同阶段,它们不愧是穿越时空来追随技术变革的两大实验场。文章     

Geology and mammalian fauna of the Middle Pleistocene site, Medzhybozh, Ukraine

Qualitative and quantitative faunal associations of molluscs, mammals and other vertebrates (Pisces, Amphibia, Reptilia, Aves) have been determined at a new important site at Medzhybozh, in western Ukraine. Geological and taphonomic conditions of the remains’ preservation are described. The transient (Mindelian–Rissian) fauna is identified and correlated with the Middle Pleistocene Holsteinian (Likhvin) Stage. This sequence represents a warm period of the Holsteinian, followed by with a short cooling in the middle of the period. The human cultural material (of Acheulian and Upper Palaeolithic age) has also been studied at this site. It indicates the oldest migration of humans into the region.     

黄河源区第四纪地质研究的新进展

通过对黄河源区的钻孔、自然露头的研究, 建立了黄河源区的第四纪地层层序。第四纪地层可划分为下更新统、中更新统、上更新统和全新统。下更新统为河湖相沉积; 中更新统主要有湖积物、冰碛物和冰水沉积物; 上更新统主要有湖积物、冰碛物、冰水沉积物、洪积物和河流沉积物; 全新统主要由河流沉积物、洪积物和湖积物构成。黄河源区的冰期可划分为3期, 即末次冰期、倒数第二次冰期、倒数第三次冰期, 末次冰期又可分为2个冰阶。黄河源区的湖泊演化可划分为早更新世、中更新世和晚更新世—全新世3个阶段: 早更新世的湖泊范围小; 中更新世的湖泊范围明显扩大, 在位置上也较早更新世的湖泊南移; 晚更新世的湖泊经历了两次的扩张—收缩变化, 到了全新世, 除现今还发育的几个湖泊外, 大多数地区的湖水已退出, 基本上转变为河流环境。在晚更新世末期到全新世初期, 封闭黄河源区的多石峡被切开, 湖水外泄, 现今的黄河形成了, 同时发生了袭夺长江水系的水流。     

Middle Terrace Deposits of the Tagus River in Alpiarça, Portugal, in Relation to Early Human Occupation

The stratigraphy and sedimentology of the Q3 middle terrace alluvial sequence in the lower Tagus river valley, Portugal, were studied near the village of Alpiarça, approximately 40 km upstream from the estuarine area. Two main stratigraphic units were recognized, separated by an important uncomformity. The Lower Gravels unit (LG) consists of intercalations of medium to coarse gravel deposits, mainly quartzitic, with coarse sandy matrix, organized in tabular bodies. The overlying Upper Sands unit (US) consists of tabular sandy channel deposits and overbank fines, the latter containing well-developed paleosols and backswamp deposits, showing a general aggrading trend, apparently with varying rates; available data indicate that deposition of the US took place under temperate climatic conditions. Within US deposits are several paleolithic archaeological sites, the lower ones in the alluvial stratigraphy being Middle Acheulian, whereas those embedded in overlying deposits are, from bottom to top, Upper Acheulian and Micoquian. Some of these sites have been recently excavated. The quartzite artifacts were apparently abandoned by early humans on the flood plain surface during deposition of the US unit and were subjected to limited reworking during their incorporation in the alluvium. TL/OSL dating of sandy-silty sediments, though imprecise, support archaeological evidence pointing to an age of 150,000 to 70,000 yr B.P. for the US unit.     

重庆地区第四纪气候的初步研究

早更新世时,重庆地区生活着巫山人;中更新世时,生存有盐井沟动物群和歌乐山动物群;晚更新世—全新世时,合川三汇坝动物群繁衍,以水杉、红椿木和枫杨为代表的植物群遍及全区,并有水杉、银杉等孓遗植物存在。整个第四纪时期,重庆地区的气候属亚热带温暖湿润气候,没有发生过冰川活动。     

A late Early Pleistocene pollen record from Fontana Ranuccio (central Italy)

A pollen record from a sediment core drilled underneath the Acheulian site of Fontana Ranuccio provides new data on the vegetation characters in central Italy during the late Early Pleistocene, in a forest phase possibly corresponding to a substage of Marine Isotope Stage 21. A number of tree taxa nowadays extinct in Europe (Cathaya, Tsuga, Taxodium type, Carya and Pterocarya) are present at Fontana Ranuccio in a temperate phase characterised by dense forest conditions, dominated by Alnus and Pterocarya. The comparison of the Fontana Ranuccio record with other dated sites distributed along the Italian peninsula indicates that the floristic and vegetational characters of the Mid Pleistocene revolution present a rather distinct succession of biostratigraphic events. Compared to other Early Pleistocene pollen records from southern Europe, the Italian Peninsula shows greater floristic similarities with the Eastern Mediterranean regions than with the Iberian Peninsula. Copyright © 2011 John Wiley & Sons, Ltd.     

The stratigraphical significance of deer species in the cromer forest-bed formation

Deer species provide a valuable biostratigraphical tool through Cromer Forest-bed times, due particularly to species turnover between the Early and early Middle Pleistocene. This study is based on the reidentification of 348 fossil antlers. The provenance of most large mammal fossils from the CF-bF, collected over 150 years, was recorded only by the nearest coastal village. None the less, analysis of the cervid taxa by these ‘localities’ reveals interesting patterns. The fauna of the West Runton Freshwater Bed is of early Middle Pleistocene complexion, and that from the foreshore at East Runton is of Early Pleistocene. Pure or nearly pure early Middle Pleistocene assemblages also occur at Kessingland-Pakefield and at Trimingham. At Overstrand, Sidestrand, Mundesley, Bacton and Happisburgh, there is a mixture of Early and early Middle Pleistocene elements. Analysis of Savin's data shows that fossils of earlier species were generally found further down the beach than those of later ones. Late nineteenth century geological surveys, made when bone collecting was at its peak, give additional information about fossil horizons, which in several cases can be related to modern stratigraphical units. Most Early Pleistocene large-mammal bones came from Pastonian conglomerates, in contrast to small-mammal and molluscan assemblages mostly extracted from Pre-Pastonian Crag. However, the diversity of Early Pleistocene CF-bF cervid species in comparison with continental faunas, and their pattern of distribution between sites, suggests they may span more than one chronostratigraphic stage. Early Middle Pleistocene assemblages came from strata now referred to the Cromerian, and the differing proportions of taxa between sites provide limited evidence of time-transgression.     

鄂西-三峡地区的古人类资源及相关研究进展

自20世纪50年代发现长阳人化石以来,迄今在鄂西-三峡地区已经发现了至少10处古人类化石地点,时代跨越了整个更新世时期,尤其以更新世中、晚期人类化石最为丰富。除人类化石外,在这一地区还发现了大量的古人类制作的石器、骨器、艺术品、装饰品等人类活动遗存,以及与古人类伴生的动物化石。近年的野外发掘及实验室研究证明鄂西-三峡地区更新世中期以来一直是古人类生存与繁衍昌盛的区域,该地区丰富的古人类资源对研究与中国古人类演化有关的一系列关键学术问题具有非常重要的价值。     

Artifact abrasion,fluvial processes,and “living floors” from the Early Paleolithic site of ’Ubeidiya (Jordan Valley,Israel)

Early Acheulian assemblages in fluviolacustrine contexts at the Early Pleistocene site of ‘Ubeidiya (Jordan Valley, Israel) have been described as “living floors.” A study of variation in the surface abrasion of stone tools from several such “living floors” suggest a mixture of cultural and geological factors were involved in the formation of these assemblages. © 1999 John Wiley & Sons, Inc.