A method to determine warm and cool steppe biomes from pollen data; application to the Mediterranean and Kazakhstan regions

An objective method for the assignment of pollen spectra to appropriate biomes has been published recently. The aim of this paper is to improve the distinction between warm and cool steppes, thus refining vegetation and climate reconstruction, particularly during the Last Glacial Maximum. A set of modern pollen spectra from the Mediterranean and Kazakhstan regions, dominated today by open vegetation types, has been analysed statistically in order to relate pollen taxa abundances to warm and cool grass/shrub plant functional types (PFTs). A statistical test using modern pollen data shows that the method is able to distinguish between cool and warm steppe biomes with a high degree of confidence. The method has been applied to two fossil pollen records. The results of this exercise showed that cool steppe dominated in central Greece between 18 000 and 13 000 yr BP, while in western Iran the vegetation was at the boundary between cool and warm steppes. These vegetation types were replaced by warm mixed forest in Greece and warm steppe in Iran after that time span. © 1998 John Wiley & Sons, Ltd.     

Reconstructing pathways of aeolian pollen transport to the marine sediments along the coastline of SW Africa

The distribution of pollen in marine sediments is used to reconstruct pathways of terrigenous input to the oceans and provides a record of vegetation change on adjacent continents. The wind transport routes of aeolian pollen is comprehensively illustrated by clusters of trajectories. Isobaric, 4-day backward trajectories are calculated using the modelled wind-field of ECHAM3, and are clustered on a seasonal basis to estimate the main pathways of aeolian particles to sites of marine cores in the south-eastern Atlantic. Trajectories and clusters based on the modelled wind-field of the Last Glacial Maximum hardly differ from those of the present-day. Trajectory clusters show three regional, and two seasonal patterns, determining the pathways of aeolian pollen transport into the south-eastern Atlantic ocean. Mainly, transport out of the continent occurs during austral fall and winter, when easterly and south-easterly winds prevail. South of 25°S, winds blow mostly from the west and southwest, and aeolian terrestrial input is very low. Generally, a good latitudinal correspondence exists between the distribution patterns of pollen in marine surface sediments and the occurrence of the source plants on the adjacent continent. The northern Angola Basin receives pollen and spores from the Congolian and Zambezian forests mainly through river discharge. The Zambezian vegetation zone is the main source area for wind-blown pollen in sediments of the Angola Basin, while the semi-desert and desert areas are the main sources for pollen in sediments of the Walvis Basin and on the Walvis Ridge. A transect of six marine pollen records along the south-western African coast indicates considerable changes in the vegetation of southern Africa between glacial and interglacial periods. Important changes in the vegetation are the decline of forests in equatorial Africa and the north of southern Africa and a northward shift of winter rain vegetation along the western escarpment.     

Relationship between pollen assemblages in surface soil and modern vegetation and climate in the western Tianshan Mountains,Xinjiang北大核心CSCD

Based on the pollen spectrum characteristics of 46 surface samples collected in the western Tianshan Mountains of Xinjiang and a plant community quadrat survey, we analyzed the relationship between pollen assemblages and modern climate and vegetation distribution in the region. The results showed the surface pollen assemblages were consistent with the distribution of modern vegetation. Based on the results of cluster analysis, the study area could be divided into three pollen assemblage zones（mountain steppe zone, desert vegetation zone and typical desert zone）. In general, pollen assemblage can roughly reflect the main characteristics of local vegetation, but there are significant differences in the percentage of some pollen types and the coverage of related plants. The pollen of Picea and Pinus in arbor plants, Ephedra in shrub plants, Chenopodiaceae and Artemisia in herb plants were high represented due to the influence of natural wind, water flow and their own pollen yield. While Rosaceae pollen in shrubs and Poaceae in herbs were less representative in a modern plant community with its own dominant species;The Artemisia and Chenopodiaceae（A/C）ratio can distinguish the desert zone from the mountain steppe zone, and could clearly reflect the aridity level. However, when using this indicator to reconstruct the climate environment, it is necessary to combine the characteristics of pollen assemblage and the influence of changes in vegetation composition and other factors to distinguish;Altitude, average annual precipitation, and average annual temperature all affect the distribution of surface pollen in the area. © 2022 The authors.     

川北若尔盖高原玛曲-红原一带表土中孢粉组合的特征

通过对四川若尔盖高原玛曲—红原一带20个表土孢粉样的分析,研究了该区域表土孢粉组合与现代植被的关系。结果显示,玛曲—红原一带表土的孢粉组合基本反映了现代植被的整体特征,与现代植被在植被类型、植被带的空间排列、主要建群种属和优势种属等方面基本一致。依据孢粉的百分含量,可以划分为亚高山灌丛植被区,亚高山灌丛、亚高山草甸植被区和高寒草甸、草原植被区3个植被带。表土花粉的代表性因种属的不同而各有差异,松属和菊科花粉具超代表性,云杉属花粉的代表性较好,禾本科和莎草科具低代表性,而桦木属则不具有超代表性,可能与桦木属花粉不易搬运有关。3个植被区中的表土孢粉的种类、比例与现代植被均存在一定的差别,这可能与植被退化有关。     

Vegetation in western Central Mexico during the last 50 000 years: modern analogs and climate in the Zacapu Basin

The late glacial and the transition towards the Holocene marked a period of significant environmental change at a global scale. In western Central Mexico, few records span beyond the Holocene, and little is known about Pleistocene climatic and environmental variability. Here we report on the pollen record of a composite sequence made of three cores that cover the last 50 ka (thousands of calibrated radiocarbon years before present) in the Zacapu Basin, western Central Mexico (～1970 m asl). The conjunction of modern pollen rain from the area with the composite fossil pollen sequence from cores Cantabria, Cantabria 1 and CEMCA Point‐1 provided a detailed history of the vegetation of the area. Recognition of modern environmental patterns through individual modern pollen taxa proved difficult, but multivariate analysis separated different vegetation types. No‐modern‐analog communities relative to the modern vegetation cover of the area were evident, especially during periods of drought. Apparently, summer precipitation was significant before 40 ka and after the deglaciation. However, this moisture source declined from ～40 to 10 ka because tradewinds from the Gulf of Mexico were blocked by the eastern mountains of Central Mexico as a result of lower temperatures. By contrast, westerly and northerly winds were enhanced by maximum ice cover from ～19 to 17 ka, providing abundant winter precipitation and therefore preventing extreme drought in the area. Copyright © 2012 John Wiley & Sons, Ltd.     

Lake Neor reveals how mountain vegetation responded to 7000 years of hydroclimate variability in northwestern Iran

Palynological and geochemical analyses provide valuable information about modern and past climatic regimes and vegetation. The impact of climate and humans on past vegetation in the semi-arid areas of northwestern Iran has received increased interest in the wake of warming temperatures in the Middle East. Palynological and down-core XRF elemental abundances from a peat core from Lake Neor enabled a reconstruction of vegetational changes of the past 7000 years over the highlands of northwestern Iran. Periods of increased arboreal pollen (AP) types and high (Artemisia + Poaceae)/Chenopodiaceae ratios along with low titanium abundances, high percentages of total organic carbon, more negative δD values, and higher carbon accumulation rates suggest a relatively wet climate. These conditions have persisted during the periods 6700–6200, 5200–4450 and 3200–2200 cal a bp. The overall low AP values, substantial rise of Chenopodiaceae, high Ti abundances and low values of palaeo-redox proxies, are all evidences of a drier climate, as has been reconstructed for the periods 6200–5200 and 4030–3150 cal a bp and the last 2200 years. An important feature of the last centuries is the increase of anthropogenic and pastoral indicator pollen types. Our results may provide basic data to predict future trends in vegetation dynamics under future climate change in western Asia.     

Modern pollen assemblages from surface lake sediments and their environmental implications on the southwestern Tibetan Plateau

The relationships amongst modern pollen assemblages, vegetation, climate and human activity are the basis for reconstructing palaeoenvironmental changes using pollen records. It is important to determine these relationships at regional scales due to the development of vegetation under different climatic conditions and human activities. In this paper, we report on an analysis of modern pollen assemblages of 31 surface lake samples from 31 lakes (one sample per lake) on the southwestern Tibetan Plateau where the knowledge of modern pollen and their relationships with vegetation, climate and human activities is insufficient. The region includes five vegetation zones: sub‐alpine shrub steppe, alpine steppe, alpine meadow and steppe ecotone, mountain desert and alpine desert. The lakes span a wide range of mean annual precipitation (50–500 mm) and mean annual temperature (?8 to 6 °C). Modern pollen assemblages from our samples mainly consist of herb taxa (Artemisia, Cyperaceae, Poaceae, Chenopodiaceae, etc.) and some tree taxa (Pinus, Fagaceae, Alnus, etc.). The results indicate that modern pollen assemblages are able to reflect the main vegetation distribution. Redundancy analysis for the main pollen types and environmental variables shows that precipitation is the leading factor that influences the pollen distribution in the study area with the first axis capturing 13.7% of the variance in the pollen data set. The Artemisia/Chenopodiaceae ratio is valid for separating the desert component (<2) from the steppe and other vegetation zones (>2) but is unable to distinguish moisture variations. The Artemisia/Cyperaceae ratio is able to identify meadows (<1) and steppes (>1) and can be used as a moisture index on the southwestern Tibetan Plateau. Our results show that an appropriate range is needed for a modern pollen data set in order to perform pollen‐based quantitative climate reconstructions in one region. It is essential to perform modern studies before using pollen ratios to reconstruct palaeovegetation and palaeoclimate at a regional scale.     

青藏高原现代表土中冷杉和云杉花粉的空间分布

文章通过对青藏高原北纬25°～45°,东经75°～106°范围内海拔1000～5700m的地区所采集的598个表土样品的孢粉分析,研究了现代表土中冷杉和云杉花粉的分布与植被、气候和海拔之间的关系。表土样品中冷杉和云杉花粉高含量范围对应着青藏高原南部和东部山地暗针叶林分布区,花粉等值线图较准确地反映了冷杉和云杉的丰富程度及其与植被、环境的关系;冷杉和云杉花粉百分含量随海拔和气候梯度变化表现出单峰分布模式,冷杉和云杉花粉百分含量大于20%的最适宜海拔高度是2500～4000m,最高含量中心限制在海拔3200m左右、年均温2～3℃范围内;较高的花粉百分含量带出现在年均温0～8℃、年均降水量400～850mm的区域。结果显示了青藏高原现代冷杉和云杉花粉的空间分布与植被、气候、海拔的空间变化之间很好的相关性,为利用花粉资料进行青藏高原地区的古气候重建提供了有益的资料     

The importance of plant macrofossils in the reconstruction of Lateglacial vegetation and climate: examples from Scotland,western Norway,and Minnesota,USA

Lateglacial and early Holocene (ca 14–9000 ¹⁴C yr BP; 15–10,000 cal yr BP) pollen records are used to make vegetation and climate reconstructions that are the basis for inferring mechanisms of past climate change and for validating palaeoclimate model simulations. Therefore, it is important that reconstructions from pollen data are realistic and reliable. Two examples of the need for independent validation of pollen interpretations are considered here. First, Lateglacial-interstadial Betula pollen records in northern Scotland and western Norway have been interpreted frequently as reflecting the presence of tree-birch that has strongly influenced the resulting climate reconstructions. However, no associated tree-birch macrofossils have been found so far, and the local dwarf-shrub or open vegetation reconstructed from macrofossil evidence indicates climates too cold for tree-birch establishment. The low local pollen production resulted in the misleadingly high percentage representation of long-distance tree-birch pollen. Second, in the Minnesotan Lateglacial Picea zone, low pollen percentages from thermophilous deciduous trees could derive either from local occurrences of the tree taxa in the Picea/Larix forest or from long-distance dispersal from areas further south. The regionally consistent occurrence of low pollen percentages, even in sites with local tundra vegetation, and the lack of any corresponding macrofossil records support the hypothesis that the trees were not locally present. Macrofossils in the Picea zone represent tundra vegetation or Picea/Larix forest associated with typically boreal taxa, suggesting it was too cold for most thermophilous deciduous trees to grow. Any long-distance tree pollen is not masked by the low pollen production of tundra and Picea and Larix and therefore it is registered relatively strongly in the percentage pollen spectra.Many Lateglacial pollen assemblages have no recognisable modern analogues and contain high representations of well-dispersed ‘indicator’ taxa such as Betula or Artemisia. The spectra could have been derived from vegetation types that do not occur today, perhaps responding to the different climate that resulted from the different balance of climate forcing functions then. However, the available contemporaneous plant-macrofossil assemblages can be readily interpreted in terms of modern vegetation communities, suggesting that the pollen assemblages could have been influenced by mixing of locally produced pollen with long-distance pollen from remote vegetation types that are then over-represented in situations with low local pollen production. In such situations, it is important to validate the climate reconstructions made from the pollen data with a macrofossil record.     

Connection between South Mediterranean climate and North African atmospheric circulation during the last 50,000 yr BP North Atlantic cold events

High-resolution clay mineralogical analyses were performed on sediment deposited during the last 50,000 yr in the Alboran sea (ODP Site 976). The clay mineral record is compared with pollen assemblages and with annual precipitation (Pann) and mean temperatures of the coldest month (MTCO) reconstructed with the modern analog technique (MAT). Enhanced contribution of palygorskite, a typical wind-blown clay mineral, characterizes the North Atlantic cold climatic events. Coeval development of the semi-arid vegetation (Artemisia rich) associated with a drastic fall of reconstructed precipitations and temperatures, suggest cold and arid continental conditions in the West Mediterranean area during North Atlantic cold events. The clay mineral association, especially the palygorskite content and the illite-to-kaolinite ratio, indicate western Morocco as one of the major source of the clay-size fraction during the North Atlantic cold events. The maximum abundance of Artemisia associated with the presence of Argania pollen both indicate Morocco as the main origin for pollen during these cold periods. The comparison of these pollen and clay mineral-specific features allows us to pinpoint western Morocco as the dominant source of wind-blown particles during North Atlantic cold events. These specific mineralogical composition and palynological assemblages reveal enhanced aridity over North Africa and intensification of winds favouring dust erosion and transport from North Africa toward the Alboran Sea during the North Atlantic cold events. According to atmospheric models, such a meridian transport (1) likely results from the development of strong and stable anticyclonic conditions over the tropical Atlantic and North Africa, similar to today's summer meteorological configuration and (2) implies a northward position of the westerly winds during North Atlantic cold events. Finally the synoptic situation over the West Mediterranean during the North Atlantic cold events is compared with the North Atlantic Oscillation (NAO), suggesting that during the cold Atlantic events, weather regimes over Europe and North Africa may have been systematically shifted towards a positive NAO situation.     

Modelling the vegetation response to the 8.2 ka bp cooling event in Europe and Northern Africa

The 8.2 ka bp cooling event is assumed to be the most clearly marked abrupt climate event in the Holocene at northern mid‐ to high latitudes. In this study, we simulate the vegetation responses to the 8.2 ka bp climate change event over Europe and Northern Africa. Our results show that all dominant plant functional types (PFTs) over Europe and North Africa respond to these climate changes, but the magnitude, timing and impact factor of their responses are different. Compared with pollen‐based vegetation reconstructions, our simulation generally captures the main features of vegetation responses to the 8.2 ka bp event. Interestingly, in Western Europe, the simulated vegetation after perturbation is different from its initial state, which is consistent with two high‐resolution pollen records. This different vegetation composition indicates the long‐lasting impact of abrupt climate change on vegetation through eco‐physiological and ecosystem demographic processes, such as plant competition. Moreover, our simulations suggest a latitudinal gradient in the magnitude of the event, with more pronounced vegetation responses to the severe cooling in the north and weaker responses to less severe cooling in the south. This effect is not seen in pollen records. © 2019 The Authors. Journal of Quaternary Science Published by John Wiley & Sons, Ltd.     

Late Pleistocene vegetation of Kings Canyon,Sierra Nevada,California

Seven packrat midden samples make possible a comparison between the modern and late Pleistocene vegetation in Kings Canyon on the western side of the southern Sierra Nevada. One modern sample contains macrofossils and pollen derived from the present-day oak-chaparral vegetation. Macrofossils from the six late Pleistocene samples record a mixed coniferous forest dominated by the xerophytic conifers Juniperus occidentalis, Pinus cf. ponderosa, and P. monophylla. The pollen spectra of these Pleistocene middens are dominated by Pinus sp., Taxodiaceae-Cupressaceae-Taxaceae (TCT), and Artemisia sp. Mesophytic conifers are represented by low macrofossil concentrations. Sequoiadendron giganteum is represented by a few pollen grains in the full glacial. Edaphic control and snow dispersal are the most likely causes of these mixed assemblages.The dominant macrofossils record a more xeric plant community than those that now occur on similar substrates at higher elevations or latitudes in the Sierra Nevada. These assemblages suggest that late Wisconsin climates were cold with mean annual precipitation not necessarily greater than modern values. This conclusion supports a model of low summer ablation allowing for the persistence of the glaciers at higher elevations during the late Wisconsin. The records in these middens also suggest that S. giganteum grew at lower elevations along the western side of the range and that P. monophylla was more widely distributed in cismontane California during the Pleistocene.     

Pollen assemblages and their environmental implications in the Qaidam Basin,NW China

The Qaidam Basin is one of the most sensitive areas to climate change in China, owing to its unique geographical position and ecological condition. In this study, 32 surface‐soil pollen samples were collected to reveal the relationship between modern pollen assemblages, vegetation and precipitation in the eastern region of the Qaidam Basin. The results show that Chenopodiaceae (3.8–87%, average 48%), Artemisia (1.7–64.2%, average 17.5%) and Ephedra (0–90%, average 16.3%) are the dominant pollen types in all samples, and that different pollen assemblages correspond to different vegetation types. DCA and CCA of major pollen types demonstrate that precipitation is an important factor in the control of the distribution of vegetation in the study area. The content and concentration of the three major pollen types (Artemisia, Chenopodiaceae and Ephedra) change with the mean annual precipitation, and the optimum mean annual precipitation for Ephedra, Chenopodiaceae and Artemisia is <80, 80–200 and >160 mm, respectively. Correlation analysis between the variation in grain size of the three major pollen types and the main environmental variables shows that the grain size of the three pollen types is positively correlated with precipitation in the Qaidam Basin. The results confirm that precipitation is the most important environmental factor in the Qaidam Basin, and that it has an important effect on pollen grain size in the study area.     

Pollen taphonomy in a canyon stream

Surface soil samples from the forested Chuska Mountains to the arid steppe of the Chinle Valley, Northeastern Arizona, show close correlation between modern pollen rain and vegetation. In contrast, modern alluvium is dominated by Pinus pollen throughout the canyon; it reflects neither the surrounding floodplain nor plateau vegetation. Pollen in surface soils is deposited by wind; pollen grains in alluvium are deposited by a stream as sedimentary particles. Clay-size particles correlate significantly with Pinus, Quercus, and Populus pollen. These pollen types settle, as clay does, in slack water. Chenopodiaceae-Amaranthus, Artemisia, other Tubuliflorae, and indeterminate pollen types correlate with sand-size particles, and are deposited by more turbulent water. Fluctuating pollen frequencies in alluvial deposits are related to sedimentology and do not reflect the local or regional vegetation where the sediments were deposited. Alluvial pollen is unreliable for reconstruction of paleoenvironments.     

重庆中梁山岩溶槽谷区表土孢粉与现代植被的关系研究

本文选取重庆中梁山岩溶槽谷区的表土和苔藓样品做孢粉分析,探讨其与现代植被的关系,并初步推断中梁山岩溶洼地的花粉来源,为其洼地剖面花粉来源提供参考。结果表明:表土花粉组合与现代植被类型具有较好的对应关系,耕地、撂荒地、灌木丛、马尾松林以及竹林能够较好地反映母体植被的群落组成;混交林与母体植被群落对应关系较差。由于洼地对区域花粉的汇聚作用,其表土花粉谱更能反映整个区域的植被状况。洼地表层样品中松属花粉来源是中梁山地区与西南风区域的花粉;落叶栎属、桤木属、枫杨属、葎草属等的主要来源是周围山地;禾本科花粉来源相对复杂。本研究可为在这一地区利用孢粉学开展第四纪古植被、古气候和古环境等研究提供参考性依据。      

Vegetation patterns, recent pollen deposition and distribution of non-pollen palynomorphs in a polygon mire near Chokurdakh (NE Yakutia, NE Siberia)

Fine-scaled distribution patterns of vegetation and palynomorphs in a polygon mire in NE Siberia were studied in 31 × 31 plots of 1 m² (vegetation) and along a transect at intervals of 1 m (palynomorphs). Pollen and non-pollen-palynomorph contents clearly reflect differences between polygon ridges and centres. For most single pollen types, however, no unambiguous relation with actual plant abundances and distribution exists. High pollen values are restricted to the immediate vicinity of the pollen sources, which is attributed to the low height of the vegetation. For several plant taxa, high pollen values were found to be restricted to particular plant specimens in the population. High values of pollen ascribed to taxa currently absent at the relevant plot show that the sampled mosses contain a pollen signal of several years. Betula exilis produces two morphologically clearly distinct pollen types. The distribution of selected non-pollen palynomorphs correlates to microrelief, moisture regime and/or trophic conditions. The study shows that single palaeo-sections from arctic peatlands should not be over-interpreted with respect to their palaeoecological value. Local vegetation types and site conditions, however, can be accurately reconstructed from the total assemblages and from combined occurrences of specific palynomorph types.     

Vegetation,fire, and climate history of the northwestern Great Basin during the last 14,000 years

The northwestern Great Basin lies in the transition zone between the mesic Pacific Northwest and xeric intermountain West. The paleoenvironmental history based on pollen, macroscopic charcoal, and plant macrofossils from three sites in the northwestern Great Basin was examined to understand the relationships among the modern vegetation, fire disturbance and climate. The vegetation history suggests that steppe and open forest communities were present at high elevations from ca 11,000 to 7000 cal yr BP, and were replaced by forests composed of white fir, western white pine, and whitebark pine in the late Holocene. Over the last 11,000 years, fires were more frequent in mid-elevation forests (10–25 fire episodes/1000 years) and rare in high-elevation forests (2–5 fire episodes/1000 years). Applying modern pollen–climate relationships to the fossil pollen spectra provided a means to interpret past climate changes in this region. In the past 9000 years summer temperatures decreased from 1 to 4 °C, and annual precipitation has increased 7–15%. These results indicate that the millennial-scale climate forcing driving vegetation changes can be quantified within the intermountain West in general and northwestern Great Basin in particular. In addition, fire can be considered an important component of these ecosystems, but it does not appear to be a forcing mechanism for vegetation change at the resolution of these records.     

川北若尔盖高原表土孢粉的特征及其与现代植被的关系

通过对若尔盖高原不同植被带34个表土样的孢粉分析,研究了各植被带的表土孢粉组合与现代植被的关系。结果显示：表土孢粉组合基本上能够反映出现代植被的整体特征,与现代植被带在空间分布、建群种和优势种属方面基本相同,但不完全等于植被组成,影响其差异的原因可能是人类活动、孢粉保存、孢粉产量、外来孢粉和孢粉鉴定。表土花粉的代表性因种属的不同而各有差异。乔木花粉中,松属具有超代表性,云杉属代表性较好,而桦木属不具有超代表性;草本花粉中,菊科具较好的代表性,蒿属和藜科等具代表性,而禾本科和莎草科具低代表性;蕨类孢子具等代表性。     

柴达木盆地东部36～18kaB.P.期间的孢粉记录及其气候环境

对柴达木盆地察尔汗古贝壳堤剖面的沉积物进行孢粉分析,结果显示这个地区植被与气候在晚更新世中晚期经历以下几个阶段的变化:36.2～31.2kaB.P.(未校正14C年代,下同)期间,以禾本科、藜科、蒿属、莎草科为主,发育草原-草甸植被,气候温和湿润,盘星藻出现较多,反映淡水湖泊,水深在10m左右;31.2～27.6kaB.P.期间,松属、云杉属、桦属等为主的木本植物的含量增加,周围山地森林发育,表明气候温暖,降水量增多。但由于蒸发量大,有效湿度下降,荒漠成分柽柳属等增加,盘星藻在30kaB.P.以后消失,反映湖泊盐度增大;27.6～23.3kaB.P.期间,植被中荒漠成分显著增加,周围山地森林萎缩,气候趋向相对寒冷干旱,湖面积缩小;23.3～18.0kaB.P.,孢粉浓度很低,蒺藜科、藜科等荒漠成分明显增加,植被稀疏,已趋向荒漠化草原,反映气候寒冷干旱。从整个剖面来看,主要的陆生植物孢粉类型为禾本科、柽柳属、蒺藜科、麻黄属、松属、云杉属、柏科、胡桃属和桦属等,藜科和蒿属含量很少,这与柴达木盆地东部地区的表土分析结果完全不同,也与其他草原以及荒漠草原的表土花粉结果相异。这说明晚更新世中晚期柴达木盆地东部地区的植被和现在无法进行比较,气候环境与现在显著不同。