Toarcian–Aalenian (Early–Middle Jurassic) radiolarian fauna from the Los Molles Formation,Neuquén Basin,Argentina: Taxonomy and paleobiogeographic affinities

This paper reports the occurrence of a Toarcian–Aalenian (Early–Middle Jurassic) radiolarian fauna in the Los Molles Formation, Neuquén Basin, Argentina, as well as comments on its paleobiogeographic affinities. The micropaleontologic analysis was carried out in fine-grained rocks from a turbiditic section of the Los Molles Formation. These samples were first chemically processed using only hydrogen peroxide (H₂O₂), and afterward treated with acetic (CH₃COOH) and hydrofluoric (HF) acids. The first chemical procedure permitted the recovery of only few spongy spumellarians, while the second one enabled to recover more diversified radiolarian assemblages. In general, the studied fauna presents low diversity and abundance, with a strong dominance of spumellarians over nassellarians. The fauna is composed by the genera Paronaella, Homoeoparonaella, Praeconocaryomma, Archaeocenosphaera, Orbiculiformella, Praeparvicingula, and some unidentified spumellarians and nassellarians. According to paleobiogeographic models, the studied Toarcian–Aalenian fauna presents a mid to high latitude affinity. It is possible to infer from those data a bipolar distribution of some taxa, such as Praeparvicingula and probably Praeconocaryomma, between the Northern and Southern hemispheres since the Early Jurassic (Toarcian).     

Stratigraphy and ammonite fauna of the upper Shemshak Formation (Toarcian–Aalenian) at Tazareh,eastern Alborz,Iran

With a thickness of 3900 m, the Tazareh section is one of the thickest developments of the Shemshak Formation in the Alborz range. It overlies with sharp and disconformable contact the limestones and dolomites of the Lower–Middle Triassic Elikah Formation and is topped, again with a disconformable contact, by the marls and limestones of the Middle Jurassic Dalichai Formation. The nearly exclusively siliciclastic succession represents a range of environments, from fluvial channels, flood plains, swamps and lake systems to storm-dominated shelf, and a comparatively deep marine and partly dysoxic basin. The segment of the section between 2300 and 3500 m is exclusively marine and contains a moderately diverse ammonite fauna, ranging from the Middle Toarcian to the Upper Aalenian. The ammonite fauna comprises 21 taxa, among them the new genus Shahrudites with two new species, Shahrudites asseretoi and S. stoecklini from the Middle Aalenian Bradfordensis Zone. The other ammonites from the Shemshak Formation at Tazareh (as elsewhere in North and Central Iran) are exclusively Tethyan in character and closely related to faunas from western and central Europe. An ammonite-based correlation of Toarcian–Aalenian successions of the eastern Alborz with time-equivalent strata of the Lut Block, part of the Central-East Iranian Microcontinent (ca. 500 km to the south), suggests a strong influence of synsedimentary tectonics during the deposition of the upper Shemshak Formation.     

“Cretaceous black flysch” in the Pieniny Klippen Belt, West Carpathians: a case of geological misinterpretation

This is a critical assessment of the paper by Oszczypko et al. (2004: Cretaceous Research 25, 89–113), in which they tried to prove a mid-Cretaceous age for the Szlachtowa (“black flysch”) and Opaleniec Formations, in the Pieniny Klippen Belt, West Carpathians, both of which had previously been shown to be of Jurassic age. We argue that the mid-Cretaceous age assignment is a misinterpretation, primarily resulting from their field samples having been collected from some Cretaceous lithostratigraphic units, tectonically associated with the Jurassic formations, and/or from tectonic contact-breccias involving Jurassic and Cretaceous strata. In addition, we suggest that they have overlooked a number of significant palaeontological papers, published since 1962, which record the presence of in situ ammonites, aptychi, belemnites, thin-shelled bivalves (Bositra), gryphaeids, foraminifera, and ostracod assemblages, all indicating a Jurassic (mainly Aalenian), and not a Cretaceous, age for the Szlachtowa Formation, and also the in situ Jurassic (Bajocian) ammonites and thin-shelled bivalves (Bositra), Bositra-microfacies, and age-diagnostic foraminiferal assemblages of the Opaleniec Formation.Our presentation here of recently published dinocyst data from well-preserved assemblages further supports the Jurassic ages for the Szlachtowa (“black flysch”) and Opaleniec Formations.     

Lytoceratid ammonites from the Inferior Oolite Formation (Middle Jurassic,Aalenian and Bajocian) of Dorset (United Kingdom)

Ammonites of the family Lytoceratidae from the Middle Jurassic Inferior Oolite Formation, Aalenian to lowermost Bathonian, of Dorset are rarely described, perhaps because the macroconchs are often very large and difficult to collect intact and the microconchs are very small and easily overlooked. Detailed stratigraphical collecting over several years has shown them to be a minor part of the ammonite fauna but more common at certain horizons. Four genera, Lytoceras, Megalytoceras Nannolytoceras and Pleurolytoceras have been shown to have different stratigraphical ranges and abundances in the Aalenian and Bajocian but they have not been found in the Lower Bathonian of Dorset. The taxonomic basis of several of the well-known species is poorly described in the literature and is remedied here.     

Jurassic–Cretaceous radiolarian biostratigraphy and sedimentary environments of the Ceno-Tethys: records from the Xialu Chert in the Yarlung-Zangbo Suture Zone,southern Tibet

Radiolarian biostratigraphic research has been carried out along two continuous sections through the Xialu Chert, one of the accreted sheets included in the Yarlung-Zangbo Suture Zone. Six radiolarian zones have been identified as follows: Laxtorum(?) jurassicum Zone (Aalenian), Tricolocapsa plicarum Zone (Bajocian–lower Bathonian), Stylocapsa(?) spiralis Zone (upper Callovian–Oxfordian), Hsuum maxwelli Zone (Kimmeridgian), Pseudodictyomitra carpatica Zone (upper Tithonian–lower Valanginian), and Turbocapsula costata Zone (Aptian).A reconstructed stratigraphy of the Xialu Chert, based on lithological succession and radiolarian dating, indicates that the chert exhibits a long depositional history, at least from early Middle Jurassic (Aalenian) to late Early Cretaceous (Aptian). The separation of the Lhasa Block from the northern Gondwana margin must, therefore, be dated before the Aalenian. The absence of calcareous sediments in the Xialu Chert indicates that the oceanic basin was deeper than the CCD throughout the depositional history. The transition from chert to siliceous mudstone is recorded sometime in Early Cretaceous, most probably around the Barremian/Aptian boundary. This means that the oceanic plate had already started being consumed at a trench by that time. The accretion of the Xialu Chert occurred after the Aptian time.     

High resolution ammonite–benthic foraminiferal biostratigraphy across the Aalenian–Bajocian boundary in the Lusitanian Basin (Portugal)

This work describes the ammonite and benthic foraminiferal assemblages recorded across the Aalenian–Bajocian boundary of the Serra da Boa Viagem II section, located about 6 km to the east of the Bajocian GSSP (Murtinheira, Portugal), and calibrated to the standard ammonite zonation previously established for the Lusitanian Basin. A total of 220 ammonite specimens referred to 30 fossiliferous levels were collected and identified throughout the section, enabling the recognition of the Concavum Zone (Concavum and Limitatum subzones) of the upper Aalenian, and the Discites Zone of the lower Bajocian. A total of 2356 foraminifers were obtained from the 16 samples collected along the section, corresponding to 4 suborders, 8 families, 16 genera and 44 species. The occurrence of Lenticulina quenstedti (Gümbel) has enabled the recognition of the Lenticulina quenstedti Zone, ranging from the Bradfordensis Zone (middle Aalenian) to the lower Discites Zone (lower Bajocian). The first record of Ramulina spandeli Paalzow, whose occurrence, up to now, was limited in the Lusitanian Basin to the Murtinheira section (the Bajocian GSSP), highlights the usefulness of the Ramulina spandeli Zone, with its lower boundary referred to the lower Discites Zone (lower Bajocian). Other bioevents displaying local, basinal or regional biostratigraphic interest have also been identified. The benthic foraminiferal record here presented, accurately calibrated with the ammonite record, aims at contributing to support the recognition of the Lenticulina quenstedti Zone and the Ramulina spandeli Zone as formal biostratigraphic units integrating the biostratigraphic scale based on benthic foraminifers for the Aalenian–Bajocian boundary in the Lusitanian Basin (Portugal). Copyright © 2014 John Wiley & Sons, Ltd.     

Stable isotope records (δO and δC) of Lower-Middle Jurassic belemnites from the Western Balkan mountains (Bulgaria): Palaeoenvironmental application

Stable O and C isotope data of 110 Upper Pliensbachian-Lower Bajocian belemnites have been obtained and used to attempt a reconstruction of palaeotemperature and its variation in two epicontinental depositional environments from the Western Balkan mountains (Bulgaria). The samples were collected from 3 sections with high-resolution ammonite subdivision. Initially taphonomic, cathodoluminescence and geochemical analyses were used for evidence of diagenetic alteration. Non-luminescent parts of the belemnite rostra have been sampled for isotope analyses and 76 samples, having δ¹⁸O < −4‰ (PDB), δ¹³C > −0.5‰ (PDB), Fe < 250 ppm, Mn < 50 ppm, Sr > 950 ppm and Sr/Mn ratio > 80 were used for palaeotemperature interpretations. The O and C isotope data generally exhibit little stratigraphical variability with minor fluctuations, however, there are prominent positive C isotope excursions and coeval negative O isotope shifts detected in the Lower Toarcian Tenuicostatum, Falciferum and Bifrons Zones. The O isotope data, interpreted in terms of palaeotemperature, revealed relatively high seawater temperatures during the Toarcian, Aalenian and Early Bajocian, with detectable temperature rises during the Early Toarcian (maximum value of 29.6 °C). Both C isotope maxima and O isotope minima are used as evidence of the Early Toarcian anoxic event reported from many localities of the same age and in similar facies in Western Europe. In the study the latter is recognized as 3 episodes, which are closely related with the seawater temperature maxima. This isotope record pattern is considered as a consequence of a global Tethyan transgression during the Early Toarcian.     

The oldest known metriorhynchid crocodylian from the Middle Jurassic of North-eastern Italy: Neptunidraco ammoniticus gen. et sp. nov.

Metriorhynchidae is a clade of marine-adapted crocodilians known from several Middle Jurassic–Early Cretaceous specimens collected predominantly in South America and Europe, but poorly known in the northern margin of Gondwana. The “Portomaggiore crocodile” is the most complete specimen of an Italian metriorhynchid to date: it consists of a partial skeleton that has been provisionally referred to an unnamed species of Late Jurassic Metriorhynchus or Geosaurus. The specimen is preserved in the reddish, nodular limestone of the Rosso Ammonitico Veronese Formation (Bajocian–Tithonian); new data on microfossil associations constrain the age of the metriorhynchid to the late Bajocian–earliest Bathonian. On the basis of cranial synapomorphies, the “Portomaggiore crocodile” falls as the closest sister-taxon of the Late Jurassic–Early Cretaceous geosaurines, and is referred to Neptunidraco ammoniticus gen. et sp. nov. It is unique among Middle Jurassic metriorhynchids in showing an incipient streamlining of the skull, shared with Late Jurassic and Cretaceous taxa. Since Neptunidraco is the oldest known member of Metriorhynchidae, its phylogenetic position supports the hypothesis that the timing of the initial metriorhynchid and geosaurine diversifications should start in the Bajocian.     

吐鲁番拗陷中侏罗世孢粉植物群及古环境

根据吐鲁番拗陷葡１井和草南１井中侏罗世的孢子花粉化石及其组合特征，研究孢粉植物群、古植被面貌及古生态环境，探讨中侏罗世的古气候性质、演变及其对聚煤作用和油气生成的影响。     

Synrift sedimentation on the northern Tethys margin: an example from the Ligurian Alps (Upper Triassic to Lower Cretaceous,Prepiedmont domain,Italy)

The Prepiedmont domain succession of the Ligurian Alps is formed by a thick Mesozoic sedimentary cover tectonically detached from its substratum. The Arnasco–Castelbianco unit preserves the most complete record of the Ligurian Prepiedmont, although completely overturned and deformed due to Alpine tectonics. It is composed of carbonate and clastic rocks deposited during the Upper Triassic to Lower Cretaceous interval. This paper is focused on the stratigraphy of the Jurassic series and its relationships to the Tethyan rifting. Each term of the sedimentary record is seen as a witness of the several phases through which the rifting took place. An early rifting phase (Late Hettangian to Early Sinemurian) brought to the formation of a normal fault system affecting the carbonate platform and favoured the development of condensed sedimentation on pelagic highs. The rapid transition from open-platform carbonates to slope-basin cherty limestones testifies the increased subsidence of the margin in the Late Sinemurian, during which moderate fault activity is recorded (intraformational breccia horizons). Until the Early Pliensbachian, a tectonic pause brought to the sedimentation of a succession of pelagic carbonates, occasionally interrupted by clastic flows. During the Late Pliensbachian (?) to Toarcian, the rifting phase followed, evidenced by the large amount of clastics and generated by renewed fault activity. Clastics flowed down into the basin as fluxoturbidites first, and then passed to breccias during the maximum tectonic pulse. In the Late Toarcian to Aalenian (?), the thermal uplift of the Briançonnais shoulder generated a basin fill of fine clastics. The following thermal subsidence (Aalenian to Tithonian) favoured the restoration of quiet basinal conditions evidenced by the deposition of radiolarites.     

New records of rare ammonite species from the Inferior Oolite Formation (Middle Jurassic,Aalenian and Lower Bajocian) of Dorset

Rare ammonite species are reported from the Inferior Oolite Formation of Dorset. In the Aalenian the ammonite Shahrudites is described from the Scissum Zone for the first time and Malladaites is recorded from the Murchisonae Zone. Tmetoceras regleyi occurs in the Scissum and Murchisonae zones and Tmetoceras henriquesae is described from the Concavum Zone. In the Lower Bajocian, specimens of Zurcheria are presented from the Discites Zone. A revised zonal scheme is used for the Aalenian Stage and two new biohorizons are introduced. The Leioceras subglabrum biohorizon is erected in the Opalinum Subzone and Leioceras evolutum in the Bifidatum Subzone.     

Early Jurassic climatic trends in the south-Tethyan margin

The Early Jurassic period was characterized by extreme environmental changes, as reflected by major global carbon isotope anomalies and abrupt changes in oxygen isotope and elemental records of marine organisms. Available data suggest an overall warm Early Jurassic climate interrupted by periods of severe cooling, with a climatic optimum during the early Toarcian. Available geochemical studies, however, have mainly focused on the northern margin of the Tethys Ocean, so that the palaeogeographic extent of these environmental perturbations, latitudinal palaeotemperature gradients and climate belt boundaries remain poorly constrained. Here we report the first stable isotope records of brachiopod shells (δ¹³C and δ¹⁸O values) from the Upper Sinemurian-Middle Toarcian interval in the southern margin of the Tethys Ocean (northwest Algeria). These data were used to better constrain the palaeoenvironmental evolution of the North Gondwana margin during the Early Jurassic, which likely played an important role on supra-regional climate. The diagenetic history of the analysed brachiopod shells was monitored using scanning electron microscopy, and elemental (manganese and strontium) compositions. The brachiopod δ¹³C and δ¹⁸O data show very similar trends as those reported for various Tethyan regions, and record negative carbon and oxygen isotope excursions near the Sinemurian–Pliensbachian and Pliensbachian–Toarcian transitions and during the Toarcian oceanic anoxic event (T-OAE). Despite these similarities, the carbon and oxygen isotope records are systematically offset towards more positive δ¹³C values (average +0.5‰) and more negative δ¹⁸O values (−1.0‰) compared to those obtained from sites of higher palaeolatitudes in the northern Tethyan margin. These offsets suggest a spatial heterogeneity in the stable isotope composition of dissolved inorganic carbon in the Early Jurassic Ocean and a marked latitudinal temperature gradient between the southern and northern margins of the Tethys.     

Correlation of the Jurassic-Cretaceous siliceous-volcanogenic sediments in northwestern surroundings of the Pacific (Koryak Upland)

In distribution areas of the Pekul’neiveem and Chirynai formations customary distinguishable in the Koryak Upland, complicated tectonostratigraphic units are composed of alternating thrust sheets of different lithologic composition and age, which are juxtaposed because of widespread thrust faulting, as is proved by the radiolarian analysis. Nineteen radiolarian assemblages of different age are first established here in the Lower Jurassic-Hauterivian succession of siliceous-volcanogenic sediments. In the Lower Jurassic interval, the lower and upper Hettangian, lower and upper Sinemurian, and Pliensbachian beds are recognized. Paleontological characterization is also presented for the Aalenian (or Toarcian?-Aalenian), upper Bajocian, lower and upper Bathonian, and Callovian beds of the Middle Jurassic. Within the Upper Jurassic, the Oxfordian-early Kimmeridgian, late Kimmeridgian-early Tithonian, Tithonian, and late Tithonian-early Berriasian radiolarian assemblages are distinguished. The late Berriasian-early Valanginian, middle-late Valanginian, and Hauterivian radiolarian assemblages are first recognized or compositionally revised. Radiolarians and lithofacies data are used to correlate the tectonostratigraphic units and individualize the jasper-alkali basaltic (lower Hettangian), chert-terrigenous (Hettangian-Sinemurian), jasper-cherty (Pliensbachian-Aalenian), jasper (Bajocian-Hauterivian), jasper-basaltic (upper Bajocian-Valanginian), Fe-Ti basaltic (upper Bajocian-Bathonian), tuffitejasper-basaltic (Bathonian-Hauterivian), and terrigenous-volcanogenic (Bajocian-Valanginian) sequences. The correlation results are extrapolated into other continental areas flanking the Pacific, i.e., to the western Kamchatka, northern and northwestern coastal areas of the Sea of Okhotsk, where the analogous radiolarian assemblages are characteristic of comparable allochthonous units of terrigenous-siliceous-volcanogenic sediments.     

New belemnites from Toarcian and Aalenian of Siberia

From the Jurassic (Toarcian and Aalenian) of the Lena river basin the following Belemnites are described as new: Sachsibeius mirus, Lenobelus lenensis, Lenobelus gravis, Lenobelus reconditus, Dactyloteuthis yacutiensis. Sachsibelus (type species S. mirus) and Lenobelus (type species L. lenensis) are new genera. The phylogenetic relationships of these genera are discussed. Sachsibelus is assigned to the Hastitinae; Lenobelus is provisionally placed in the same subfamily. --E. T. Tozer.     

Carbon and oxygen isotopes: a tool for Jurassic and early Cretaceous pelagic correlation (southern Spain)

The isotopic evolution of δ¹³C and δ¹⁸O is reported for the Jurassic and early Cretaceous in two pelagic sections of the External Zones in the Betic Cordilleras (SE Spain). Stable isotope curves from pelagic trough and swell sections display similar patterns. Variations in δ¹⁸O and δ¹³C values from strata at equivalent age probably reflect both early diagenetic cementation and later temperature‐related burial diagenesis. Comparison of global isotope curves with those presented in this work allows the differentiation of global from local events. Thus, the anoxic event during the early Toarcian (falciferum Zone) is characterized by elevated δ¹³C and depressed δ¹⁸O values. The isotopic record also allows the detection of the middle Oxfordian transgression. There are other peaks for the late Toarcian, early Bajocian, Callovian and early Berriasian that can also be used as tools for correlation purposes. Copyright © 2002 John Wiley & Sons, Ltd.     

Marine Early Bajocian Deposits of the Lower Volga Region (Volgograd Region) and Their Belemnite-Based Stratigraphy

A representative early Bajocian belemnite assemblage containing four genera and 11 species (two identified in open nomenclature and five new: Eocylindroteuthis weisi sp. nov., E. mariottii sp. nov., E. yarkovi sp. nov., Homaloteuthis volgogradensis sp. nov., and Hastites orphana sp. nov.) is described from a section near the Dubovoi hamlet in the Greater Don Bend area (southern termination of the Don-Medveditsa dislocations). Some members of this assemblage were previously known from the Caucasus; however, it is possible to state the discovery of a Euroboreal belemnite fauna completely new for Russia, previously described only from Central Europe and almost unknown for Eastern Europe. The age of the studied assemblage collected from the lower part of the section, traditionally considered to be Upper Bajocian, corresponds to the Laeviuscula ammonite Chronozone and is the first reliable evidence of marine settings in the Volga Region for the early Bajocian. Two new biostratigraphic units, Beds with Eocylindroteuthis weisi and Beds with H. orphana, are introduced. The former, judging from the occurrences of its characteristic species in Western Europe, has a broad correlative potential. On the basis of datings obtained, the scheme of the formational subdivision of the Middle Jurassic of the Volga Region near Volgograd is revised, and the formerly abandoned Bakhtemir Formation is reinstated as a valid unit. Its total range is reestablished as corresponding to the upper part of the Discites (?)/Laeviuscula Chronozone (lower Bajocian)–Garantiana Chronozone (upper Bajocian).     

A revision of some British Lower Bajocian stephanoceratid ammonites

Members of the ammonite family Stephanoceratidae from the Middle Jurassic, Lower Bajocian, Laeviuscula to Humphriesianum zones of South West England are revised. These comprise faunas resulting from periodic migrations and possibly hybridization. The taxonomic positions of Teloceras Mascke, 1907, Kumatostephanus Buckman, 1922, Gibbistephanus Buckman, 1928 and Pseudoteloceras Pavia and Fernández-López, 2016 are considered in the light of new research. A new subfamily Kumatostephaninae is erected and hypotheses are presented regarding the evolution of some Early Bajocian stephanoceratids. Pseudoteloceras digbyi sp. nov. represents the earliest ‘Teloceras-like’ morphology from the late Sauzei Zone. Refinement is made of the scheme of faunal horizons for the Lower Bajocian.     

Aalenian to Cenomanian Radiolaria of the Bermeja Complex (Puerto Rico) and Pacific origin of radiolarites on the Caribbean Plate

The study of the radiolarian ribbon chert is a key in determining the origins of associated Mesozoic oceanic terranes and may help to achieve a general agreement regarding the basic principles on the evolution of the Caribbean Plate. The Bermeja Complex of Puerto Rico, which contains serpentinized peridotite, altered basalt, amphibolite, and chert (Mariquita Chert Formation), is one of these crucial oceanic terranes. The radiolarian biochronology presented in this work is mainly based by correlation on the biozonations of Baumgartner et al. (1995) and O??Dogherty (1994) and indicates an early Middle Jurassic to early Late Cretaceous (late Bajocian?Cearly Callovian to late early Albian?Cearly middle Cenomanian) age. The illustrated assemblages contain about 120 species, of which one is new (Pantanellium karinae), and belonging to about 50 genera. A review of the previous radiolarian published works on the Mariquita Chert Formation and the results of this study suggest that this formation ranges in age from Middle Jurassic to early Late Cretaceous (late Aalenian to early?Cmiddle Cenomanian) and also reveal a possible feature of the Bermeja Complex, which is the younging of radiolarian cherts from north to south, evoking a polarity of accretion. On the basis of a currently exhaustive inventory of the radiolarite facies s.s. on the Caribbean Plate, a re-examination of the regional distribution of Middle Jurassic sediments associated with oceanic crust, and a paleoceanographic argumentation on the water currents, we come to the conclusion that the radiolarite and associated Mesozoic oceanic terranes of the Caribbean Plate are of Pacific origin. Eventually, a discussion on the origin of the cherts of the Mariquita Formation illustrated by Middle Jurassic to middle Cretaceous geodynamic models of the Pacific and Caribbean realms bring up the possibility that the rocks of the Bermeja Complex are remnants of two different oceans.