Stratigraphy and paleobathymetric interpretation of the Cretaceous Karai shale formation of Uttatur Group, TamilNadu, India

The Karai shale Formation of the Uttatur Group is exposed in a bad land area at the western margin of the Cauvery Basin. This shale has been investigated based on foraminiferal fauna and clay minerals. The foraminiferal assemblages obtained contain predominantly calcareous benthic foraminifera, rare planktic and arenaceous foraminifera. The planktic foraminiferal index taxa Planomalina buxtorfi, Rotalipora reicheli, Praeglobotruncana stephani, and Hedbergella portsdownensis suggest the late Albian to middle Turonian age. The benthic assemblage dominated by Lenticulina, Gavelinella, Osangularia and Quadrimorphina, suggests an outer neritic (100–200 m) environment. The clay mineral content dominated by kaolinite-illite-montmorillonite indicates that the Karai shale was formed from weathering of igneous rocks.     

Palaeoecological significance of turritelline gastropod-dominated assemblages from the mid-Cretaceous (Albian-Cenomanian) of Texas and Oklahoma,USA

The faunas of three previously poorly known and highly fossiliferous limestones from the upper Lower Cretaceous of Texas are dominated by turritelline gastropods. These faunas consist of turritelline-dominated assemblages in the Whitestone Limestone Member of the Walnut Formation in Travis County (middle Albian), the Keys Valley Marl Member of the Walnut Formation in Coryell County (middle Albian), and the Fort Terrett Formation in Kimble County (middle Albian). A fourth high-spired gastropod assemblage in the Segovia Formation in Pecos County (upper Albian) is not dominated by turritellines. Two other turritelline-dominated assemblages in non-carbonate rocks from the Albian and Cenomanian of Texas and Oklahoma are also described. These turritelline-dominated assemblage occurrences add considerably to our knowledge of the facies occurrence of Cretaceous turritelline-dominated assemblages, and they are consistent with the global facies distribution of these assemblages: i.e., although they are widespread in siliciclastic facies from Cretaceous to Recent, turritelline-dominated assemblages in carbonate facies occur almost exclusively in the Cretaceous and Paleogene.     

The Albian fern flora of Alexander Island,Antarctica

The Albian Alexander Island macrofossil flora from the Antarctic Peninsula preserves a diverse community of liverworts (Marchantiophyta), ferns (Polypodiopsida), Lycopodiales, Equisetales, Cycadales, Ginkgoales, seed-ferns (Bennettitales and Pentoxylales), Coniferales, and the first representatives of angiospermous leaves in Antarctica. Despite the presence of angiosperms in this assemblage, ferns are the most diverse element of the flora and are also ecologically dominant, while angiosperms contribute a smaller component to floristic diversity and have low abundance. Here we describe 11 fern taxa from this assemblage. The fossils are assigned to Cladophlebis, Sphenopteris and two newly created genera. The new genera and species are described under Adiantitophyllum serratum gen. et. sp. nov. and Nunatakia alexanderensis gen. et. sp. nov., and the new species are recognized as Cladophlebis dissecta sp. nov., Cladophlebis drinnanii sp. nov., Cladophlebis macloughlinii sp. nov. and Sphenopteris sinuosa sp. nov. In total, there are 24 fern species known from Alexander Island. In comparison to older floras (Jurassic) there is a greater diversity of ferns, while latest Cretaceous floras preserve significantly fewer fern species and more angiosperms. Possible factors that might account for such high fern diversity are high rainfall or generally humid conditions, regular disturbances by flooding and occasionally fire, and the preservation of a diverse range of fern communities that represent several palaeoenvironments.     

Plant assemblage from Opole,southern Poland: New data on Late Cretaceous vegetation of the northern part of the European Province

This paper presents the results of studies on plant macroremains found in the upper Turonian of the Folwark Quarry, Opole, Poland, associated with palynological studies of the host rocks. In addition to a few macrofossils (gymnosperm wood, conifer Geinitzia reichenbachii and fern ?Didymosorus) rich sporomorphs (bryophyte, lycopod and fern spores, conifer and angiosperm pollen grains) and marine palynomorphs (mainly dinoflagellate cysts) were recorded. The palynological analysis revealed that the vegetation on the neighbouring land (the East Sudetic Island) in the late Turonian was much more diverse than could be reconstructed based on only macrofossil remains. The latter are taxonomically restricted and dominated by one gymnosperm species (Geinitzia reichenbachii), which make them similar to most neighbouring, coeval Central European assemblages. Its over representation is, thus, a result of taphonomy.     

Late Pleistocene vegetation of Kings Canyon,Sierra Nevada,California

Seven packrat midden samples make possible a comparison between the modern and late Pleistocene vegetation in Kings Canyon on the western side of the southern Sierra Nevada. One modern sample contains macrofossils and pollen derived from the present-day oak-chaparral vegetation. Macrofossils from the six late Pleistocene samples record a mixed coniferous forest dominated by the xerophytic conifers Juniperus occidentalis, Pinus cf. ponderosa, and P. monophylla. The pollen spectra of these Pleistocene middens are dominated by Pinus sp., Taxodiaceae-Cupressaceae-Taxaceae (TCT), and Artemisia sp. Mesophytic conifers are represented by low macrofossil concentrations. Sequoiadendron giganteum is represented by a few pollen grains in the full glacial. Edaphic control and snow dispersal are the most likely causes of these mixed assemblages.The dominant macrofossils record a more xeric plant community than those that now occur on similar substrates at higher elevations or latitudes in the Sierra Nevada. These assemblages suggest that late Wisconsin climates were cold with mean annual precipitation not necessarily greater than modern values. This conclusion supports a model of low summer ablation allowing for the persistence of the glaciers at higher elevations during the late Wisconsin. The records in these middens also suggest that S. giganteum grew at lower elevations along the western side of the range and that P. monophylla was more widely distributed in cismontane California during the Pleistocene.     

Transport, sorting, and reworking of late Wisconsinan plant macrofossils from Lake Erie, Canada

Stratigraphic, sedimentological, and plant macrofossil studies on proglacial lacustrine and deltaic sequences dating from the Mackinaw Interstadial on the north shore of Lake Erie, Canada, indicate long-distance transport, selective sorting and reworking of the plant fossils. Transport, sorting, and reworking may make detailed paleoclimatic indices irrelevant and may account for incongruent phytogeographic indicators in the fossil assemblage. Details of the stratigraphic and sedimentological setting of the study site and the state of preservation of the fossils should be considered more carefully as aids to the interpretation of macrofossil assemblages.     

Charophyte biostratigraphy of the non-marine Lower Cretaceous in the Central Tunisian Atlas (North Africa): Palaeobiogeographic implications

The chronostratigraphic framework of the non-marine deposits of the Central Tunisian Lower Cretaceous (Kebar Formation) is reviewed from a biostratigraphic viewpoint. The outcrops located in the Jebel Kebar, Jebel Ksaïra and Jebel Koumine localities provided charophyte assemblages belonging to two biochronozones: Ascidiella cruciata-Pseudoglobator paucibracteatus (upper Barremian–lower Aptian) and Clavator grovesii lusitanicus (upper Aptian–lower Albian). Clavatoraceans from the upper Barremian–lower Aptian in the Tunisian Atlas are reported here for the first time. The assemblage consists of Atopochara trivolvis var. triquetra, Ascidiella iberica var. inflata, Globator maillardii var. trochiliscoides, Globator maillardii var. biutricularis, Echinochara peckii var. lazarii, Clavator harrisii var. harrisii and Clavator harrisii var. reyi. In addition, a new characean species, Mesochara magna nov. sp. Trabelsi and Martín-Closas, is described herein. The results show that the Kebar Formation is diachronous in Central Tunisia, with a more complete record to the north (Jebel Koumine) than in the type locality (Jebel Kebar). Barremian–Aptian diapiric activity is proposed as the factor that controls the diachronous nature of this formation.The late Barremian–early Aptian charophyte assemblages from the Kebar Formation display strong affinities with the contemporaneous floras of the European basins, thus suggesting that intense supraregional floristic exchanges occurred between the Tethyan islands scattered throughout what is now Western Europe and North Africa. The biogeographic distribution of these charophytes leads to the hypothesis that the peri-Tethyan Archipelago acted as an effective bridge for the intercontinental exchanges of these plants between Laurasia and Gondwana.     

Association of macro- and microfossils in the Vendian (Ediacaran) postglacial successions in Western Mongolia

The Vendian (Ediacaran) beds of the Zavkhan Basin, in the upper part of the Tsagaanolom Formation (<632 ± 14 Ma), yielded a new “Zavkhan” association of algae, microfossils, and problematic organisms, which is established in the series of alternating chert-carbonate shale with phosphorite interbeds. This association is distinct in the predomination of large (250 μm and over) sphaeromorphic microfossils of the genera Tasmanites, Archaeooides, and Leiosphaeridia, whereas acanthomorph acritarchs are represented by rarely found Cavaspina sp. and Tanarium sp. Multicellular algae included fragments of encrusting or foliate thalli with pseudoparenchymatous structure of polygonal cells characteristic of Rhodophyta algae (Thallophycoides sp.), and cordlike thalli of Vendotaenid algae Tyrasotaenia podolica. These layers of siltstone contain imprints of the problematic Vendian macrofossil Beltanelliformis brunsae. In their stratigraphic position, chemostratigraphic data, and fossil assemblage, the “Zavkhan” association can be assigned to the Upper Vendian.     

Plant remains from the Lower Cretaceous Fossil-Lagerstätte of Pietraroja,Benevento, southern Italy

Plant macrofossils with epidermal features from the Lower Albian (Lower Cretaceous) Plattenkalk 2 as exposed in “Le Cavere” quarry within Pietraroja Fossil-Lagerstätte, southern Italy, include sterile foliage-bearing shoots and reproductive structures of gymnosperms and possible angiosperm leaves occurring in a controversial marine depositional environment. The following taxa are recorded: Brachyphyllum sp., Frenelopsis sp., Cheirolepidiaceae gen. et sp. indet. and Nageiopsis?. Bituminous strata of the lower Plattenkalk 2 are full of plant debris composed mainly of isolated leaves and sterile axes. Taphonomic considerations suggest parautochtonous deposition for Frenelopsis while the other plant remains are allochthonous. Among the plants studied, the cheirolepidiaceous conifers have ecological significance owing to their (debated) xeromorphic foliar features. These adaptations suggest a warm and dry, or possibly coastal palaeoenvironment. Studied plant fossils together with sedimentological and palaeozoological considerations do not exclude a lagoonal hypothesis for Pietraroja sedimentary basin. This research expands the phytotaxonomic knowledge of the emergent lands within the Early Cretaceous Apenninic Carbonate Platform (ACP).     

Early Cretaceous megaspore assemblages from southeastern Australia

Twenty-nine megaspore species including six new taxa (Bacutriletes otwayensis sp. nov.,Erlansonisporites cerebrus sp. nov., Erlansonisporites decisum sp. nov., Hughesisporites coronatus sp. nov., Hughesisporites dettmanniae sp. nov., and Verrutriletes depressus sp. nov.) are documented from Aptian and Albian strata of the Gippsland and Otway basins, southeastern Australia. Together with six taxa known only from underlying Neocomian strata, these megaspores are used to establish four provisional biozones for the Lower Cretaceous that complement existing biostratigraphic schemes based on miospores and plant macrofossils. Megaspores are best represented in silty floodbasin facies and it is likely that the parent plants predominantly occupied moist understorey to fully aquatic habitats on the floodplain. Megaspores are sparsely represented in most other fluvial facies chiefly due to reworking of floodbasin sediments into higher energy channel and crevasse deposits. The relatively high diversity of lycophyte and fern megaspores contrasts with the scarcity of these plant groups in macrofossil assemblages. The megaspore record suggests that heterosporous cryptogams may have been significantly more prominent in the vegetation of this region than previously suggested. Several megaspores from southeastern Australia are closely comparable to forms from India and Argentina indicating broad similarities between Early Cretaceous heterosporous fern and lycophyte communities across Gondwana. These similarities also suggest that megaspores may be useful for inter-continental biostratigraphic correlation.     

Climate characteristics of the eastern Mongolian Plateau,China during the early Early Cretaceous (145–132 Ma): Palynological evidence from the Tongbomiao Formation in Well Hong-6, Hailar Basin

This study identified two palynological assemblages, namely Bayanhuasporites-Cycadopites-Protoconiferus and Cicatricosisporites-Cedripites-Perinopollenites, in the Tongbomiao Formation in the Hongqi Sag in the Hailar Basin, Inner Mongolia, China for the first time. The former is distributed in the lower part of the Tongbomiao Formation and is characterized by abundant gymnosperm pollen and diverse fern spores. Among them, the gymnosperm pollen is dominated by Paleoconifer (4.98%–31.62%) and Cycadopite (8.55%–25.23%) pollen grains and also includes other pollen grains such as Classopollis, Parcisporites, Erlianpollis, Callialasporites, and Jiaohepollis. The fern spores in the former palynological assemblage contain Bayanhuasporite (0–8.96%), Granulatisporites (0.93%–6.97%), and some important Cretaceous genera, such as Cicatricosisporites, Concavissimisporites, Densoisporites, Hsuisporites, Foraminisporis, and Leptolepidites. The Cicatricosisporites-Cedripites-Perinopollenites palynological assemblage is distributed in the upper part of the Tongbomiao Formation. Gymnosperm (77.30%), Pinaceae (31.9%), and Paleoconiferus (19.02%) pollen predominate this palynological assemblage, and Quadraeculina, Erlianpollis, and Jiaohepollis pollen are also common in this assemblage. The fern spores in this palynological assemblage include abundant Cicatricosisporites (4.29%). Besides, Concavissimisporites, Aequitriradites, and Leptolepidites are also common in this palynological assemblage. No angiosperm pollen has been found in both palynological assemblages. The identification of both palynological assemblages provides important evidence for the biostratigraphic correlation between the Hailar Basin and its adjacent areas. It also enables the reconstructions of the Berriasian-Valanginian (Early Cretaceous) vegetation and the paleoclimate on the eastern Mongolian Plateau during 141–132 Ma. The vegetation reconstructed on the palynological data of the represented by Hailar Basin in eastern Mongolian Plateau (141.6–141.4 Ma), form conifer forest or conifer broad-leaved mixed forest to conifer forest with shrubs and grassland, the climate belongs to warm temperate and warm-subtropicalt, the highest temperature is estimated to reach 35–38°C. Form 132.3 Ma, the vegetation type is conifer forest, and its paleoclimate is sub-humid warm temperate, the highest temperature is estimated to reach 24–29°C.©2022 China Geology Editorial Office.     

Late Aptian angiosperm pollen grains from Patagonia: Earliest steps in flowering plant evolution at middle latitudes in southern South America

The angiosperm pollen record from the Anfiteatro de Ticó and Punta del Barco formations (Baqueró Group) is reported. The relevance of these floras is that they are accurately dated as late Aptian, and one of the oldest floras southern South America with fossil angiosperms. Twelve samples were studied, showing Clavatipollenites and Retimonocolpites as dominant types. A new species, Jushingipollis ticoensis sp. nov., is proposed. A doubtful angiosperm pollen grain, Lethomasites sp., is also identified and described. A multivariate analysis of similarities between different Early Cretaceous angiosperm pollen assemblages suggests that the Baqueró Group assemblages have great similarities with other coeval units from Argentina, Australia and United States, which were located in a similar paleolatitude.     

Late Cretaceous Kholokhovchan Flora of Northeastern Asia: Composition,age and fossil plant descriptions

The Kholokhovchan Flora comes from tuffaceous – terrigenous deposits of the Vetvinskaya Member (Chalbugchan Group) in the Penzhina and Oklan rivers interfluve, Northeastern Russia. The depositional environment of the plant-bearing deposits is interpreted to have been a freshwater lake. The Kholokhovchan Flora hosts 42 fossil plant species belonging to Marchantiopsida, Polypodiopsida, Ginkgoales, Leptostrobales, Bennettitales, Pinales and Magnoliopsida. It is characterised by diverse angiosperms, less diverse conifers and ferns, by the presence of relatively ancient Sphenobaiera, Phoenicopsis and Pterophyllum together with advanced Late Cretaceous Taxodium, Glyptostrobus and angiosperms, among which platanoids are quite diverse. The Kholokhovchan Flora is most similar to Penzhina and Kaivayam floras of the Anadyr-Koryak Subregion and Arman Flora of the Okhotsk-Chukotka volcanogenic belt (Northeastern Russia) and should be dated as Turonian–Coniacian. The Kholokhovchan Flora, that populated volcanic plateaus and intermontane valleys, are characterised by a mixture of ancient “Mesophytic” plants with typical Late Cretaceous “Cenophytic” taxa. This peculiar composition probably reflects a gradual penetration of new angiosperm-dominated plant assemblages into older floras: during the Late Cretaceous, “Cenophytic” assemblages migrated along river valleys and other disturbed habitats into the interior of Asia, eventually occupying volcanogenic uplands, and in places replacing the “Mesophytic” fern-gymnospermous communities that existed there. Two new angiosperm species, as well as four the most characteristic conifers of the Kholokhovchan Flora, are described: Cupressaceae gen. et sp. indet. cf. Widdringtonites sp., Taxodium cf. olrikii, Taxodium sp., Glyptostrobus sp., Ettingshausenia vetviensis sp. nov. and Parvileguminophyllum penzhinense sp. nov.     

Late Wisconsinan Vegetation and Environment of the Tunica Hills Region, Louisiana/Mississippi

Pollen, plant macrofossil, and radiocarbon-dating studies of seven exposures of fluvial sediments in the Tunica Hills region of southeastern Louisiana and southwestern Mississippi provide new information on late Wisconsinan vegetation, flora, and environment of the region. The assemblages date between 25,250 and 17,530 yr B.P. Pollen and macrofossil assemblages are dominated by Picea, which comprises 40-70% of the pollen assemblages. Abies and Larix pollen and macrofossils are absent, in contrast to sites to the north in the central Mississippi Valley. Deciduous hardwoods (Quercus, Fagus, Fraxinus, Carya, Juglans nigra, Acer, Ulmus) are minor components of both pollen and macrofossil assemblages. Radiocarbon dates of Picea and Quercus wood indicate that these two genera grew contemporaneously in the region. Regional upland forests were dominated by Picea. Picea cones and cone fragments are not typical of any extant North American species, and probably represent either an extinct species or an extinct variety or subspecies of Picea glauca. Late Wisconsinan climate of the region was cooler than present, but not necessarily as cool as implied by P. glauca or other "boreal" taxa.     

Smaller foraminifers from type sections of the Bolorian stage,the Lower Permian of Darvaz

Smaller foraminifers from Upper Yakhtashian and Bolorian deposits of the stratotype area (Pamir, Darvaz, Tajikistan) are investigated. Four assemblages are defined. The first assemblage is from Chalaroschwagerina vulgaris-Pamirina darvasica Zone. The second assemblage found in the transitional Yakhtashian-Bolorian beds includes Globivalvulina, Palaeotextulariidae, Hemigordiidae, and Glomospira, associated with the first Pachyphloia and Langella forms. Characteristic taxa of third assemblage from the Misellina (Brevaxina) dyhrenfurthi Zone are Geinitzinidae, Globivalvulina, Palaeotextulariidae, Glomospira, and rare Pachyphloia. The forth assemblage of Hemigordiidae, Pachyphloia, Palaeotextulariidae, Geinitzinidae, Pseudoagathammina is identified in the M. (Misellina) parvicostata Zone. The assemblages were compared with concurrent analogs from China, Japan, and Russia. New species and subspecies Glomospira paleograndis sp. nov., G. darvasica sp. nov., Agathammina darvasica sp. nov., Pachyphloia darvasica sp. nov., Nodosinelloides cubanicus elongatus subsp. nov., and Hemigordius saranensis darvasicus subsp. nov. are described.     

Micropaleontological record of Holocene estuarine stages in the Bahía Blanca estuary,Argentina

Holocene paleoenvironmental changes have been interpreted on the basis of benthic foraminifera and calcareous nannofossils recovered in samples from Napostá Grande Stream, Bahía Blanca estuary, southern Buenos Aires Province. Samples are fine sands and clay sediments from a Holocene outcrop and were studied with quantitative techniques. The benthic foraminiferal assemblage is dominated by Ammonia parkinsoniana, Ammonia tepida, Bolivina pseudoplicata, Bolivina striatula, Bolivina sp., Buccella peruviana, and Elphidium spp. The calcareous nannofossil assemblage recovered is a typical cold-water association, dominated by Calcidiscus leptoporus, Coccolithus pelagicus, Emiliana huxleyi and Gephyrocapsa oceanica. A dendrogram classification by cluster analysis was made for each microfossil group. The results of these analyses were coincident, showing a liaison between changes in the assemblages of benthic foraminifera and calcareous nannofossils. Those results, jointly with the sedimentological information, lead to the identification of three different paleoenvironments along the Napostá N1 site. The lower part of the succession represents an estuarine environment with larger marine connection. The middle part represents a gradual passage to a more restricted estuarine environment, and the upper part represents the establishment of the modern continental fresh-water environment.     

Age and palynological characteristic of the Dongning Formation (Eastern Heilongjiang,China)

The taxonomic composition of the palynological assemblage of the Dongning Formation is supplemented. The palynological assemblage corresponds to those from the Lipovtsy Formation (Aptian) in the Razdolnaya Basin of Primorye (Russia) and the Muling Formation (Aptian) in the Jixi Basin of eastern Heilongjiang (China). It is found that the age of the Dongning Formation is Aptian. The palynological assemblage is characterized by dominance of spores of Gleicheniaceae; they are accompanied by spores of Cyatheaceae. The most important feature of the palynological assemblage of the Dongning Formation is the presence of angiosperm pollen (Tricolpites sp., T. micromunus, T. vulgaris, Retitricolpites georgiensis, R. vulgaris, Clavatipollenites hughesii, Quercites sparsus, Fraxiniopollenites variabilis).     

The palynology and geology of the Lower Cretaceous (Aptian–Albian) of Munday’s Hill Quarry,Bedfordshire, UK

Early Cretaceous sediments of Aptian–Albian age outcrop at Munday’s Hill Quarry, Bedfordshire, England. Previous papers describing the section have resulted in different terminologies being applied. The Lower Cretaceous in Bedfordshire is represented by sediments belonging to the Lower Greensand Group and the Gault Clay Formation. Within the Lower Greensand Group in the study area the Woburn Sands Formation, are of Aptian–Albian age. Selected samples have been analysed for palynology. The analysis reveals diverse palynomorph assemblages, including well-preserved dinoflagellate cysts and sporomorphs. Comparison of the assemblages with published records indicates that the lower samples are of Late Aptian age. Forms recorded include common Kiokansium unituberculatum, Cerbia tabulata, Aptea polymorpha and Cyclonephelium inconspicuum. An Early Albian age is indicated for the uppermost sample.     

Chondrichthyans from the Menuha Formation (Late Cretaceous: Santonian–Early Campanian) of the Makhtesh Ramon region,southern Israel

Exposures of the Menuha Formation (Santonian–Early Campanian, Mount Scopus Group) in the Makhtesh Ramon region of the southern Negev have produced numerous chondrichthyan teeth. The isolated teeth represent at least ten different species: Cretalamna appendiculata, Cretoxyrhina mantelli, Squalicorax falcatus?, S. kaupi, Scapanorhynchus rapax, S. raphiodon?, Carcharias samhammeri, Carcharias cf. C. holmdelensis?, and two other fish (Hadrodus priscus and a pycnodont). This assemblage has important implications for Late Cretaceous chondrichthyan palaeobiogeography. The majority of teeth were contained within a glauconite-rich, yellow-brown, soft chalk that included oysters (Pycnodonte vesicularis?), trace fossils (Planolites, Thalassinoides, and Chondrites), phosphatic peloids, and foraminiferans (globigerinids). The teeth were collected mainly through surface-sampling and sieving. The Menuha Formation probably represents a temperate to subtropical, shallow, open-shelf environment deposited during the formation of the Ramon anticline. Reworked conglomeratic chalks in the western section represent marginal facies derived from this structural uplift. With little to no published material describing the chondrichthyan fauna of the Menuha Formation, these data improve interpretations of its palaeoenvironment. Interpretation of the palaeoenvironment of the formation is important for understanding the larger stratigraphic/tectonic framework of the Ramon monocline region of southern Israel.     

Sedimentology of the Madumabisa Mudstone Formation (Late Permian), Lower Karoo Group,mid-Zambezi Valley Basin,southern Zambia

Sediments of the Upper Carboniferous to Lower Jurassic Karoo Supergroup (∼ 4.5 km thick) were deposited in the mid-Zambezi Valley Basin, southern Zambia. The Upper Palæozoic Lower Karoo Group in this area ends with a Late Permian sedimentary unit called the Madumabisa Mudstone Formation. The formation is 700 m thick and comprises four lithofacies grouped into two facies assemblages, collectively interpreted as lacustrine deposits. Sediments of a massive mudrock facies assemblage were deposited from suspension, probably from sediment-laden rivers entering a lake. Concretionary calcilutite beds probably mark the positions of palæosediment-water interfaces where calcite was precipitated. A laminated mudrock facies assemblage is attributed to lacustrine deposition from inflowing rivers at the lake margins and shallow parts of the lake. Repeated thickening-upward cycles are evidence of upward shallowing, interrupted by events of more abrupt deepening. Sandstone interbeds are interpreted as fluvial deposits laid down during low lake stands, with cross-lamination and asymmetrical ripples indicating current rather than wave deposition. A fossil assemblage of ostracods, bivalves, gastropods, fish scales, the alga Botryococcus sp. and fossil burrows is consistent with a lacustrine origin for the formation.