Transfer of results from reed research into practice as illustrated by reed protection measures in Berlin

There is great interest in solving the question of how far the successful establishment of common reed for shore restoration is influenced by genetically determined characteristics of different reed clones. For this purpose, ten reed clones were planted at six artificial sand banks in 1995, according to a special planting design. The growth and expansion performance of the clones was compared.The reed plants grew without significant stress in the first vegetation period, resulting in newly established reed stands in a very good condition at the end of the season. Despite the same site conditions at the experimental plots, the planted clones differed significantly in their morphology and stand structure. By comparison, reed plants of the same clone exhibited strong similarity on all experimental plots. These results appear to justify the assertion that, at least during the initial phase of establishing a reed stand, the development of stand structure is determined more by genetically controlled programmes than by site conditions. Only long-term investigations will be able to clarify to what extent stable stand structures are formed by genetically controlled programmes after polycormons reach maturity.     

玉门关段汉长城墙体结构抗震性能研究

基于野外调查和室内试验建立了玉门关段汉长城结构的三维动力分析模型,其中以长城墙体结构中的夯土层、芦苇层和芦苇绳为主要研究对象。采用有限差分法计算了该段长城结构在地震动作用下的抗震性能。计算结果表明:在自重作用下,芦苇层与夯土层变形机制相互补充,芦苇绳的拉结作用明显,整个墙体变形合理;从墙体底部到顶部,地震波的高频成分减少,低频成分增加;在强震作用下,墙体芦苇层的最大应力远小于单根芦苇纤维的抗拉强度,处在比较安全的状态;采用格栅单元和锚索单元分别模拟芦苇层、芦苇绳是合理的。研究成果可为河西地区土遗址保护提供参考。     

苏北溱湖芦苇和芦苇+香蒲群落中植物对湿地土壤N、P的固持效果

湿地土壤是湿地生态系统固持氮(N)、磷(P)的重要库,水生高等植物在湿地土壤固持N、P过程中起到了非常重要的作用.本研究采用室外取样与室内实验结合的方法,对溱湖湿地两种主要湿地类型(芦苇(Phragmites australis)群落和芦苇+香蒲(Typha latifolia)群落)影响湿地土壤N、P固持过程的规律展开研究.分析了芦苇、香蒲各器官生物量和总氮(TN)、总磷(TP)含量及储量对于土壤各土层TN和TP含量的影响.结果显示:(1)溱湖湿地对于水体TN和TP有一定的削减作用,且对TN的削减作用更大;(2)芦苇可以增强湿地土壤(30 cm以下的土层)富集N的效率,并且芦苇+香蒲群落中土壤固N效率更高,芦苇植株内TN和TP储量都是根茎叶穗,而香蒲则是根穗茎叶;(3)芦苇和香蒲茎、叶器官的TN和TP含量在夏季均显著高于其他几个季节,尤其是芦苇茎的TN含量在夏季高出其他几个季节70%~84%,而TP含量甚至高出其他几个季节81%~92%;(4)芦苇、香蒲对于P贫瘠的响应机制不同,导致芦苇会消耗土壤P,而香蒲的介入可以抵消这一消耗过程.因此,芦苇+香蒲的植物配置模式可以提高湿地土壤固持N、P的综合效率.     

Geschichte und Ursachen des Röhrichtrückgangs am Ammersee (Deutschland)

The development of aquatic reed stands at Lake Ammersee was documented by using vertical airborne photographs which were taken during four partial and five complete flights beginning in 1944. A dramatic decline of reed, especially of those parts which stood in deep water, was demonstrated. Between the years 1963 and 1969 60% and until 1992 another 30% of the Phragmitetum disappeared. Only at the east bank did the lake reed beds slightly expand.The examination of possible factors affecting the development of reed at Lake Ammersee lead to the following scenario: Until 1940 reed stands at Lake Ammersee spread and covered large aquatic areas. Only the east bank of the lake, where the influence of wind and waves was high, and banks near towns and recreation areas remained uncovered. Since the regulation of the River Ammer between 1920 and 1938 flood occurrence increased enormously. The flood disaster of the year 1965 coincided with the period of the highest rate of reed decline. The peak of eutrophication in the lake was reached in 1975. Huge carpets of green algae caused further retreat of reeds.At the present time recovery is not possible because of the negative influence of human recreation and of reed consuming waterfowl. Interestingly enough an expansion of reed has been observed at the east bank of Lake Ammersee at places were there was no Phragmites previously.     

Dynamik der Schilfröhrichte am Bodensee unter dem Einfluss von Wasserstandsvariationen

On the basis of different sets of aerial photos the dynamics of the reed bed areas of Lake Constance were investigated in relation to the dynamics of the water levels. The objectives of the study were to quantify the changes of reed areas due to different flood events in the last decades and their recovery in the time periods between these events. The results should given information of the relevance of water level variations on reed bed dynamics and the regeneration times of reed beds after extreme disturbance events.Following the extreme flood at Lake Constance in 1999 the reed belts of Lake Constance lost approximately 30 ha (24%) of the lakeside reed beds. The loss is comparable to the situation in the late 1960s, when approximately 40 ha died back due to the extreme flood in 1965 and the high spring water levels in the subsequent years. In the time period between the extreme floods of 1965 and 1999, the reed areas expanded to nearly 85% of the area before 1965. The expansion rates increased with increasing distance to the flood event of 1965. Especially in periods with series of years of low spring water level the expansion rates were high.The damage degrees of the reed areas in the years 2000 and 2002 showed a clear relation to the elevation (i.e. average water level) of the stands. The damage degree increased with decreasing elevation. Furthermore the regeneration process of severely damaged stands was related to the elevation level of the stands. Whereas stands at high elevation regenerate fast, those at low elevation died off completely in the years after the extreme flood. This supports the hypothesis that the water level flutuations play a major role in the reed dynamics of Lake Constance.As a consequence of the climate change an increase in the frequency of high spring water levels is expected. Thus, it seems unlikely that reed stands will ever expand again to the same area as before 1965.     

Einsatz von Pflanzenkläranlagen zur Behandlung von schadstoffbelastetem Oberflächenabfluß städtischer Straßen

A combination of retention pond and reed bed was tested for its effectiveness in reducing non-point source runoff pollution from an urban area. This paper presents data on the development of reed plants (Phragmites australis Cav. Trin.), on the effectiveness of runoff purification and on the accumulation of contaminants in the pond sediment and the reed bed for the years 1993–1995. P. australis was well established and grew according to normal development. The measured length and biomass values of P. australis were larger on sand than on gravel and coarse materials. Toxic levels of heavy metals in the plants were not reached. Harvested plants can be composted. The purification system is effective within the following ranges: The rate of retention of suspended solids and heavy metals varies between 16% and 91%. For chemical oxygen demand the average values is 25%. Poor retention rates usually occur along with very low input concentrations. Generally retention within the reed bed is higher than in the pond. Between 1993 and 1995 the concentration of contaminants within the organic layer of the reed bed increased by 50% for lead and by 90% for poly aromatic hydrocarbons. Until 1994 the concentrations of mineral oil hydrocarbons also increased by 60%. Between 1994 and 1995 however the concentration of mineral oil hydrocarbons in the reed bed decreased rapidly by more than 50%: Mineral oil hydrocarbons underlie biological degradation within the reed bed. In the pond sediment a significant accumulation of heavy metals, poly aromatic hydrocarbons and mineral oil hydrocarbons has also been found. The concentrations of these contaminants are still far from inhibiting the function of the system.     

Distribution of the aphid Hyalopterus pruni Geoffr. within and between habitats of Common Reed Phragmites australis (Cav.) Trin. ex Steudel as a result of migration and population growth

The aphid Hyalopterus pruni migrates to its secondary host Common Reed in late spring and early summer. Reed fields are known to be more heavily infested at the edges compared to the centre, as the migrants are attracted to the colour of the reed. There is also evidence that reeds growing at wet sites are more heavily infested than at dry sites. This paper shows that both proceses interact in determining the distribution of aphids in different habitats.The observations were carried out in the vast reed field of the Zuidelijk Flevoland polder (The Netherlands) during a number of years after its reclamation in 1968. In the period of migration flying aphids were caught with sticky traps situated in and around the reed field. The distribution of aphids on the reed was determined at different times during the season along transects through the centre of the polder. The existence early in the season (June) of a higher infestation at the edges than in the centre of the field was confirmed. The density of winged aphids in this early period was higher in wet than in dry habitats, which may have been a result of a preference of the settling migrants for reed standing in water. Later in the season (August) the dry habitat still had a relatively low infestation in the centre, but at the wet sites the infestation was high everywhere. This could be attributed to the much quicker population growth that was found in the wet habitat and subsequent short range movement of newly born winged aphids.     

Stickstoff- und Kohlenhydratspeicherung in Rhizomen von Phragmites australis (Cav.) Trin. ex Steudel an unterschiedlichen aquatischen Standorten oberbayerischer Seen

Nitrogen and carbohydrate utilization in Phragmites australis rhizomes was investigated under natural conditions at 10 sites in 5 lakes in Upper Bavaria over a period of two years. Nitrate and ammonium of surface and interstitial water was determined. The results indicate that conditions of habitat and environment, such as climate and fluctuation of water level, lead to different adaptations of reed. These parameters had an effect on the storage of nitrogen and carbohydrate. The reed stands exposed to high water level during spring and summer have a high carbohydrate content in the rhizomes. A positive correlation was shown between the aboveground biomass and the content of storaged carbohydrates during the vegetation period. No relationship could be demonstrated between the nitrogen availability of the habitats and the type of storage of carbohydrates and nitrogen. The nitrogen content in the rhizomes can be used as an indicator of the ability of reed stands to produce new secondary shoots after shoot loss. The differently established reed genotypes are not able to compensate short-term environmental changes, such as growing leisure problems and higher shoot loss by the increasing appearance of waterfowl, from their reserve storage.     

洞庭湖区不同退田还湖模式下湿地植被恢复特征的比较

以2006年野外调查资料为依据,对洞庭湖区3种"双退"恢复模式(自然恢复、种植荻和种植杨树)下的植被特征进行了比较分析.结果表明,自然恢复模式下的物种数最为丰富,达79种,而种植荻模式下的物种数最为稀少,仅36种;自然恢复模式下湿生植物种类最为丰富,占物种总数的81.0%,而种植杨树和荻模式下所占比例分别为78.7%和...     

Influence of reed stem density on foredune development

Vegetation density on foredunes exerts an important control on aeolian sediment transport and deposition, and therefore on profile development. In a long‐term monitoring field experiment, three plots were planted with regular grids of reed bundles in three different densities: 4, 2 and 1 bundles per m². This study reports on the differences in profile development under the range of vegetation densities. Topographic profiles were measured between May 1996 and April 1997. Results indicate important differences in profile development for the three reed bundle densities: in the highest density plot a distinct, steep dune developed, while in the lowest density a more gradual and smooth sand ramp was deposited. When the stems had been completely buried, differences in profile evolution vanished. After a second planting of reed stems in January 1997 the process was repeated. In May 1997, all plots had gained a sand volume ranging from 11·5 to 12·3 m³ m^?1, indicating that the sediment budget is relatively constant, regardless of the particular profile evolution. The field evidence is compared with simulations of profile development, generated by the foredune development model SAFE. The model successfully reproduces the overall profile development, but in general, the equations used for vegetation–transport interaction overestimate the effect of vegetation. This causes some deviations between field and model results. Several reasons for this are discussed. Based on the experiments reported here, recommendations are given for further research. Copyright © 2001 John Wiley & Sons, Ltd.     

Effects of flooding on the amino acid and carbohydrate patterns of Phragmites australis

Changes in the contents and composition of dissolved amino acids and carbohydrates were monitored in different Phragmites australis stands in Germany. Four different reed clones planted in 1997 in a degraded fen area in NE-Germany were compared with respect to the seasonal development in the total amount of amino acids and sugars in the basal culm internodes. The individual Phragmites clones showed significant differences in the absolute content of both parameters indicating an influence of the genotype. Flooding affected the ratio of amino acids to carbohydrates within the reed clones. Plants grown in flooded parts of the fen area revealed a significantly higher total amino acid content together with a lower total content of carbohydrates.Furthermore, the effects of extreme flooding on the content and composition of amino acids and carbohydrates in the basal culm internodes of Phragmites australis were studied in nine reed stands with different degrees of damage on Lake Constance. Shoots from extremely damaged reed stands were characterised by a significant decrease in the total content of carbohydrates, an increase in the total content of amino acids and an increase in the share of γ-aminobutyric acid. These changes in the content and composition of carbohydrates and amino acids were discussed with respect to their value as stress indicators, such as for oxygen deprivation which causes a hypoxic metabolic state in Phragmites australis stands.     

Carbohydrate and nutrient content in rhizomes of Phragmites australis from different habitats of Lake FertÕ/Neusiedlersee

Changes in the soluble sugar, starch, total non-structural carbohydrate (TNC) and nutrient (N, P, K) concentration in the reed rhizome were investigated in different reed habitat types in Lake Fertõ/Neusiedlersee during the vegetation period in 1993 to 94. Three die-back and four vigorous reed stands were monitored.For biomass estimations rhizomes were collected twice, in early summer and in autumn. They were calculated to be 1.9 to 2.7 kg/m² at die-back and 3.1 to 6.1 kg/m² at vigorous sites.The carbohydrate and nutrient concentrations of the reed rhizomes were high at the beginning of the vegetation period; they reached their minima in May–June, which was followed by a gradual increase until the end of the vegetation period. From autumn to spring the concentrations decreased, e.g. TNC concentration by 15 to 40% at vigorous, 50 to 70% at die-back sites.In both years the lowest carbohydrate concentrations were recorded at the end of May, a month before the inflorescence developed (vigorous sites: 50–70 mg TNC/g, die-back sites: 70–120 mg TNC/g). In that period the standing stock of the TNC in the rhizome did not separate the sites clearly, and the smallest carbohydrate standing stock was measured at a vigorous site with a high water cover.At the beginning of summer and in autumn the soluble sugar, TNC and N concentration values were higher at the die-back than at the vigorous sites but the standing stock was higher at the vigorous sites. In autumn the TNC standing stock of the rhizomes was between 780 and 1200 g/m² at vigorous, between 380 and 690 g/m² at die-back sites. This is especially important for the reactions and resistance of die-back reed stands to unfavourable conditions.     

The impact of reed-protecting structures on littoral zones

The impact of several types of reed-protecting structures on hydrodynamic conditions, sediment properties and littoral water quality were investigated. All types of embankment lead to a dissipation of wave energy. The embankment with fagots (type 1) shows a limited porosity in contrast to the other types examined. All kinds of embankment enhance the accumulation of organic matter at the land/water-interface. Hence, they protect littoral accumulation zones from erosion. The embankment with fagots causes changes in sediment properties at the seaward side of reed belts. The surface of mineral sediments is turned into a fine-grained mud, with a high content of organic matter. Accelerated sedimentary microbial processes (oxygen demand) are produced by increasing nutrient availability (carbon, nitrogen). As a result, the O₂-saturation of water bodies is significantly reduced. This is in contrast to the embankment with wooden partitions and palisades (type 2 and 3), which had no significant effects on sediment conditions and water quality. The consequences of changes in type 1-protected reed stands are discussed. Practical recommendations are given for the further use of reed-protecting structures.     

井下甚宽频带地震仪关键装置研究进展

井下甚宽频带地震仪关键装置研究进展主要包括:分向核心芯体机械及簧片系统、核心反馈调理系统及井下定位定向系统等。      

富营养化湖泊蓝藻-芦苇碎屑堆积分解过程及其潜在的共代谢效应

为探究富营养化浅水湖泊所富集的有机物对湖泊碳循环和水质的影响,本研究构建微宇宙系统,模拟蓝藻和芦苇碎屑单独分解及混合分解过程.通过测定各组上覆水营养盐浓度、有机质含量及结构的变化,揭示富营养化湖泊藻草残体混合分解过程中养分和有机碳的释放特征.结果表明,在实验0～88 h内,在添加相同的碳源条件下,蓝藻和芦苇混合处理组总碳(TC)释放量显著高于理论值,表明藻草碎屑混合分解存在共代谢效应.在培养初期,沉积物通过共代谢效应对水质产生了较大的影响,加速向水体中释放氮、磷物质.相较单独的植物分解,混合处理组中总氮(TN)、总磷(TP)的最大释放量分别提高了13.49%和26.84%;通过三维荧光光谱表征的类富里酸荧光强度变化也表明:较芦苇处理组,混合处理组中芦苇的分解速率更快.在培养开始后,各处理组均快速转变为厌氧状态,TC、TN、TP浓度随时间变化总体上呈先快速上升再逐渐平缓的趋势,分别在第228、108和324小时达最大值(372.4±2.98)、(138.45±2.97)和(7.95±1.11) mg/L.细菌特异性脂肪酸含量变化表明,将蓝藻碎屑添加到芦苇碎屑中,会增加芦苇碎屑中细菌的丰度,从而提高分解速率,激发共代谢效应.在全球气候变暖的背景下,随着富营养化湖泊藻类暴发频次增加,共代谢效应可能还会进一步加强,对富营养化湖泊水质将会持续产生影响.     

Reed litter Si content affects microbial community structure and the lipid composition of an invertebrate shredder during aquatic decomposition

The decomposition of plant litter is an important mechanism in regard to energy and nutrient dynamics of ecosystems. Silicon concentration of plant tissue can affect these processes by changing litter quality, i.e. nutrient stoichiometry and cellulose and phenols content. To determine which group of microbial decomposers benefits from high Si content in plants and how this impacts on animal decomposers, a batch experiment was conducted with reed leaf litter (Phragmites australis) differing in Si content in the presence/absence of invertebrate shredders (Gammarus pulex). Lipid concentration of G. pulex, in reed litter and fine particulate matter (FPOM) were examined. High Si concentration in reed resulted in a decline of gram positive bacteria in the heterotrophic biofilm and of gram negative bacteria in FPOM. The lipid composition in the next trophic level, the decomposer G. pulex, changed too, indicating a diet shift in favor of bacteria and algae with increasing litter Si concentration. Thus, basal decomposers were affected by the Si availability in plant resources, and these effects likely persist along the food chain, as FPOM is a dominant food supply for other groups, e.g. collectors. This impact of Si content on plant substrate quality for decomposer food webs may have global relevance, due to related modifications in carbon and nutrient cycling during litter decomposition.     

江苏滆湖北部区整治后浮游植物时空分布及环境因子变化规律

滆湖是我国长江中下游典型的浅水型湖泊,为了解其治理后浮游植物群落时空分布规律,2013年1 12月对其北部区浮游植物及环境因子进行调查.调查期间共检出浮游植物7门43属61种,春、冬季以栅藻(Scenedesmus)和小环藻(Cyclotella)为主要优势种属,夏、秋季以微囊藻(Microcystis)和颗粒直链藻(Melosira granulata)为主要优势种属,采样期间浮游植物生物量最高值为90.6 mg/L,出现在8月份,铜绿微囊藻(Microcystis aeruginosa)占绝对优势.浮游植物平均密度呈现由西向东递减的趋势,植被覆盖区低于敞水区.环境因子分析表明:总氮浓度、总磷浓度、水温是影响滆湖北部区浮游植物密度和生物量的主要因子.比较相同月份湖区内部菱角芦苇区和未治理的敞水区的平均生物量,菱角区生物量较敞水区低约72.7%~91.1%,芦苇区生物量较敞水区低约63.9%~83.7%.在8、9月湖区内敞水区暴发水华时菱角区浮游植物生物量仅为敞水区的14.6%,芦苇区为敞水区的30.3%.     

Early succession of the macroinvertebrate community in a shallow lake: Response to changes in the habitat condition

In the present study the initial succession of the aquatic macroinvertebrate communities of a newly created shallow lake in a wetland area was monitored during 15 months. Three different types of macroinvertebrate samples (multihabitat, associated to the Phragmites australis stands and associated to the sediment) were collected monthly from May 2004 to July 2005. Additional samples were collected in the reed stand in 2007, when the reed belt had become much taller and thicker. Colonization of the lake was fast, and the colonization sequence was mainly related to the dispersal abilities of the taxa. Habitat-specific changes in the assemblage's structure and composition were registered. The communities associated to the sediment showed a decrease in overall biomass, density and species richness along time. The density of the gastropod Physella acuta and the chironomid species Dicrotendipes pallidicornis, Polypedilum nubifer and Tanytarsus horni decreased significantly, after Chara stands declined; while the chironomid Chironomus riparius became dominant. The epiphytic macroinvertebrate communities associated with the reed stands followed a seasonal pattern, with a warm-period community dominated by the chironomid species Ablabesmyia monilis and Psectrocladius sordidellus-group, and a cold-period community dominated by the chironomid species Cricotopus ornatus and D. pallidicornis.     

Susceptibility of lakeside phragmites reeds to environmental stresses: Examples from Lake Constance-Untersee (SW-Germany)

The lakeside border of Phragmites reeds is under the control of many environmental factors, among which the flooding height and the degree of mechanical load by waves and washes are the most prominent. Examples from Lake Constance-Untersee show that the reed front retreats under enhanced stresses. Models are presented that describe the susceptibility of lakeside reeds to mechanical damage and to flooding events. A monitoring programme which ran from 1984/85 to 1992/93 showed a slow lakeward progression of the reeds in a decade of abnormally low water conditions.     

Characteristics of reeds on Lake Peipsi and the floristic consequences of their expansion

The reeds of Lake Peipsi (surface area 3555 km², mean depth 7.1 m, max. depth 15.3 m, not-regulated) have increased through the last 30 years. The most significant increase has taken place in the larger formerly mesotrophic northern part where the mean air-dry biomass of reed shoots has increased from 20 g m⁻² in 1970 through 789 g m⁻² in 1989 to 1563 g m⁻² in 2002. The data of 34 transects for 2001–2002 along the Estonian coast and aerophotos made in 2001 showed that in places reeds reach a water depth of 1 m (mean 0.64 m) and a width of more than 200 m (mean 89.5 m). Their area of 9.3 km² accounts for about 0.6% of the Estonian territory of the lake. The longer and thicker shoots grew in the lakeward and intermediate zones of reeds. Near villages the shoots were weaker and their biomass lower. Phosphorus content of shoots, being higher near settlements, was generally low; in some cases content of phosphorus was ≤0.001%. The monotonous reed belt occupies the growth area of shallow-water submergent and shorter emergent species and favours the accumulation of organic sediments. The frequency of several species demanding light and sandy bottom has decreased, as well has the overall diversity of the macrovegetation. The main reasons for expansion of the reed could be eutrophication, combined with low-water periods, and the decline of cattle breeding in the shore areas. Some suppressed species have reappeared on stretches cleaned from reeds.