Anak Krakatau — a colonization model within a colonization model?

The ppaer re-examines three suggestions previously made concerning the colonization of the Krakatau islands since the extirpating 1883 eruption that involve the more recently emergent volcanic island Anak Krakatau, which itself suffered a devastating eruption in 1952. The suggestions re-addressed in the light of recent comments by other workers are:

Emplacement of a Debris Avalanche during the 1883 eruption of Krakatau (Sunda Straits,Indonesia)

Conclusion The data collected during the Mentawai cruise help to clarify understanding of the 1883 eruption of Krakatau. We have previously discussed the weaknesses of the interpretation of Williams (1941) and others (Self and Rampino 1981) and emphasized that only a Mount St. Helens-type collapse during the course of the eruption could account for all the characteristics of the eruption and of the related deposits.The discovery on land of deposits attributable to a debris-avalanche, in the stratigraphic position where they were expected, is a strong argument for the validity of our scenario.Marine surveys confirm that the sea bottom around Krakatau is covered by a thick ignimbritic deposit. But the presence of this deposit does not invalidate the presence of a debris-avalanche deposit under the ignimbrites. The hummocky morphology favours this hypothesis.Flank-failure of volcanoes is generally considered as a very efficient mechanism for triggering tsunamis (Kienle et al. 1987; Siebert et al. 1987). However, the majority of the volcanoes where flank-failure has been described are tall and bulky and the collapse of a broad edifice like Krakatau may be surprising. However the geological evidence shows that such a mechanism can act at various scales; for example the flank collapse of Mayu Yama volcano (height 700 m, volume 0,3 km³), a parasitic cone of Unzen volcano (Japan), triggered a debris-avalanche into the sea that was 1 km long, with a characteristic hummocky surface; the resulting tsunami killed 9528 people (Katayama 1974). In the same way, a partial collapse of Iliwerung volcano, Indonesia (50 × 10⁶ m³) in July 1979, triggered a tsunami which killed several hundred people (McClelland et al. 1989). At Krakatau, the main summit was 822 m asl; the collapse took place along the edge of the prehistoric caldera and this structural unconformity probably facilitated the triggering of the process.     

Anak Krakatau's vegetation and flora circa 1991, with observations on a decade of development and change

Ecological surveys were conducted over the period 1989 to 1991 on the island of Anak Krakatau. The aims of the study were to detail a key phase of rapid change in the vegetation succession and plant colonization of this young volcano and to compare results with those from earlier studies.Floristic composition was determined by reconaissance of all vegetated areas during the period 1989 to 1991. Colonization has been rapid since 1983 with 53 species newly recorded for Anak Krakatau, comprising 8 Pteridophyta and 45 Angiospermae. Of the latter, a significant element (13) were of species believed to be human-dispersed. Comparison of all floral surveys has revealed a core of strand-line and pioneer colonists which has been common to each restart of colonization. The sea-dispersed component is already comparable in size to that of Rakata and is thought unlikely to increase significantly from this point. The accumulation of animal-dispersed species of interior habitats has only recently begun to pick up speed.Vegetation was characterised by means of plot-based sampling, including the establishment of three permanent plots, and by mapping at the whole-island scale. Variation in composition within the island is discussed, with particular reference to the three vegetated forelands. Anak Krakatau now supports the most extensive and advanced vegetation in its disrupted 61-year history. A more or less closed cover extends along the eastern/north-eastern coast-line, but the xeric interior remains only sparsely vegetated. It is concluded that because of volcanic disturbance a base-line for ecosystem development on Anak Krakatau cannot be calculated but that the pattern of change is not inconsistent with significant disturbance ca. 1972 and the subsequent relaxation of these peculiar environmental constraints.     

Middle Weichselian environments on western Yamal Peninsula, Kara Sea based on pollen records

Pollen data from two sections from a coastal cliff on the western Yamal Peninsula (69°43.27′N, 66°48.80′E) document the environmental history during the Karginsky (Middle Weichselian) interstadial. Low pollen concentrations, high amounts of redeposited pollen, and relatively high presence of Artemisia pollen characterize sediments deposited at about 33,000 ¹⁴C yr B.P. Grass-sedge plant associations with few other herbs occupied the area during the late Karginsky interstadial. Artemisia pollen may indicate rather xerophytic vegetation and disturbed soils in the area. The dominance of redeposited pollen reflects scarce (disturbed) vegetation cover and low pollen productivity. The climate was relatively cold and dry. Sediments dated to 32,400 ¹⁴C yr B.P. contain fewer redeposited pollen and concentration of non-redeposited pollen is significantly higher. Pollen contents indicate the dominance of tundra-like grass-sedge vegetation and more humid conditions. Pollen records dated between 30,100 and 25,100 ¹⁴C yr B.P. also reflect scarce tundra-like vegetation during this interval. The presence of Betula nana and Salix pollen may reflect limited presence of shrub communities. This suggests that the climate was somewhat warmer during the latter part of the interstadial. However, generally the pollen records show that harsh environmental conditions prevailed on the Yamal Peninsula during the Karginsky interstadial.     

Late Quaternary vegetation reconstruction from the Eastern Arc Mountains, Tanzania

Pollen, spore, macrofossil and stable isotope (C and N) analyses from a 266-cm sediment core collected from a swamp on the Eastern Arc Mountains, Tanzania, are used to reconstruct vegetation and environmental history. An estimated time scale based on five ¹⁴C ages records approximately 38,000 yr. This palaeorecord is the first from this biodiversity hotspot and importantly extends through the last glacial maximum (LGM). The altitudinal transition from montane to upper montane forest shifted from 1700-1800 m (38,000 ¹⁴C yr BP) to 1800-1900 m (35,000-29,000 ¹⁴C yr BP). From 29,000 to 10,000 ¹⁴C yr BP, it shifted from 1850-1950 m across the LGM to 1750-1800 m (during 10,000-3500 ¹⁴C yr BP), and to present-day elevations at 2000 m during the last 3500 ¹⁴C yr BP. The relative ecosystem stability across the LGM may be explained by the Indian Ocean's influence in maintaining continuous moist forest cover during a period of East African regional climate aridity. During the late Holocene, presence of abundant coprophilous fungi and algal blooms demonstrates increasing human impact. Neurospora spores indicate frequent fires, coinciding with clear signals of decline in Podocarpus and Psychotria trees that possibly represent selective logging.     

Drastic desalination of small lakes in East Siberia (Russia) in the early twentieth century: inferred from sedimentological, geochemical and palynological composition of small lakes

Reconstruction of temporal and spatial climate development on a seasonal basis during the last few centuries may help us better understand modern-day interplay between natural and anthropogenic climate variability. The objective of this paper is to reconstruct hydrology and landscape changes of East Siberia during the termination of the Little Ice Age and the subsequent Recent Warming. We analysed sediment samples from the saltwater Sulfatnoe Lake, Bolshoye Alginskoe and freshwater Shuchie Lake using high-resolution X-ray fluorescence spectroscopy at 1-mm scan resolution, Fourier-transform infrared techniques and pollen analyses. The depth–age models of the cores were constructed by ²¹⁰Pb activity using the constant rate of supply model. The lake sediment cover of these lakes began to form from ca. 1870. Three significant periods (1870–1895, 1895–1925 and from 1925 to the present) were defined in hydrology and chemical regime of these lakes for the past 140 years. Lake levels were extremely low and high saturated with salts during the final period of the Little Ice Age. Lake levels began to slowly rise from 1870 to 1895 and vegetation was poor at that period. Intensive desalination of the lakes occurred in 1895–1925, and environment conditions were temperate and favourable for the majority of the taxa of the regional vegetation. Regional precipitation significantly increased and water saturation of the catchments was high from 1925 to the present. The chemical precipitation of carbonate stopped completely in Lake Shichie and reduced considerably in Lake Sulfatnoe and B. Alginskoe. Strong increasing trend of weathering of the lake catchments began in 1970 and still continues.     

A methodological framework for assessing and reducing temporal uncertainty in paleovegetation mapping from late-Quaternary pollen records

Mapping past vegetation dynamics from heterogeneous databases of fossil-pollen records must face the challenge of temporal uncertainty. The growing collection of densely sampled fossil-pollen records with accurate and precise chronologies allows us to develop new methods to assess and reduce this uncertainty. Here, we test our methods in the context of vegetation changes in eastern North America during the abrupt climate changes of the last deglaciation. We use the network of fossil-pollen records in the Neotoma Paleoecology Database (www.neotomadb.org) and data contributed by individual investigators. Because many of these records were collected decades before the current generation of ¹⁴C and age-model technologies, we first developed a framework to assess the overall reliability of ¹⁴C chronologies by systematically evaluating individual ¹⁴C ages and associated chronologies. We developed a qualitative ranking scheme for individual ¹⁴C ages that combines information about their accuracy and precision. ‘Benchmark’ pollen records were defined to have at least one ¹⁴C age with an accuracy within 250 years and a precision less than 500 years that is within 1000 years of the time interval of interest, and at least five pollen samples per 1000 years across this time period. Only 22 of >350 late-Pleistocene pollen cores in eastern North America met the benchmark criteria.We then used Bayesian change-point analysis to identify widespread ecological events (Picea decline, Quercus rise, and Alnus decline), and interpolated the ages of these events from the benchmark sites to non-benchmark sites. Leave-one-out cross-validation analyses with the benchmark sites indicated that the spatial error associated with interpolation was less for inverse distance-weighting (IDW) than thin-plate splines (TPS) and was about 500 years for the three biotic events. By comparison, the difference between the original ages of events at poorly constrained sites and the biostratigraphic ages interpolated from the benchmark sites was close to 1000 years, suggesting that the use of biostratigraphic ages can significantly improve the age models for poorly constrained sites. Overall, these analyses suggest that the temporal resolution of multi-site syntheses of late-Pleistocene fossil-pollen data in eastern North America is about 500 years, a resolution that allows analysis of ecological responses to millennial-scale climate change during the last deglaciation.     

Multiproxy record for the last 4500 years from Lake Shkodra (Albania/Montenegro)

A multi‐proxy record is presented for approximately the last 4500 cal a BP from Lake Shkodra, Albania/Montenegro. Lithological analyses, C/N ratio and δ¹³C of the organic and inorganic carbon component suggest that organic matter and bulk carbonate are predominantly authigenic. The δ¹⁸O record of bulk carbonate indicates the presence of two prominent wet periods: one at ca. 4300 cal a BP and one at ca. 2500–2000 cal a BP. The latter phase is also found in southern Spain and Central Italy, and represents a prominent event in the western and central Mediterranean. In the last 2000 years, four relatively wet intervals occurred between ca. 1800 and 1500 cal a BP (150–450 AD), 1350–1250 (600–700 AD), 1100–800 (850–1150 AD), and at ca. 90 cal a BP (1860 AD). Between ca. 4100 and 2500 cal a BP δ¹⁸O values are relatively high, with three prominent peaks indicating drier conditions at ca. 4100–4000 cal a BP, ca. 3500 and at ca. 3300 cal a BP. Four additional drier events are identified at 1850 (ca. 100 AD), 1400 (ca. 550 AD), 1150 (800 AD) and ca.750 cal a BP (1200 AD). The pollen record does not show changes in accordance with these episodes owing to the poor sensitivity of vegetation in this area, which is dominated by an orographic rainfall effect and where changes in altitudinal vegetation belts do not affect the pollen rain in the lake catchment. However, since ca. 900 cal a BP a significant decrease in the percentage arboreal pollen and in pollen concentrations suggest major deforestation produced by human activities. Copyright © 2012 John Wiley & Sons, Ltd.     

Late Holocene environmental reconstruction using cave sediments from Belize

Cave sediments collected from Reflection Cave on the Vaca Plateau, Belize show variations in the δ¹³C values of their fulvic acids (FAs), which indicate periods of vegetation change caused by climatic and Maya influences during the late Holocene. The δ¹³C values range from − 27.11‰ to − 21.52‰, a shift of ∼ 5.59‰, which suggests fluctuating contributions of C₃ and C₄ plants throughout the last 2.5 ka, with C₄ plant input reflecting periods of Maya agriculture. Maya activity in the study area occurred at different intensities from ∼ 2600 cal yr BP until ∼ 1500 cal yr BP, after which agricultural practices waned as the Maya depopulated the area. These changes in plant assemblages were in response to changes in available water resources, with increased aridity leading to the eventual abandonment of agricultural areas. The Ix Chel archaeological site, located in the study area, is a highland site that would have been among the first agricultural settlements to be affected during periods of aridity. During these periods, minimal water resources would have been available in this highly karstified, well-drained area, and supplemental groundwater extraction would have been difficult due to the extreme depth of the water table.     

Climate change and local adaptation strategies in the middle Inner Mongolia,northern China

According to records of 17 meteorological stations distributed in the study area, climate change of the middle Inner Mongolia in northern China was analyzed in this paper. Based on SPOT VGT data, combined with field investigation, local vegetation change was detected in the last 10 years. The results show that annual mean air temperature obviously rose, while precipitation slightly decreased in fluctuation in the study area during the last 50 years. Air temperature increasing rates are +0.318°C 10 year⁻¹ during 1960-2009 and +0.423°C 10 year⁻¹ during 1980–2009, while precipitation decreasing rates are −2.91 mm 10 year⁻¹ during 1960–2009. There were five different dry or wet periods from the 1960s to the 2000s in order, and the wetter 1990s and the drier 2000s changed dramatically in the study area. Local climate totally tend to warm–dry conditions during the last 50 years. According to coefficient of variation (Cv) of yearly growing-season cumulative NDVI value and yearly NDVI maximum in pixel scale, vegetation had experienced huge temporal and spatial variation during the last 10 years. Recently, frequent droughts and dust storms seriously affected local agriculture and grazing activities, and resulted in heavy economic loss, especially over the drought period of 1999–2001. Faced with those drought disasters accompanied with strong dust storms, the local authorities proposed the enclosing-transferring strategy and made great efforts to adapt overt climate change and improve environment, including making selective emigration, decreasing livestock numbers, fencing grasslands and building forage production bases with irrigation instruments and actively adjusting industry structure. However, some effects and potential problems of this adaptation strategy still need to be comprehensively assessed further in longer time scales and aimed at different sub-regions.     

Primary succession and the effect of first arrivals on subsequent development of forest types

The three Krakatau islands in Indonesia were completely denuded by the great eruption in 1883. One century after the eruption, the vegetation of Rakata on the one hand and of Panjang and Sertung on the other are quite different. Indigenous or exogenous Neonauclea seeds successfully formed Neonauclea forest on Rakata island but compltely failed to do so on Panjang and Sertung islands. The dispersal of seeds, soil conditions and volcanic activity of Anak Krakatau were all found to be contributing factors.The Timonius and Dysoxylum forests found on Panjang and Sertung originated on both islands after the appearance of Anak Krakatau. These dominants have larger seeds than Neonauclea and were able to germinate and grow under the regenerated mixed forest canopy but it was not so easy for them to colonize the Neonauclea forest. At the present time, their invasion progresses only gradually on Rakata. Ardisia scrub on steep coastal slopes on the Satsunan islands, SW Japan, is thought to be a somewhat parallel case of the retarding effect on the subsequent development of communities of the first arrivals. Ardisia scrub in these habitats appears to be a stable community, withstanding salt-laden wind. This shrub produces a number of stem and root sprouts, in a mop-headed growth. Their drupes are carried by birds and few seedlings were found in the scrub. The scrub floor was heavily shaded when its canopy was covered by lianes. The deep shade and frugivore activity prevent the invasion of other dominant species which would change the scrub into another type of forest. In milder habitats the scrub gradually becomes tall and the number of sprouts decreases, the forest then changes into another type such as Persea or Castanopsis forest.     

Late Quaternary vegetation and climate dynamics in central-eastern Brazil: insights from a ~35k cal a bp peat record in the Cerrado biome

The late Quaternary evolution of central-eastern Brazil has been under-researched. Questions remain as to the origin of the Cerrado, a highly endangered biome, and other types of vegetation, such as the Capões – small vegetation islands of semi-deciduous and mountain forests. We investigated the factors that influenced the expansion and contraction of the Cerrado and Capões during the late Quaternary (last ~35 ka), using a multi-proxy approach: stable isotopes (δ¹³C, δ¹⁵N), geochemistry, pollen and multivariate statistics derived from a peat core (Pinheiro mire, Serra do Espinhaço Meridional). Five major shifts in precipitation, temperature, vegetation and landscape stability occurred at different timescales. Our study revealed that changes in the South Atlantic Convergence Zone (SACZ) seem to have been coeval with these shifts: from the Late Glacial Maximum to mid-Holocene the SACZ remained near (~29.6 to ~16.5k cal a bp ) and over (~16.5 to ~6.1 k cal a bp ) the study area, providing humidity to the region. This challenges previous research which suggested that climate was drier for this time period. At present, the Capões are likely to be a remnant of a more humid climate; meanwhile, the Cerrado biome seems to have stablished in the late Holocene, after ~3.1 k cal a bp .     

Fifty-thousand-year vegetation and climate history of Noel Kempff Mercado National Park, Bolivian Amazon

Pollen and charcoal records from two large, shallow lakes reveal that throughout most of the past 50,000 yr Noel Kempff Mercado National Park, in northeastern lowland Bolivia (southwestern Amazon Basin), was predominantly covered by savannas and seasonally dry semideciduous forests. Lowered atmospheric CO₂ concentrations, in combination with a longer dry season, caused expansion of dry forests and savannas during the last glacial period, especially at the last glacial maximum. These ecosystems persisted until the mid-Holocene, although they underwent significant species reassortment. Forest communities containing a mixture of evergreen and semideciduous species began to expand between 6000 and 3000 ¹⁴C yr B.P. Humid evergreen rain forests expanded to cover most of the area within the past 2000 ¹⁴C yr B.P., coincident with a reduction in fire frequencies. Comparisons between modern pollen spectra and vegetation reveal that the Moraceae-dominated rain forest pollen spectra likely have a regional source area at least 2-3 km beyond the lake shore, whereas the grass- and sedge-dominated savanna pollen spectra likely have a predominantly local source area. The Holocene vegetation changes are consistent with independent paleoprecipitation records from the Bolivian Altiplano and paleovegetation records from other parts of southwestern Amazonia. The progressive expansion in rain forests through the Holocene can be largely attributed to enhanced convective activity over Amazonia, due to greater seasonality of insolation in the Southern Hemisphere tropics driven by the precession cycle according to the Milankovitch Astronomical Theory.     

Photographic evidence of rate of development of plant cover on the emergent island Anak Krakatau from 1971 to 1991 and implications for the effect of volcanism

Photographs of the vegetated North and East Forelands of Anak Krakatau, which emerged from the sea in 1930 and was totally devastated by volcanic activity in 1952/53 provide some evidence of rate of change of plant cover from grassland to woodland between 1971 and 1991. The series of photographs demostrate that vegetational stages of the two forelands have not been synchronous and suggest that the North Foreland is lagging the East Foreland by about 12 years.A recent hypothesis that Anak Krakatau's vegetation was destroyed, totally or almost so, by eruptions in 1972/73 and the island's vegetation has regenerated since that time is not easily accommodated by the photographic evidence, nor by the relative rates of change on Rakata (Krakatau's post-1883 remnant) and Anak Krakatau. The photographic evidence accords more closely with a hypothesis that the 1972/73 eruptions affected the North Foreland's vegetation only, or affected it to a much greater degree than that of the East Foreland, allowing the latter's post-1952/53 succession to proceed relatively unhampered.     

A high resolution late Holocene palaeo environmental record from the central Adriatic Sea

A multi-proxy study of a Holocene sediment core (RF 93-30) from the western flank of the central Adriatic, in 77 m of water, reveals a sequence of changes in terrestrial vegetation, terrigenous sediment input and benthic fauna, as well as evidence for variations in sea surface temperature spanning most of the last 7000 yr. The chronology of sedimentation is based on several lines of evidence, including AMS ¹⁴C dates of foraminifera extracted from the core, palaeomagnetic secular variation, pollen indicators and dated tephra. The temporal resolution increases towards the surface and, for some of the properties measured, is sub-decadal for the last few centuries.The main changes recorded in vegetation, sedimentation and benthic foraminiferal assemblages appear to be directly related to human activity in the sediment source area, which includes the Po valley and the eastern flanks of the central and northern Appenines. The most striking episodes of deforestation and expanding human impact begin around 3600 BP (Late Bronze Age) and 700 BP (Medieval) and each leads to an acceleration in mass sedimentation and an increase in the proportion of terrigenous material, reflecting the response of surface processes to widespread forest clearance and cultivation. Although human impact appears to be the proximal cause of these changes, climatic effects may also have been important. During these periods, signs of stress are detectable in the benthic foram morphotype assemblages. Between these two periods of increased terrigeneous sedimentation there is smaller peak in sedimentation rate around 2400BP which is not associated with evidence for deforestation, shifts in the balance between terrigenous and authigenic sedimentation, or changes in benthic foraminifera.The mineral magnetic record provides a sensitive indicator of changing sediment sources: during forested periods of reduced terrigenous input it is dominated by authigenic bacterial magnetite, whereas during periods of increased erosion, anti-ferromagetic minerals (haematite and/or goethite) become more important, as well as both paramagnetic minerals and super-paramagnetic magnetite. Analysis of the alkenone, U₃₇^k′, record provides an indication of possible changes in sea surface temperature during the period, but it is premature to place too much reliance on these inferred changes until the indirect effects of past changes in the depth of the halocline and in circulation have been more fully evaluated.The combination of methods used and the results obtained illustrate the potential value of such high resolution near-shore marine sedimentary sequences for recording wide-scale human impact, documenting the effects of this on marine sedimentation and fauna and, potentially, disentangling evidence for human activities from that for past changes in climate.     

Vegetation and environmental conditions in the Doñana Natural Park coastal area (SW Iberia) at the beginning of the last glacial cycle

Detailed reconstructions of the vegetation of Iberia during the last glacial inception are rare due to the limited number of terrestrial sites recording this period. Active retreat of El Asperillo cliff, located on the Atlantic coast of southwestern Iberia, has exposed a fossil organic level dating back to one of the early stades of the last glacial cycle. Pollen and macrofossil analyses from this site show that the Doñana area was covered mainly by steppic vegetation; temperate trees survived the coldest periods, albeit in reduced numbers. Mediterranean taxa are extremely reduced, in contrast with other dry areas of southern Iberia over this time span. This vegetation suggests cold and arid climatic conditions, in accordance with paleoclimatic reconstructions based on several Atlantic marine cores.     

Large variations of δ13C values in stalagmites from southeastern China during historical times: implications for anthropogenic deforestation

Variations in speleothem δ¹³C values can reflect changes in overlying surface vegetation, which, over historical time scales, may represent the influence of human activities. Here, we examined δ¹³C variations in two stalagmites growing for the last 2200 years in Shennong Cave, Jiangxi Province, SE China. The two δ¹³C records corroborate well one another and show a prominent 6‰ enrichment of the δ¹³C values from AD 700 to 1100. The isotopic equilibrium for modern calcite and negative correlation between δ¹⁸O and δ¹³C values along the growth axis suggest that the influences of kinetic fractionation are negligible. Varied correlations between Mg/Ca and Sr/Ca ratios and divergent changes between δ¹³C values and Mg/Ca and Sr/Ca ratios from AD 700 to 1100 reveal that the prior calcite precipitation (PCP) and water–rock interaction did not dominate the increase of δ¹³C values. It is plausible that the obvious δ¹³C variation was largely influenced by the changes in vegetation cover overlying the cave. Our δ¹³C results, together with the records of climate and human activity from historical documentary records, suggest that: (i) prior to AD 700, small fluctuations in relatively light δ¹³C values reflect the presence of lush forest coverage above the cave, which was minimally disturbed by human activities; (ii) during AD 700–1100, the drastic increase in δ¹³C values indicates persistent and massive deforestation associated with large‐scale immigration into northern Jiangxi after the Rebellion of An & Shi (AD 755–763) in the Tang Dynasty and the subsequent development of agriculture and economic activity; and (iii) since AD 1100, fluctuations in relatively high δ¹³C values suggest that local vegetation during the last millennium has been sparse. Since the Rebellion of An & Shi, southeastern China was progressively developed, coincident with deforestation and vegetation deterioration caused by human disturbance in the form of deforestation and cultivation.     

The detailed palaeoecology of a mid‐Wisconsinan interstadial (ca. 32 000 14C a BP) vegetation surface from interior Alaska

We present a multi‐proxy reconstruction from a well‐preserved vegetation surface (ca. 32 000 ¹⁴C a BP) from the Fox Permafrost tunnel near Fairbanks, Alaska. A thick litter layer of plant material on the vegetation surface is consistent with the vegetation lacking evidence of disturbance. Plant macrofossils and graminoid cuticle analysis show the presence of a graminoid assemblage consistent with phytolith data. The pollen data indicate that trees were not local to the site and that Artemisia sp. was present in the region. The insect and bryophyte reconstructions are consistent with the vascular plant reconstruction, indicating the site was at least periodically wet. δ¹³C values from the graminoids present show a large range encompassing both the wet and dry range displayed by modern graminoids in Alaska. Sequential δ¹³C analyses conducted along the length of leaves attached to the vegetation surface indicate a seasonal shift towards relatively higher water use efficiency. The lower water use efficiency earlier in the growing season may have stemmed from the use of winter season meltwater by plants at the site – a scenario consistent with the site's cryostratigraphy. Our multi‐proxy reconstruction contributes to the limited palaeoecological data available for graminoid‐dominated vegetation present in Eastern Beringia and particularly the interior of Alaska during the mid‐Wisconsinan interstadial. Copyright © 2011 John Wiley & Sons, Ltd.     

Development of the ecosystem on surtsey with references to Anak Krakatau

There is a strong similarity in the formation and size of the two new volcanic islands: Anak Krakatau, Indonesia (emerging 1930) and Surtsey, Iceland (emerging 1963). They are both piled up from an ocean floor of over 100 m depth, forming islands of cinder and lava covering more than 2 km² in area. They are both extremely valuable for various ecological studies, ie of investigation of means of dispersal and colonization. However, due to their tropical and subarctic locations, respectively, there is an extreme difference in climate of the two islands and thus in the kind of available life forms, their means of colonization and rate of growth.The first vascular plant was found on Surtsey in 1965, bryophytes became established in 1967 and lichens were first detected in 1970. The advancement of the vascular plant colonization has been of greatest importance in the establishment of vegetation on the island. Twenty-five species of vascular plants, all herbaceous, had been recorded on Surtsey in 1990, the dominant species being Honkenya peploides. Less than 1% ofthe island has vegetation cover, root biomass is low and the organic carbon and nitrogen status of the soil is poorly developed. Birds started nesting on Surtsey in 1970 and six species, all depending on marine life for food, had nested on the island in 1990. The land-invertebrate fauna has developed slowly on Surtsey and the island is devoid of vertebrates other than birds.