Late quaternary vegetational change in the Kotzebue sound area, northwestern Alaska

Two sediment cores from Kaiyak and Squirrel lakes in northwestern Alaska yielded pollen records that date to ca. 39,000 and 27,000 yr B.P., respectively. Between 39,000 and 14,000 yr B.P., the vegetation around these lakes was dominated by Gramineae and Cyperaceae with some Salix and possibly Betula nana/glandulosa forming a local, shrub component of the vegetation. Betula pollen percentages increased about 14,000 yr B.P., indicating the presence of a birchdominated shrub tundra. Alnus pollen appeared at both sites between 9000 and 8000yr B.P., and Picea pollen (mostly P. mariana) arrived at Squirrel Lake about 5000 yr B.P. The current foresttundra mosaic around Squirrel Lake was established at this time, whereas shrub tundra existed near Kaiyak Lake throughout the Holocene. When compared to other pollen records from north-western North America, these cores (1) represent a meadow component of lowland. Beringian tundra between 39,000 and 14,000 yr B.P., (2) demonstrate an early Holocene arrival of Alnus in northwestern Alaska that predates most other Alnus horizons in northern Alaska or northwestern Canada, and (3) show an east-to-west migration of Picea across northern Alaska from 9000 to 5000 yr B.P.     

Late Pleistocene to Holocene vegetation and climate changes in northwestern Chukotka (Far East Russia) deduced from lakes Ilirney and Rauchuagytgyn pollen records

This paper presents two new pollen records and quantitative climate reconstructions from northern Chukotka documenting environmental changes over the last 27.9 ka. Open tundra- and steppe-like habitats dominated between 27.9 and 18.7 cal. ka BP. Betula and Alnus shrubs might have grown in sheltered microhabitats but disappeared after 18.7 cal. ka BP. Although the climate was rather harsh, local herb-dominated communities supported herbivores as is evident by the presence of coprophilous spores in the sediments. The increase in Salix and Cyperaceae ~16.1 cal. ka BP suggests climate amelioration. Shrub Betula appeared ~15.9 cal. ka BP, and became dominant after ~15.52 cal. ka BP, whilst typical steppe communities drastically reduced. Very high presence of Botryococcus in the Lateglacial sediments reflects widespread shallow habitats, probably due to lake level increase. Shrub Alnus became common after ~13 cal. ka BP reflecting further climate amelioration. Simultaneously, herb communities gradually decreased in the vegetation reaching a minimum ~11.8 cal. ka BP. A gradual decrease of algae remains suggests a reduction of shallow-water habitats. Shrubby and graminoid tundra was dominant ~11.8–11.1 cal. ka BP, later Salix stands significantly decreased. The forest-tundra ecotone established in the Early Holocene, shortly after 11.1 cal. ka BP. Low contents of green algae in the Early Holocene sediments likely reflect deeper aquatic conditions. The most favourable climate conditions were between ~10.6 and 7 cal. ka BP. Vegetation became similar to the modern after ~7 cal. ka BP but Pinus pumila came to the Ilirney area at about 1.2 cal. ka BP. It is important to emphasize that the study area provided refugia for Betula and Alnus during MIS 2. It is also notable that our records do not reflect evidence of Younger Dryas cooling, which is inconsistent with some regional environmental records but in good accordance with some others.     

Late Holocene δC and pollen records of paleosalinity from tidal marshes in the San Francisco Bay estuary, California

Records of stable carbon isotopes (δ¹³C) are presented from cores collected from four San Francisco Bay marshes and used as a proxy for changes in estuary salinity. The δ¹³C value of organic marsh sediments are a reflection of the relative proportion of C₃ vs. C₄ plants occupying the surface, and can thus be used as a proxy for vegetation change on the marsh surface. The four marshes included in this study are located along a natural salinity gradient that exists in the San Francisco Bay, and records of vegetation change at all four sites can be used to infer changes in overall estuary paleosalinity. The δ¹³C values complement pollen data from the same marsh sites producing a paleoclimate record for the late Holocene period in the San Francisco Bay estuary. The data indicate that there have been periods of higher-than-average salinity in the Bay estuary (reduced fresh water inflow), including 1600-1300 cal yr B.P., 1000-800 cal yr B.P., 300-200 cal yr B.P., and ca. A.D. 1950 to the present. Periods of lower-than-average salinity (increased fresh water inflow) occurred before 2000 cal yr B.P., from 1300 to 1200 cal yr B.P. and ca. 150 cal yr B.P. to A.D. 1950. A comparison of the timing of these events with records from the California coast, watershed, and beyond the larger drainage of the Bay reveals that the paleosalinity variations reflected regional precipitation.     

滇西北格贡错那卡湖沉积记录揭示的晚全新世气候变化

云南省西北部高山湖泊格贡错那卡湖（海拔4214 m）地处西南季风区,对气候变化十分敏感。文章以在格贡错那卡湖湖心钻取的1.84 m长的沉积岩芯为研究对象,对粒度、磁化率和总有机碳（TOC）含量等环境指标进行了高分辨率的分析,在精确测年的基础上（²¹⁰Pb测年结合陆生植物残体的AMS ¹⁴C测年建立年代序列）,重建了研究区距今约3570年来的气候与环境变化。结果表明,格贡错那卡湖所在地区近3570年来的气候与环境变化经历了5个较明显的演化阶段：1）在3570~3490 cal.a B.P.期间,气候明显降温、减湿;2）3490~1710 cal.a B.P.期间,气候总体偏冷、较干,其中3490~2860 cal.a B.P.期间气候处于较稳定的最冷干时期,随后在2860~1710 cal.a B.P.期间气候波动地有所增温、增湿;3）1710~930 cal.a B.P.（即240~1020A.D.）期间,为较稳定的暖湿期,年均温较高,温差较小;4）930~80 cal.a B.P.（即1020~1870A.D.）期间,气候总体偏冷较湿,但存在多次短暂变暖;降水总量或冰雪融水可能有所减少,但降水强度或冰雪融水强度变化较大;5）自80 cal.a B.P.（即1870 A.D.）以来,温度明显增加,可能是过去3570年以来温度最高的时期;降水有所减少,气候总体为暖偏湿。该气候特征与相邻地区泸沽湖的乔木花粉含量反映的气候变化非常一致,揭示了研究区气候变化的区域性。

     

Reconstruction of the Late Holocene climate and environmental history from North Bolgoda Lake,Sri Lanka,using lipid biomarkers and pollen records

The catastrophic impact and unpredictability of the Indian Ocean Monsoon (IOM) over South Asia are evident from devastating floods, mudslides and droughts in one of the most densely populated regions of the globe. However, our understanding as to how the IOM has varied in the past, as well as its impact on local environments, remains limited. This is particularly the case for Sri Lanka, where erosional landscapes have limited the availability of well-stratified, high-resolution terrestrial archives. Here, we present novel data from an undisturbed sediment core retrieved from the coastal Bolgoda Lake. This includes the presentation of a revised Late Holocene age model as well as an innovative combination of pollen, source-specific biomarkers, and compound-specific stable carbon isotopes of n-alkanes to reconstruct the shifts in precipitation, salinity and vegetation cover. Our record documents variable climate between 3000 years and the present, with arid conditions c. 2334 and 2067 cal a bp. This extreme dry period was preceded and followed by more wet conditions. The high-resolution palaeoenvironmental reconstruction fills a major gap in our knowledge on the ramifications of IOM shifts across South Asia and provides insights during a time of major redistribution of dense human settlements across Sri Lanka.     

A late quaternary pollen record from the Transvaal bushveld, South Africa

Pollen spectra from cores of organic spring deposits from the Transvaal provide evidence for the climatic evolution of the province during the last 35,000 yr B.P. or more. The past climatic phases are derived from palynological reconstructions of past vegetation types by comparison of fossil pollen data with modern surface pollen spectra from various localities. Evidence is provided for an early moist, cool phase with relatively mesic bushveld and expanded montane forest in the central Transvaal, followed by a drier period with drier bushveld which probably lasted until approximately 25,000 yr B.P. During the next phase, which at the latest ended about 11,000 yr B.P., the temperatures were probably 5°–6°C cooler than at present. At that time bushveld vegetation in the central Transvaal was replaced by open grassland with macchia elements. Climatic amelioration came and semiarid savanna returned to the plains, at first gradually and then developing into a warm Kalahari thornveld-type vegetation. After 6000 yr B.P. it apparently became slightly wetter and a more broad-leafed bushveld developed. About 4000 yr B.P. it again became cooler and slightly wetter and the bushveld vegetation on the central and northern plains was comparable to present open upland types. After 2000 yr B.P. conditions gradually became warner until about 1000 yr B.P., when the modern climate of the central Transvaal bushveld originated.     

Postglacial vegetation community change over an elevational gradient on the western Kenai Peninsula,Alaska: pollen records from Sunken Island and Choquette Lakes

The Kenai Peninsula of south‐central Alaska is a region of high topographic diversity with a complex glacial history. The sedimentary record of two small lakes [Sunken Island (SIL; 76 m a.s.l.) in the Kenai Lowlands; Choquette (CL; 527 m a.s.l.) in the Caribou Hills upland] exemplifies the postglacial development of the conifer–hardwood forest over an elevational range there. A herb–shrub tundra was established at both sites after deglaciation. By ~10.7 ka, poplar (Populus sp.) and alder (Alnus) dominated the lowland forest, while alder with minor poplar occurred at the upland site. Lake levels lower than today occurred during the early Holocene until ~8 ka. Subsequently at SIL, the near‐modern Kenai birch (Betula kenaica) – white spruce (Picea glauca) forest maintained prominence throughout the Holocene. However, at CL, alder dominated with dwarf birch and other subshrubs; small amounts of white spruce arrived ~5.2 ka. Black spruce (Picea mariana) grew around SIL by ~4 ka, but never gained prominence at CL. Fire, a prominent agent of disturbance in the Kenai Lowlands since ~8 ka, was essentially absent at the hardwood‐dominated upland site before ~6 ka, and rare thereafter. This suggests an important link between fire and spruce in Kenai forests.     

Late pleistocene paleoenvironmental studies from the Rocky Mountain Region: A comparison of pollen and insect fossil records

A regional paleoenvironmental reconstruction is presented for a north-south transect of late glacial pollen and insect fossil sites from the Rocky Mountain region. Altitudinal treelines were depressed by an average of 671 m and summer temperatures were an average of 5.4° lower. the interpretation of paleoenvironmental conditions based on insect fossil evidence tends to show stronger departures from modern parameters than that based on palynology. While clear evidence for late glacial occupation of the region by Paleoindians is scarce, montane conditions were not so severe as to preclude human occupation, at least on a seasonal basis. Newly discovered sites of early occupation would benefit greatly from inter-disciplinary studies.     

青藏高原西北部晚第四纪以来的隆升作用 ——来自西昆仑阿什库勒多级河流阶地的证据

青藏高原的隆升是新生代最壮观的地质事件,关于青藏高原隆升研究一直是地学界的研究焦点.河流阶地的发育记录了丰富的构造运动和气候变化的信息,近年来被广泛应用于构造运动和气候演变的研究,但前人研究的河流阶地基本分布在青藏高原的周缘,阶地的形成可能是气候与构造运动共同作用的结果.本文通过高分辨率卫星影像的解译,在青藏高原内部的西昆仑阿什库勒地区发现了多达七级的河流阶地.对该处河流阶地结构、沉积特征、几何特征的研究表明该阶地是典型的构造成因阶地.野外利用全站仪对河流阶地地貌形态进行了精细的测量,获得了各级阶地的拔河高度分别为4～5m(T1)、9～ 10m(T2)、16 ～ 18m(T3)、28～31m(T4)、45～48m(T5).通过宇宙成因核素10Be测年方法对各级阶地面的暴露年龄进行了测定,获得了各级阶地的形成时代分别为7.7±0.7ka(T1)、32.7±3.lka (T2)、53.6±2.5ka(T3)、115.7±23.2ka(T4)、166.8±10.4ka (T5)、19.5±8.5ka (T6).由此确定了晚第四纪166.8ka以来不同时期的河流下切速率总体介于0.2～0.35mm/yr,该速率代表了青藏高原西北部晚第四纪166.8ka以来的平均隆升速率.     

Late Pleistocene Vertebrates and Other Fossils from Epiguruk, Northwestern Alaska

Sediments exposed at Epiguruk, a large cutbank on the Kobuk River about 170 km inland from Kotzebue Sound, record multiple episodes of glacial-age alluviation followed by interstadial downcutting and formation of paleosols. Vertebrate remains from Epiguruk include mammoth, bison, caribou, an equid, a canid, arctic ground squirrel, lemmings, and voles. Radiocarbon ages of bone validated by concordant ages of peat and wood span the interval between about 37,000 and 14,000 yr B.P. The late Pleistocene pollen record is dominated by Cyperaceae, with Artemisia, Salix, Betula, and Gramineae also generally abundant. The fossil record from Epiguruk indicates that the Kobuk River valley supported tundra vegetation with abundant riparian willows during middle and late Wisconsin time. Large herbivores were present during the height of late Wisconsin glaciation as well as during its waning stage and the preceding interstadial interval. The Kobuk River valley would have been a favorable refugium for plants, animals, and possibly humans throughout the last glaciation.     

Late Pleistocene and Holocene glaciation of the Fish Lake valley,northeastern Alaska Range,Alaska

We reconstructed a chronology of glaciation spanning from the Late Pleistocene through the late Holocene for Fish Lake valley in the north‐eastern Alaska Range using ¹⁰Be surface exposure dating and lichenometry. After it attained its maximum late Wisconsin extent, the Fish Lake valley glacier began to retreat ca. 16.5 ka, and then experienced a readvance or standstill at 11.6 ± 0.3 ka. Evidence of the earliest Holocene glacial activity in the valley is a moraine immediately in front of Little Ice Age (LIA) moraines and is dated to 3.3–3.0 ka. A subsequent advance culminated at ca. AD 610–900 and several LIA moraine crests date to AD 1290, 1640, 1860 and 1910. Our results indicate that ¹⁰Be dating from high‐elevation sites can be used to help constrain late Holocene glacial histories in Alaska, even when other dating techniques are unavailable. Close agreement between ¹⁰Be and lichenometric ages reveal that ¹⁰Be ages on late Holocene moraines may be as accurate as other dating methods. Copyright © 2009 John Wiley & Sons, Ltd.     

Middle Pleistocene age of the Nome River glaciation, northwestern Alaska

During the middle Pleistocene Nome River glaciation of northwestern Alaska, glaciers covered an area an order of magnitude more extensive than during any subsequent glacial intervals. The age of the Nome River glaciation is constrained by laser-fusion ⁴⁰Ar/³⁹Ar analyses of basaltic lava that overlies Nome River drift at Minnie Creek, central Seward Peninsula, that average 470,000 ± 190,000 yr (±1σ). Milligram-size subsamples of the lava were dated to identify and eliminate extraneous ⁴⁰Ar enrichments that rendered the mean of conventional K---Ar dates on larger bulk samples of the same flow too old (700,000 ± 570,000 yr). While the ⁴⁰Ar/³⁹Ar analyses provide a minimum limiting age for the Nome River glaciation, maximum ages are provided by a provisional K---Ar date on a basaltic lava flow that underlies the Nome River drift at nearby Lave Creek, by paleomagnetic determinations of the drift itself at and near the type locality, and by amino acid epimerization analysis of molluscan fossils from nearshore sediments of the Anvilian marine transgression that underlie Nome River drift on the coastal plain at Nome. Taken together, the new age data indicate that the glaciation took place between 580,000 and 280,000 yr ago. The altitude of the Anvilian deposits suggests that eustatic sea level during the Anvilian transgression rose at least as high as and probably higher than during the last interglacial transgression; by correlation with the marine oxygen-isotope record, the transgression probably dates to stage 11 at 410,000 yr, and the Nome River glaciation is younger still. Analyses of floor altitudes of presumed Nome River cirques indicate that the Nome River regional snowline depression was at least twice that of the maximum late Wisconsin. The cause of the enhanced snowline lowering appears to be related to greater availability of moisture in northwestern Alaska during the middle Pleistocene.     

Vegetation and climate history in the Laptev Sea region (Arctic Siberia) during Late Quaternary inferred from pollen records

Paleoenvironmental records from a number of permafrost sections and lacustrine cores from the Laptev Sea region dated by several methods (¹⁴C-AMS, TL, IRSL, OSL and ²³⁰Th/U) were analyzed for pollen and palynomorphs. The records reveal the environmental history for the last ca 200 kyr. For interglacial pollen spectra, quantitative temperature values were estimated using the best modern analogue method. Sparse grass-sedge vegetation indicating arctic desert environmental conditions existed prior to 200 kyr ago. Dense, wet grass-sedge tundra habitats dominated during an interstadial ca 200–190 kyr ago, reflecting warmer and wetter summers than before. Sparser vegetation communities point to much more severe stadial conditions ca 190–130 kyr ago. Open grass and Artemisia communities with shrub stands (Alnus fruticosa, Salix, Betula nana) in more protected and moister places characterized the beginning of the Last Interglacial indicate climate conditions similar to present. Shrub tundra (Alnus fruticosa and Betula nana) dominated during the middle Eemian climatic optimum, when summer temperatures were 4–5 °C higher than today. Early-Weichselian sparse grass-sedge dominated vegetation indicates climate conditions colder and dryer than in the previous interval. Middle Weichselian Interstadial records indicate moister and warmer climate conditions, for example, in the interval 40–32 kyr BP Salix was present within dense, grass-sedge dominated vegetation. Sedge-grass-Artemisia-communities indicate that climate became cooler and drier after 30 kyr BP, and cold, dry conditions characterized the Late Weichselian, ca 26–16 kyr BP, when grass-dominated communities with Caryophyllaceae, Asteraceae, Cichoriaceae, Selaginella rupestris were present. From 16 to 12 kyr BP, grass-sedge communities with Caryophyllaceae, Asteraceae, and Cichoriaceae indicate climate was significantly warmer and moister than during the previous interval. The presence of Salix and Betula reflect temperatures about 4 °C higher than present at about 12–11 kyr BP, during the Allerød interval, but shrubs were absent in the Younger Dryas interval, pointing to a deterioration of climate conditions. Alnus fruticosa, Betula nana, Poaceae, and Cyperaceae dominate early Holocene spectra. Reconstructed absolute temperature values were substantially warmer than present (up to 12 °C). Shrubs gradually disappeared from coastal areas after 7.6 kyr BP when vegetation cover became similar to modern. A comparison of proxy-based paleoenvironmental reconstructions with the simulations performed by an Earth system model of intermediate complexity (CLIMBER-2) show good accordance between the regional paleodata and model simulations, especially for the warmer intervals.     

川西北地区晚三叠世卡尼期洪水事件沉积的碳、氧同位素记录*

川西北地区绵竹和安县等地发育的下卡尼阶岩性以灰色鲕粒灰岩、海绵礁丘粘结及障积成因的凝块石灰岩为主,上卡尼阶岩性突变为灰黑色页岩、粉砂岩,这与西特提斯地区卡尼阶在岩性和颜色上有很大的相似性。本次研究用激光微区碳、氧同位素测试方法分析了川西北地区卡尼阶中有孔虫、双壳类及腕足类化石壳体和植物叶片化石的碳、氧同位素,结果显示上卡尼阶化石中氧同位素值较下卡尼阶略有变小,上卡尼阶泥质粉砂岩中化石的碳同位素较下卡尼阶灰岩明显变小,可能反映了淡水输入的影响。结合研究区卡尼阶的沉积学特征,笔者认为川西北地区同样受到卡尼期洪水事件的强烈影响,潮湿气候条件下的淡水输入是造成川西北地区海绵礁（丘）大量周期性死亡及最后消失的影响因素之一。     

Paleoecology of late-glacial peats from the bering land bridge, Chukchi Sea shelf region, northwestern Alaska

Insect fossils and pollen from late Pleistocene nonmarine peat layers were recovered from cores from the shelf region of the Chukchi Sea at depths of about 50 m below sea level. The peats date to 11,300−11,000 yr B.P. and provide a limiting age for the regional Pleistocene-Holocene marine transgression. The insect fossils are indicative of arctic coastal habitats like those of the Mackenzie Delta region (mean July TEMPERATURES = 10.6–14°C) suggesting that 11,000 yr ago the exposed Chukchi Sea shelf had a climate substantially warmer than modern coastal regions of the Alaskan north slope. The pollen spectra are consistent with the age assignment to the Birch Interval (14,000–9000 yr B.P.). The data suggest a meadow-like graminoid tundra with birch shrubs and some willow shrubs growing in sheltered areas.     

Late Pleistocene(MIS 2) climate change on the Northern Tibetan Plateau inferred from pollen and geochemical records from Lake Gomo Co

正Introduction A 405-cm sediment section GM02 from Gomo Co salt lake in the northern Tibetan Plateau was used to reconstruct climate changes during marine isotope stage(MIS)2,including the Last Glacial Maximum(LGM)and last     

Long-term river discharge and multidecadal climate variability inferred from varved sediments, southwest Alaska

Sedimentological analyses of 289 years (AD 1718-2006) of varved sediment from Shadow Bay, southwest Alaska, were used to investigate hydroclimate variability during and prior to the instrumental period. Varve thicknesses relate most strongly to total annual discharge (r² = 0.75, n = 43, p < 0.0001). Maximum annual grain size depends most strongly on maximum spring daily discharge (r² = 0.63, n = 43, p < 0.0001) and maximum annual daily discharge (r² = 0.61, n = 43, p < 0.0001), while varve thickness is poorly correlated with maximum annual grain size (r² = 0.004, n = 287, p = 0.33). Relations between varve thickness and annual climate variables (temperature, precipitation, North Pacific (NP) and Pacific Decadal Oscillation (PDO) indices) are insignificant. On multidecadal timescales, however, regime shifts in varve thickness and total annual discharge coincide with shifts in NP and PDO indices. Periods with increased varve thickness and total annual discharge were associated with warm PDO phases and a strengthened Aleutian Low. The varve-inferred record of PDO suggests that any periodicity in the PDO varied over time, and that the early 19th century marked a transition to a more frequent or detectable shifts.     

Late quaternary history of Lake Manitoba,Canada

The postglacial history of Lake Manitoba has been deduced from a study of the changes in physical, mineralogical, and chemical variables in sediment cores collected from the lake. Six lithostratigraphic units are recognized in the South Basin of the lake. Weakly developed pedogenic zones, reflecting dry or extremely low water conditions in the basin, separate five of these six units. The initial phase of lacustrine sedimentation in the Lake Manitoba basin began shortly after 12,000 yr B.P. as water was impounded in front of the receding glacier to form Lake Agassiz. By 11,000 yr ago, continued retreat of the ice sheet opened lower outlets to the east and much of Lake Agassiz drained, including the Lake Manitoba basin. Water levels again rose at 9900 yr B.P., but by about 9200 yr B.P. the South Basin was again dry. For the next 4700 yr there was an alternation of wet and dry conditions in the basin in response to the interaction of a warmer and drier climate and differential crustal rebound of the basin. About 4500 yr ago a new phase of Lake Manitoba sedimentation was initiated when the Assiniboine River began to discharge into the South Basin. The Assiniboine River was diverted out of the Lake Manitoba watershed about 2200 yr ago. Erosion and redistribution of the sandy deltaic sediments deposited by the Assiniboine River has created the barrier beach that now separates the extensive marsh to the south of the lake from the main lake.     

广西那艺泥盆系五指山组下段地层研究

从广西那艺五指山组下段识别出三个晚泥盆世牙形石标准带，自上而下为：混杂上ｃｒｅｐｉｄａ带，混杂中ｃｒｅｐｉｄａ带至ｔｒｉａｎｇｕｌａｒｉｓ带，下ｔｒｉａｎｇｕｌａｒｉｓ带。该段含不少再沉积化石，岩性以灰质拟角砾岩为主夹瘤状灰岩，从而确定其为碳酸盐浊积岩，浊积的时代为法门期ｔｒｉａｎｇｕｌａｒｉｓ带至上ｃｒｅｐｉｄａ带。该段属陆壳上的盆地相，靠近台地斜坡，至少存在两种类型的浊积岩。另外，本剖面尚发现大量微球粒。     

Stratigraphy,pollen analysis,and paleoclimatic interpretation of Pulbeena Swamp,northwestern Tasmania

Sedimentary, palynologic, and ¹⁴C analysis of 480 cm of freshwater marl and swamp-peat deposits, formed under the influence of fluctuating artesian springs, provides a paleoenvironmental and paleoclimatic record of approximately 65,000 yr for northwestern Tasmania.The Holocene (Pollen Zone 1, 11,000-0 yr B.P.) climate was warm and moist, and forest vegetation was dominant throughout the area. During the later part of the last glacial stage (Pollen Zone 2, 35,000–11,000 yr B.P.) the climate was generally drier, and grassy open environments were widespread. The driest part of this period occurred between 25,000 to 11,000 yr B.P., when temperatures in western Tasmania were markedly reduced during the last major phase of glaciation. Prior to 35,000 yr B.P. (Pollen Zones 3–9) a long “interstadial complex” dating to the middle of the last glacial stage is recognized. During this period the climate was generally moist, and forest and scrub communities were more important than during the later part of the last glacial stage, except during Pollen Zone 5 when high Gramineae plus Compositae values suggest drier conditions. High Gramineae and Compositae values also occur in Pollen Zone 10 at the base of the diagram. They suggest that a phase of drier and cooler climatic conditions occurred during the early part of the last glacial stage.