Local records of long-term dynamics of bamboo gregarious flowering in northern Laos and regional synchronicity of <Emphasis Type="Italic">Dendrocalamus membranaceus</Emphasis> in two flowering sites

Bamboos are widely distributed in mainland Southeast Asia and have abundant biomass. They are characterized by prolonged vegetative growth and semelparity. Where bamboos are dominant, their synchronous flowering and death has a major impact on forest vegetation. Although the small-scale dynamics of this process have become clearer in recent years, the history, geographical scale and synchronicity of bamboo flowering over broad areas remains unknown. This study focused on the flowering history of six bamboo species, Bambusa tulda, Cephalostachyum virgatum, Dendrocalamus hamiltonii, Dendrocalamus membranaceus, Indosasa sinica and Oxytenanthera parvifolia, over 40 years across a broad area of northern Laos. We also examined the synchronicity of flowering in D. membranaceus. We visited 49 villages in northern Laos and surveyed knowledgeable inhabitants about bamboo flowering history. The timing, scale and synchronicity of gregarious flowering varied among species. D. hamiltonii and D. membranaceus showed higher flowering synchronicity than other species. All the species except I. sinica had both sporadic and gregarious flowering traits, and showed conspicuous variability in their flowering scale. The flowering bamboo population at two gregarious flowering sites for D. membranaceus was surveyed. While this species had the highest synchronicity in this study, its synchronicity was lower than other species in previous studies worldwide. We found that the gregarious flowering of bamboos in northern Laos over the last 40 years showed lower synchronicity than bamboo flowering reported in other areas of the world. The historical dynamics and scale of bamboo flowering must be further clarified to understand the vegetation composition of this area.     

老挝北部Sentinel-2 A/B云覆盖适宜阈值界定及其影像获取概率时空差异

光学卫星影像云覆盖时空特征评估是衡量其作为重要遥感监测数据源的前提。Sentinel-2 A/B影像因其免费获取、多光谱（红边）、更高时空分辨率等优势,已在全球不同尺度陆面植被与生态监测中受到重视。相较于Landsat等同类影像产品,有关Sentinel-2 A/B的云覆盖分析还未见报道。本文利用2016—2018年老挝北部所有5288景Sentinel-2 A/B影像（Granule/Tile）的云覆盖元数据,基于不同云覆盖阈值（0~100%）水平下的影像获取概率差异确定了影像获取概率分析的云覆盖适宜阈值,并揭示了云量特征阈值水平下的影像获取概率时空差异。主要结论如下:① Sentinel-2 A/B影像获取概率分析云覆盖适宜特征阈值为20%（即云覆盖≤20%）,该阈值水平下老挝北部Sentinel-2 A/B影像的逐月累积平均获取概率最高（约27.41%）;② 在20%云覆盖阈值水平下,老挝北部Sentinel-2 A/B影像逐月累积平均获取概率差异在时间上与旱季（11月—次年4月）雨季（5月—10月）的时间分布较为吻合。旱季获取概率约为42.91%,3月概率（50.27%）最大,4月与2月次之,时间上与刀耕火种焚烧与橡胶林落叶特征吻合;雨季相应概率约为11.81%,6月最低（约1.26%）;③ 老挝北部Sentinel-2 A/B影像逐月累积平均获取概率在空间上存在东西差异,旱季西部省域单元（如琅南塔）影像获取概率远高于东部,雨季西部地区影像获取概率则略低于东部地区。本研究既可为后续开展大区及全球Sentinel-2 A/B影像云量分析提供借鉴,也对开展联合国减少森林砍伐和退化排放（UN-REDD）计划引发的土地利用变化如刀耕火种农业演变、橡胶林扩张等遥感监测有指导意义。     

Spatio-Temporal Variations in Co-Occurrence Patterns of Fish Communities in Haizhou Bay,China: Null Model Analysis

Co-occurrence pattern of fish species plays an important role in understanding the spatio-temporal structure and the stability of fish community. Species coexistence may vary with time and space. The co-occurrence patterns of fish species were examined using the C-score under fixed-fixed null model for fish communities in spring and autumn over different years in the Haizhou Bay, China. The results showed that fish assemblages in the whole bay had non-random patterns in spring and autumn over different years. However, the fish co-occurrence patterns were different for the northern and southern fish assemblages in spring and autumn. The northern fish assemblage showed structured pattern, whereas the southern assemblage were randomly assembled in spring. The co-occurrence patterns of fish communities were relatively stable over different years, and the number of significant species pairs in northern assemblage was more than that in the southern assemblage. Environmental heterogeneity played an important role in determining the distributions of fish species that formed significant species pairs, which might affect the co-occurrence patterns of northern and southern assemblages further in the Haizhou Bay.     

基于网格的街区尺度城市绿度度量方法

本文提出一种基于网格的街区尺度城市绿度度量方法。其以900 m×900 m的网格单元作为以城市主干道划分所形成街区尺度的表征,根据网格单元内植被覆盖度与植被构成之间的关系构建绿度指标。通过与传统的基于植被面积占比的格网法所计算的绿度指数进行比较。发现本文方法建立的绿度指标能在街区尺度上提供更加丰富的城市绿度信息,有效地解决了传统方法难以反映的植被覆盖度近似区域因其内部植被构成不同所带来的绿度差异问题。为城市规划、设计以及管理提供了一种新视角。基于该绿度指标,利用Landsat TM/OLI 2009、2015和2019年不同季节的遥感数据分析北京市城市绿化的时空变化特征。结果表明:北京市主城区城市绿化的面积和质量在2009—2019年显著提升,且秋季最为明显。绿化的变化在四环内外呈现出不同的模式。四环内主要为绿化面积的增加,四环外则在绿化面积和绿化质量上都有明显的增长。     

Regional-scale identification of three-dimensional pattern of vegetation landscapes

The altitudinal pattern of vegetation is usually identified by field surveys,however,these can only provide discrete data on a local mountain.Few studies identifying and analyzing the altitudinal vegetation pattern on a regional scale are available.This study selected central Inner Mongolia as the study area,presented a method for extracting vegetation patterns in altitudinal and horizontal directions.The data included a vegetation map at a 1∶1 000 000 scale and a digital elevation model at a 1∶250 000 scale.The three-dimensional vegetation pattern indicated the distribution probability for each vegetation type and the transition zones between different vegetation landscapes.From low to high elevations,there were five vegetation types in the southern mountain flanks,including the montane steppe,broad-leaved forest,coniferous mixed forest,montane dwarf-scrub and sub-alpine shrub-meadow.Correspondingly,only four vegetation types were found in the northern flanks,except for the montane steppe.This study could provide a general model for understanding the complexity and diversity of mountain environment and landscape.     

Understory vascular plant community assembly in relation to time-since-fire and environmental variables in a Chinese boreal forest

Understanding the response of understory vegetation to fire disturbance is vital to biodiversity conservation and management of boreal forests. We surveyed understory vascular plant richness and composition, and measured related environmental variables along a toposequence within three successional stages, initial (3 years post-fire), early (13 years post-fire) and late (>100 years post-fire) successional stages. Using permutation multivariate ANOVA and nonmetric multidimensional scaling, we analyzed how understory species richness and composition change as time-since-fire proceeds, and their correlative relationships with environmental variables. Species richness and composition showed significant differences among the three successional stages. Understory species richness and abundance were significantly associated with time-since-fire, topographic position, elevation and organic layer depth. Among these variables, time-since-fire had the strongest effect and topographic position was the second major factor on affecting understory community assembly. In addition, time-since-fire overwhelmed the effects of soil pH in the initial successional stage and gravimetric soil moisture in early and late successional stages on understory species composition     

Vegetation-environment relationship in conifer dominating forests of the mountainous range of Indus Kohistan in northern Pakistan

Environmental variables play a crucial role in shaping vegetation structure,mainly in mountainous ecosystems.Different studies have attempted to identify the environment-vegetation relationship of Conifer Dominating Forests(CDF) worldwide.However,due to differences in local climate and soil composition,different environmental drivers can be found.By applying multivariate analysis techniques,this study investigated the vegetation-environment relationship of CDF of Indus Kohistan in northern Pakistan.Our results showed that CDF of Indus Kohistan are distributed in five distinct ecological groups,which are dominated by different trees and understory species.A total of 7 trees and 71 understory species were recorded from the sampling sites.Cedrus deodara was the leading species among four groups,having the highest importance value(IV),density and basal area.Group I was dominated by Pinus wallichiana with the second highest importance value,density and basal area.In addition,elevation,slope,maximum water holding capacity(MWHC),soil moisture(SM),total organic matter(TOM),sodium,phosphorus and nickel showed highly significant influence on composition and distribution pattern of Indus Kohistan vegetation.Therefore,this study shows a new evidence of vegetation-environment relationship,pointing out specific drivers of vegetation structure in CDF of Indus Kohistan region in northern Pakistan.     

Effects of nitrogen deposition on tundra vegetation undergoing invasion by Deyeuxia angustifolia in Changbai Mountains

In recent years, herbaceous species such as Deyeuxia angustifolia(Kom.) Y. L. Chang has invaded alpine tundra regions of the western slope of the Changbai Mountains. Because atmospheric nitrogen deposition is predicted to increase under a warming climate and D. angustifolia is sensitive to nitrogen addition, field experiments were conducted from 2010 to 2013 to determine the effect of increased nitrogen deposition on the mechanisms of D. angustifolia invasion. The goal of this study is to evaluate the impact of increased nitrogen deposition on the changes in alpine tundra vegetation(consisting mostly of Rhododendron chrysanthum Pall. and Vaccinium uliginosum Linn.). The results showed that: 1) simulated nitrogen deposition affected overall characteristics and structure of R. chrysanthum and V. uliginosum communities and had a positive impact on the growth of tundra vegetation invaded by D. angustifolia; 2) R. chrysanthum was more resistant to invasion by D. angustifolia than V. uliginosum; 3) simulated nitrogen deposition could improve the growth and enhance the competitiveness of D. angustifolia, which was gradually replacing R. chrysanthum and V. uliginosum and might become the dominant species in the system in future, transforming alpine tundra into alpine meadow in the Changbai Mountains.     

Pollen-based reconstructions of Holocene vegetation and climatic change of Tibetan Plateau

A synthesis of Holocene pollen records from the Tibetan Plateau shows the history of vegetation and climatic changes during the Holocene. Palynological evidences from 24 cores/sections have been compiled and show that the vegetation shifted from subalpine/alpine conifer forest to subalpine/alpine evergreen sclerophyllous forest in the southeastern part of the plateau; from alpine steppe to alpine desert in the central, western and northern part; and from alpine meadow to alpine steppe in the eastern and southern plateau regions during the Holocene. These records show that increases in precipitation began about 9 ka from the southeast, and a wide ranging level of increased humidity developed over the entire of the plateau around 8-7 ka, followed by aridity from 6 ka and a continuous drying over the plateau after 4-3 ka. The changes in Holocene climates of the plateau can be interpreted qualitatively as a response to orbital forcing and its secondary effects on the Indian Monsoon which expanded northwards     

Changes in microbial community composition in the leaf litter of successional communities after volcanic eruptions of Mount Usu,northern Japan

Changes in the fungal and bacterial biomass and community structure in litter after the volcanic eruptions of Mount Usu, northern Japan were investigated using a chronosequence approach, which is widely used for analyzing vegetation succession. The vegetation changed from bare ground(10 years after the eruptions) with little plant cover and poor soil to monotonic grassland dominated by Polygonum sachalinense with undeveloped soil(33 years) and then to deciduous broad-leaved forest dominated by Populus maximowiczii with diverse species composition and well-developed soil(100 years). At three chronosequential sites, we evaluated the compositions of phospholipid fatty acids(PLFAs), carbon(C) and nitrogen(N) contents and the isotope ratios of C(δ13C) and N(δ15N) in the litter of two dominant species, Polygonum sachalinense and Populus maximowiczii. The C/N ratio, δ13C and δ15N in the litter of these two species were higher in the forest than that in the bare ground and grassland. The PLFAs gradually increased from the bare ground to the forest, showing that microbial biomass increased with the development of the soil and/or vegetation. The fungi-to-bacteria ratio of PLFA was constant at 5.3 ± 1.4 in all three sites, suggesting that fungi were predominant. A canonical correspondence analysis suggested that the PLFA composition was related tothe successional ages and the developing soil properties(P 0.05, ANOSIM). The chronosequential analysis effectively detected the successional changes in both microbial and plant communities.     

Alpine tundra species phenology is mostly driven by climate-related variables rather than by photoperiod

The study of plant phenology has frequently been used to link phenological events to various factors, such as temperature or photoperiod.In the high-alpine environment, proper timing of the phenological cycle has always been crucial to overcome harsh conditions and potential extreme events(i.e. spring frosts) but little is known about the response dynamics of the vegetation, which could shape the alpine landscape in a future of changing climate. Alpine tundra vegetation is composed by an array of species belonging to different phytosociological optima and with various survival strategies, and snowbed communities are a relevant expression of such an extreme-climate adapted flora.We set eight permanent plots with each one in a snowbed located on the Cimalegna plateau in Northwestern Italy and then we selected 10 most recurring species among our plots, all typical of the alpine tundra environment and classified in 3different pools: snowbed specialists, grassland species and rocky debris species. For 3 years we registered the phenophases of each species during the whole growing season using an adaptation of the BBCH scale. We later focused on the three most biologically relevant phenophases, i.e., flower buds visible, full flowering, and beginning of seed dispersion. Three important season-related variables were chosen to investigate their relationship with the phenological cycle of the studied species:(i) the Day Of Year(DOY),the progressive number of days starting from the 1 st of January, used as a proxy of photoperiod,(ii) Days From Snow Melt(DFSM), selected to include the relevance of the snow dynamics, and(iii) Growing Degree Days(GDD), computed as a thermal sum. Our analysis highlighted that phenological development correlated better with DFSM and GDD than with DOY.Indeed, models showed that DOY was always a worse predictor since it failed to overcome interannual variations, while DFSM and marginally GDD were better suited to predict the phenological development of most of the species, despite differences intemperature and snowmelt date among the three years. Even if the response pattern to the three variables was mainly consistent for all the species, the timing of their phenological response was different.Indeed, species such as Salix herbacea and Ranunculus glacialis were always earlier in the achievement of the phenophases, while Agrostis rupestris and Euphrasia minima developed later and the remaining species showed an intermediate behavior. However, we did not detect significant differences among the three functional pools of species.     

Plant Diversity Performance After Natural Restoration in Reclaimed Deyeuxia angustifolia Wetland

Deyeuxia angustifolia wetlands were widely distributed in the Sanjiang Plain in Northeast China. Due to strong demand for food production, large-area wetlands were reclaimed to farmlands, which threatened regional ecological security greatly. Since the 21 th century, returning farmlands to wetlands was widely adopted for natural restoration in the Sangjiang Plain. As the first reflection of wetland restoration, vegetation succession of restored D. angustifolia wetlands should be fully assessed. In this study, vegetation investigation was carried out in three restored D. angustifolia wetlands with 5, 8 and 12 yr restoration, respectively. Meanwhile, a natural D. angustifolia wetland was selected as reference wetland. Results showed that community composition changed greatly and there was visible community succession. Community dominant species changed from composite to gramineae as restoration time increasing.At first, weeds community appeared in the restored wetlands, especially the xerophytes developed to the pioneer species rapidly. And then, mesophytes and wetland species became the dominant species in the restored wetlands. Finally, wetland species, especially D. angustifolia, occupied the dominant position of restored community. Shannon-wiener index(H) and Simpson index(D) both decreased to close to natural D. angustifolia wetlands. Compared with natural D. angustifolia wetland, species composition and diversity in restored wetlands were more complex and higher. As restoration time increasing, there were not significant differences between community characteristics of restored wetlands and natural wetland. All these suggested that vegetation in reclaimed D. angustifolia wetland could be restored naturally, but its restored period is 10 yr at least. From another angle, it is important to protect current natural wetlands.     

中街山水域虾类组成及其群落多样性

根据2010~2011年中街山列岛水域4个季节的渔业资源调查资料,分析了该水域虾类资源密度和群落多样性,并探讨了底层温度、底层盐度和水深对虾类数量分布和多样性的影响.结果表明:共有虾类20种,隶属于9科18属;广温广盐种无论在种类数还是尾数密度与生物量均占主导地位;虾类生物量和尾数密度在季节分布上,以冬季最高,冬季均与春、夏和秋三季呈显著差异(p值<0.05);平面分布上,生物量与密度指数总体上呈南北高、中间低,东部高西部低的特征;生物量与平均底温显著相关(P<0.05),与平均底盐和水深相关性较差(p值>0.05);葛氏长臂虾、细巧仿对虾、中国毛虾、日本鼓虾和中华管鞭虾为四季的优势种,且均属广温广盐种;虾类群落Margalef丰富度指数为0.61~1.09,Shannon-Wiener多样性指数为0.76~1.31,Pielou均匀度指数为0.50~0.70,其中丰富度指数和多样性指数春季与冬季均存在显著差异(P<0.05),均匀度指数秋季与冬季存在显著差异(p值<0.05).     

老挝稀土资源特征及勘查开发前景

稀土是世界各国的战略性关键金属矿产资源,而中重稀土尤为重要。老挝境内分布着大量不同时代花岗岩类,且气候湿热多雨,风化作用强烈,为离子吸附型稀土矿的形成创造了有利条件。为查明老挝稀土资源成矿地质特征和资源分布特征,分析其投资开发前景,对整个老挝境内开展了大量系统的野外调研及相关分析测试工作,并且汇总了老挝稀土资源勘查工作进展。研究结果表明,老挝稀土资源成矿特征可类比中国南方地区,属于典型的风化壳淋积型矿床,矿体主要富集于花岗岩类风化壳全风化层,盖层厚度相对中国较大。矿体呈层状、似层状和透镜体状,厚度数米到二十多米。离子相稀土平均品位为0.4‰~0.7‰,大部分稀土配分模式同江西赣州信丰稀土矿相似,属富集中重稀土元素的中钇富铕型。老挝稀土资源主要分布于北部的川圹省和华潘省,预计两省稀土资源总量可达到60万t。随着老挝稀土政策的开放和中国稀土战略的逐步完善,投资开发老挝稀土资源,建立稀土资源的海外战略基地,可以长期稳定地支撑中国对离子吸附型稀土资源的需求,同时也可以为加强两国合作、共建中老命运共同体做出突出贡献。      

2001-2013年洞庭湖流域植被覆盖度时空变化特征

本文基于2001-2013年MODIS NDVI多时序数据,采用像元二分模型估算了洞庭湖流域植被覆盖度,分析了区域近13年来植被覆盖度的变化特征及趋势,并结合同期气象数据,阐明了植被覆盖度变化对气候因素的响应。结果表明：（1）近13年洞庭湖流域植被覆盖度的整体变化较为稳定,呈微弱减少趋势,速率为-0.3%/10a。（2）洞庭湖流域绝大部分区域植被覆盖状况良好,植被覆盖度呈自西向东递减趋势,高植被覆盖度及中高植被覆盖度占整个流域面积的88.63%,水体或低植被覆盖度及中低植被覆盖度仅占2.57%。（3）洞庭湖流域植被覆盖度变化趋势为北部强于南部、东部强于西部。流域内植被覆盖度极显著与显著减少的面积比例为5.30%、增加面积的比例为4.29%,植被覆盖度变化不显著占90.40%。该区域植被覆盖度变化受人为因素影响更大。     

Succession Features and Dynamic Simulation of Subalpine Forest in the Gongga Mountain, China

The Gongga Mountain of eastern Tibet Plateau is a representative of the alpine regions with high peaks and deep valleys. Climate change over the last thousand years has controlled the dynamics of glacier and debris flow occurrence, which resulted in substantial changes in the mountainous environment. The authors surveyed the community structure of primary forests in Gongga Mountain and forest successign processes in woodland plots. The changing features in the subalpine environment are discussed in this paper. Tree species and sizes between the glacier shrinking areas and debris flow fans in Hailuogou Valley are compared. The pioneer species that settle in debris flow fans and the glacier shrinking areas are Salix spp. and Populus purdomii. Abies fabri and Picea brachytyla are the climax tree species. The succession process of primary vegetation in Hailuogou (2700 ～ 3200 m) can be divided into four stages:     

长江三角洲城市化地区植被初级生产力的时空变化研究

城市化过程对植被初级生产具有重要影响。以往研究主要集中于城市用地扩张对植被初级生产力的直接影响分析,而较少关注其间接效果。本文以长江三角洲地区为例,分别从地区尺度和城市尺度分别分析了2000-2013年植被初级生产力的时空变化,探讨了其与气温、降水量及城市建成区绿化覆盖率的关系。研究表明：地区尺度上,2000-2013年长江三角洲植被初级生产力呈现不断增加,其中城市建成区植被初级生产力呈现显著增加的趋势（P<0.05）;城市尺度上,城市建成区内植被初级生产力主要呈现增加的趋势,而其外围缓冲区内则与此相反。在当前气候变化背景下,这可能与城市建成区绿化覆盖率不断增加,及快速的城市扩张有关。     

Soil microbial population dynamics along a chronosequence of moist evergreen broad-leaved forest succession in southwestern China

Little is known about whether soil microbial population dynamics are correlated with forest succession.To test the hypotheses that(1) soil microbial composition changes over successional stages,and(2) soil microbial diversity is positively correlated with plant species diversity,we determined the soil microbial populations,community composition,and microflora diversity in evergreen broad-leaved forests along a chronosequence of vegetation succession from 5 to 300 years in southwestern China.The soil microbi...     

Variation of Zooplankton Ecological Group During Spring in China Estuaries

Zooplankton was major indicator species of the environment. To explore the effect of marine environment on zooplankton distribution in the coast of China, the zooplankton samples from stations in Guanhe Estuary(GE), Changjiang(Yangtze River) Estuary(CE), Oujiang Estuary(OE), Jiulongjiang Estuary(JE) and Beilun Estuary(BE)(covering 14 latitudes) in spring were surveyed and the variation of zooplankton ecological group was researched. According to the adaptability temperature, the zooplankton was divided into two ecological groups: warm-temperate species and subtropical species. The warm-temperate species was the main dominant species and subtropical species was only dominant species in BE. Calanus sinicus, a warm-temperate species, was the only dominant species in all five estuaries. From north to south, the proportion of warm-temperate species in the five estuaries gradually decreased depends on the number, were 83.33%, 48.39%, 45.00%, 43.75% and 30.43%, respectively. In contrary, the proportion of subtropical species was gradually increased. The warm-temperate species predominated over the total abundance and the percentage was whopping high in the north estuaries, including GE(96.16%), CE(95.57%), OE(97.83%) and JE(95.53%). The abundance percentage of subtropical species have remarkably higher(82.39%) in BE. Five estuaries zooplankton community was subdivided into three zooplankton groups, which were the northern warm temperate zooplankton group, the southern subtropical zooplankton group and the estuarine brackish-water group. The water temperature and coastal current of China Sea are recognized as the main factors determining the distribution and community structure of estuarine zooplankton in spring.     

Sea surface height variations in the Mindanao Dome region in response to the northern tropical Pacific winds

Sea surface height (SSH) variability in the Mindanao Dome (MD) region is found to be one of the strong variations in the northern Pacific. It is only weaker than that in the Kuroshio Extension area, and is comparable to that in the North Pacific Subtropical Countercurrent region. Based on a 1.5-layer reduced gravity model, we analyzed SSH variations in this region and their responses to northern tropical Pacific winds. The average SSH anomaly in the region varies mainly on a seasonal scale, with significant periods of 0.5 and 1 year, ENSO time scale2-7years, and time scale in excess of 8 years. Annual and long-term variabilities are comparably stronger. These variations are essentially a response to the northern tropical Pacific winds. On seasonal and ENSO time scales, they are mainly caused by wind anomalies east of the region, which generate westward-propagating, long Rossby waves. On time scales longer than 8 years, they are mostly induced by local Ekman pumping. Long-term SSH variations in the MD region and their responses to local winds are examined and discussed for the first time .