Paleoenvironmental and biostratigraphic significance of uvigerinids and other foraminifera from the Bhuban Formation,Assam-Arakan Basin,Mizoram

Planktic and benthic foraminifera including uvigerinids are documented from the Upper Bhuban Formation, exposed at Thingdawl village, Kolasib district, Mizoram. The foraminiferal assemblage is poorly preserved and consists of index fossils useful for precise biochronology and interpretation of the depositional environment. A total of ten benthic and six planktic foraminiferal species are described. Six species belong to the genus Uvigerina, four to Ammonia, one species each of Globorotalia, Globigerinoides, Clavatorella, Praeorbulina and two species of the genus Orbulina. Based on lithological and foraminiferal assemblage, middle neritic to upper part of outer neritic paleobathymetry is inferred for the deposition of this part of the Upper Bhuban Formation. The foraminiferal assemblage suggests late Early Miocene to early Middle Miocene, (∼16 Ma) equivalent to planktic foraminiferal zones N8-N9 for the Upper Bhuban Formation.     

Selective feeding of the oystercrassostrea gigas fed on a natural microphytobenthos assemblage

In estuarine ecosystems, microphytobenthos resuspended by tidal currents often represents a large part of the food supply available to bivalves which feed selectively. This study investigated the feeding behavior of the Pacific oysterCrassostrea gigas (Thunberg) relative to a natural microphytobenthic assemblage and the effect on the structure of this assemblage. Oysters were fed only benthic microalgae collected on the intertidal mud flats of Bourgneuf Bay (France) at a suspended particulate matter concentration above the threshold of pseudofeces production. All species in the assemblage were endemic diatoms characteritic of tidal mud flat environments. Four dominant ones, which were all solitary cells with spear-shaped frustules [Navicula ammophila (Grunow),Navicula rostellata (Kützing),Plagiotropis lepidoptera (Kuntze), andStaurophora amphioxys (Mann)] represented more than 95% of the 16 species involved. Analysis of feeding processes showed that the retention of the four main diatoms was not significantly different, but that two species, the smallest (N. ammophila [22×4 μm]) and the largest (P. lepidoptera [60×15 μm]), were preferentially ingested. The study of post-ingestive selection revealed that these two species were also preferentially digeted, i.e., preferentially directed into the digestive diverticulum, when they passed through the gut ofC. gigas. Cell size and shape did not appear to account for pre-ingestive and post-ingestive selection. The composition of the assemblage was significantly modified by oyster filtration, although the retention rates of the four main species were not significantly different. The composition of microalgae in pseudofeces and feces as a result of pre-ingestive and post-ingestive selection differed from that in seawater.     

Distribution of foraminifera and ostracods in the Onkaparinga Estuary,South Australia

Analysis of sediment and aquatic vegetation samples collected along the Onkaparinga estuary, South Australia, revealed the distribution patterns of benthic and epi-benthic micro-organisms, specifically foraminifera and ostracods. The total assemblage provided an assemblage ‘snap shot,’ contemporaneous with the environmental conditions at the time of sampling, as well as seasonally and time-averaged distribution information. Species richness is low in the upper reaches of the estuary and favours species with a wide salinity tolerance. Observed species richness increases seawards. Species richness and abundance within species is greater where there is conspicuous aquatic vegetation. Thus, species richness and distribution appear to be related to the proximity to the sea and the provision of diverse habitable substrate. Foraminiferal species, Ammonia beccarii and Elphidium excavatum, and ostracods belonging to the genera Paracypria, Xestolebris and Leptocythere dominate the total assemblage. These species are characteristic of euryhaline conditions. The mutual maximum occurrences of Reophax barwonensis, Ammobaculites barwonensis and Trochammina inflata signify mid-estuarine conditions. In the lower regions of the estuary, the appearance of shallow marine species of foraminifera such as Elphidium crispum and Quinqueloculina poeyana, and ostracods Bairdoppilata sp., Hemicytherura spp. and Paranesidea spp. signifies interactions with the adjacent inner shelf coastal waters. Throughout the entire length of the estuary, the presence of vegetal substrate created a strong numerical bias towards live occurrences of the ostracods Paracypria sp. and Xestolebris cedunaensis. These species are potentially useful as proxies for paleoenvironmental interpretations of estuarine sediments.     

The Influx of the Tsushima Current into the Central Ulleung Basin of East Sea (Sea of Japan), Korea

A total of 85 samples, collected from the UBGH1-9 core taken from the Ulleung Basin, East Sea, Korea, were analyzed using diatom assemblages. 111 diatom species belonging to 46 genera were identified, and three diatom assemblage zones were established on the basis of occurrence and distribution pattern of diatoms. Diatom assemblage zone I(134.10–174 m) is characterized by a relatively high abundance of marine species, while the increased number of the brackish species is recorded in diatom assemblage zone II(75–125 m). The assemblage zones IIIa became drastic drop of valve abundances and brackish planktons, whereas it became increase during the IIIb. High Td values which indicate an influence of warm current are recorded both in diatom assemblage zone I and III, and low Td values in diatom assemblage zone II. Analysis of diatom assemblages indicating that the depositional condition moved from oceanic to littoral-neritic environments and that paleotemperature underwent a shift from warm to cold condition at the middle interval, and from cold to warm condition in the upper interval of the UBGH1-9 core. This suggests that the lower(130–162 m) and upper intervals(0–20 m) of the UBGH1-9 core were deposited in the warm current condition(Tsushima Warm Current).     

山东诸城晚白垩世孢粉组合的发现及其地质意义

山东晚白垩世沉积以往未见孢粉化石。近年,通过对山东诸城侯家屯钻井岩心的研究,发现了丰富的孢粉化石,为确定地层时代和恢复古气候提供了有力的依据。其孢粉组成如下:     

Rhaxellid sponge microscleres from the Portlandian of Dorset,UK

Three previously described species and two new species of rhaxellid microscleres, Rhaxella winspitensis sp. nov. and R. elongata sp. nov., are described from cherts collected from the Cherty Series of the Portlandian of Dorset. The classification of Rhaxella is reviewed, retaining the genus within the family Geodiidae Gray. The results of this study suggest that three time-dependent rhaxellid assemblages can be recognized which, despite being restricted to certain depositional environments, may have biostratigraphic applications. A Portlandian assemblage described here is species-rich in comparison with known Oxfordian-Kimmeridgian assemblages. Bajocian-Callovian assemblages contain only R. sorbyana. The increased faunal diversity with time is speculatively described by an evolving Rhaxella lineage.     

The Darriwilian acritarch assemblage from Ordovician deposits of the Arkhangelsk Oblast, the northern Russian Plate

Borehole 2506 drilled in the northern area of the Arkhangelsk Oblast penetrated through the Paleozoic sedimentary block isolated in the Vendian thick sequence. A diverse acritarch assemblage has been established within the depth interval of 119.9–217.5 m. The assemblage comprises more than 70 taxa, including species characteristic of the boundary interval between the Volkhov and Kunda horizons of the East European Platform (the graptolite Didymograptus hirundo Zone). Stratigraphic position of host deposits was established within the Darriwilian Stage of the Middle Ordovician. The described assemblage of microphytofossils is similar to coeval assemblages from NW Russia, Baltic region, and Scandinavia, being typical of the Baltic phytoplankton province of temperate latitudes. A great number of species in common suggests that the assemblage under consideration is correlative with coeval assemblages of southern China thus offering a possibility of remote correlation.     

Disperse megaspores in the Lower Visean deposits of the Kaluga oblast (1P/A Aleksandrovskaya borehole)

Disperse megaspores were first found in the Lower Visean deposits of the Kaluga region (1P/A borehole). A layered distribution of the megaspores in the studied section is shown. A large number of megaspores were established in the lower part of the section of the Bobrikovskian deposits. Higher along the section, there is a trend to a decrease in the number of megaspores until they disappear completely in the Tula deposits. Two megaspore assemblages were identified. A great variety of species was observed in the lower assemblage; the dominant megaspores are Zonalesporites brasserti and Aphanozonatisporites triglobatus. The only dominant species was recorded in the upper assemblage, Zonalesporites brasserti, with a low species diversity of the entire assemblage. The miospore and megaspore assemblages that are typical of the Lower Visean in the section of the analyzed borehole are compared.     

The upper Cenomanian (Cretaceous) ammonite Neolobites vibrayeanus () in the Middle East: taxonomic and palaeoecologic remarks

In the Middle East and Africa, the Cenomanian ammonite genus Neolobites occurs, partly in great abundance, in shallow marine shelf settings. The genus includes several species but it has remained uncertain to what extent these species reflect biospecies, chronospecies or morphological variants. Based on material from Jordan and Egypt, the morphological variation of Neolobites vibrayeanus is described and discussed. From the several species of Neolobites described in the literature, only N. vibrayeanus, N. fourtaui and N. peroni appear to deserve specific separation. Sedimentological and lithological data, notably the common occurrence in strata that were possibly deposited above storm wave base, suggest that Neolobites was able to inhabit shallow marine settings. In these settings, it seems to form a distinct assemblage with the nautiloid Angulithes. Although N. vibrayeanus shows morphological variability between distinct populations, local N. vibrayeanus assemblages seem to develop morphologically stable populations that show only a little morphological overlap with those of other areas. This may be the reason for the earlier taxonomic splitting. The occurrence in habitats in which other stratigraphically significant ammonites are often missing may be the reason for the previous poor dating of Neolobites assemblages.     

Palaeobiology of Mesoproterozoic Salkhan Limestone, Semri Group, Rohtas, Bihar, India: Systematics and significance

Mesoproterozoic (∼ 1600 Ma old) Salkhan Limestone (Semri Group) of the Vindhyan Supergroup, exposed in Rohtas district of Bihar, India, preserves an abundant and varied ancient microbial assemblage. These microfossils are recorded in three distinctly occurring cherts viz., bedded chert, stromatolitic chert and cherty stromatolites. 27 morphoforms belonging to 14 genera and 21 species have been recognized. Six unnamed forms are also described. The microbial assemblage, almost exclusively composed of the remnants of cyanobacteria, is dominated by entophysalidacean members and short trichomes and can be termed as ‘typical Mesoproterozoic microbiotas’. The assemblage includes characteristic mat-forming scytonematacean and entophysalidacean cyanobacteria.Eoentophysalis is the dominant organism in the assemblage. Ellipsoidal akinetes of nostocalean cyanobacteria(Archaeollipsoides) and spherical unicells also occur; both are distinct from mat forming assemblage, allochthonous and possibly planktic. Co-occurrence of the microbiotas and precipitates is related to the depositional environment of the Mesoproterozoic tidal flats with high carbonate saturation.     

Environmental influences on the demersal fish assemblages in the Sepetiba Bay, Brazil

The structure of the demersal fish assemblage in Sepetiba Bay, southeast Brazil (22°54′–23°04′S, 43°34′–44°10′W) was described and related to environmental variables. A 3-yr long trawl survey (from July 1993 to June 1996) was conducted using a stratified random sampling strategy at 158 stations. One hundred and seven species of fishes in 80 genera and 44 families were identified. Gerreidae and Ariidae made up 30.5% and 28.6%, and 19.8% and 28.1% of the total number and weight, respectively. Sciaenidae, Engraulididae, Sparidae, and Carangidae were other prominent families. Three assemblages existed in the bay: one inhabited shallow, less saline and less transparent water in the inner zone (depth <5 m, transparency <2 m, salinity <30 psu, and slightly higher temperature); another in the outer zone with the opposite situation (depth >10 m, transparency >3 m, salinity >30 psu, and slightly lower temperature); and a third assemblage in the middle zone likely to prefer average values or show no clear preferences. Most species decreased in abundance over the 3-yr period of study. No seasonal change was detected in the assemblage structure, but many species were rare, indicating high spatial and temporal patchiness. Canonical correlation analyses were used to describe and compare the fish assemblages in the three different habitats. The outer zone had the greatest number of species recorded, lowest abundance, highest diversity and evenness, and was comprised mainly byCynossion leiarchus, Prionotus punctatus, Anchoa tricolor, Haemulon steindachneri, Diplectrum radiale, Etropus crossotus, andSphoeroides greeleyi; the inner zone had the lowest number of species recorded but the greatest number of species per sample, highest abundance, lowest diversity and evenness, being comprised mainly byDiapterus rhombeus, Genidens genidens, Cathrops spixii, Chloroschombrus chrysurus, Sciadeichthyes luniscutis, Cetengraulis edentulus, andArchorsargus rhomboidalis; while the third group had the lowest number of species per sample, being comprised by species widely distributed asGerres aprion, G. genidens, Gerres gula, andMicropogonias furnieri. Depth, followed by transparency and salinity, were the primary factors influencing assemblage distribution.     

黔北务川—正安—道真地区下二叠统大竹园组和梁山组孢粉学研究

文中材料采自黔北务川—正安—道真地区下二叠统大竹园组和梁山组。大竹园组孢粉化石类型单调,保存较差;而梁山组孢粉化石丰富,保存良好。孢粉组合特征为:(1)大竹园组包含丰富的Calamospora和Florinites及少量Torispora verrucosa,Gulisporites,Alisporites,Hamiapollenites和Striatopodocarpites,这一孢粉组合主要出现在上石炭统至下二叠统。按照大竹园组层位和孢粉化石特征,其地质时代应为早二叠世Asselian期—Artinskian期(相当于紫松期—隆林期)。(2)梁山组中可建立孢粉组合Laevigatosporites vulgaris-Gulisporites-Florinites ovalis,共计发现孢粉化石34属49种,其中蕨类植物孢子含量为51.6％～56.1％,裸子植物花粉含量为43.9％～48.4％。与大竹园组沉积时期相比,这一时期植被极为繁盛。根据孢粉组合特征,梁山组地质时代可能为早二叠世隆林期,即Kungurian期。     

Larger foraminifera distribution and strontium isotope stratigraphy of the La Cova limestones (Coniacian–Santonian, “Serra del Montsec”, Pyrenees, NE Spain)

The Upper Cretaceous La Cova limestones (southern Pyrenees, Spain) host a rich and diverse larger foraminiferal fauna, which represents the first diversification of K-strategists after the mass extinction at the Cenomanian–Turonian boundary.The stratigraphic distribution of the main taxa of larger foraminifera defines two assemblages. The first assemblage is characterised by the first appearance of lacazinids (Pseudolacazina loeblichi) and meandropsinids (Eofallotia simplex), by the large agglutinated Montsechiana montsechiensis, and by several species of complex rotalids (Rotorbinella campaniola, Iberorotalia reicheli, Orbitokhatina wondersmitti and Calcarinella schaubi). The second assemblage is defined by the appearance of Lacazina pyrenaica, Palandrosina taxyae and Martiguesia cyclamminiformis.A late Coniacian-early Santonian age was so far accepted for the La Cova limestones, based on indirect correlation with deep-water facies bearing planktic foraminifers of the Dicarinella concavata zone. Strontium isotope stratigraphy, based on many samples of pristine biotic calcite of rudists and ostreids, indicates that the La Cova limestones span from the early Coniacian to the early-middle Santonian boundary. The first assemblage of larger foraminifera appears very close to the early-middle Coniacian boundary and reaches its full diversity by the middle Coniacian. The originations defining the second assemblage are dated as earliest Santonian: they represent important bioevents to define the Coniacian-Santonian boundary in the shallow-water facies of the South Pyrenean province.By means of the calibration of strontium isotope stratigraphy to the Geological Time Scale, the larger foraminiferal assemblages of the La Cova limestones can be correlated to the standard biozonal scheme of ammonites, planktonic foraminifers and calcareous nannoplankton. This correlation is a first step toward a larger foraminifera standard biozonation for Upper Cretaceous carbonate platform facies.     

Mollusks from the lower quaternary Chauda sediments of the Black Sea

The analysis of paleontological remains in many samples from the Lower Quaternary Chauda sediments drilled and cored on the Bulgarian shelf of the Black Sea revealed widespread mollusks of the genera Didacna and Dreissena (Didacna tschaudae guriana, D. tschaudae, D. pleistopleura, D. crassa guriensis, Dreissena rostriformis tschaudae, D. rostriformis abchasica) accompanied by reworked Neogene representatives of the genus Digressodacna. In numerous places of a continental slope and an adjacent deep-sea depressions near Crimea and Caucasus the similar mollusc assemblage is described in the redeposited state for the first time. The composition of palynological spectra and diatom assemblages in shelf sediments indicates climate changes during the Chaudan period. The Chaudan mollusk fauna from the Black Sea sediments, which is lacking Caspian Bakuan species characteristic of the Chauda stratotype, is compositionally close to the mollusk assemblage from basal layers of the Chauda Horizon in the Guriya area of Georgia (“Gurian” Chauda). These data imply repeated changes in the level of the Chaudan basin between present-day isobaths of −30...−50 to −140 m.     

Age of the Vastan lignite in context of some oldest cenozoic fossil mammals from India

A highly diverse assemblage of mammal and other continental vertebrate fossils is reported in recent years from Vastan lignite mine (21°25′47″ N; 73°07′30″ E). Its importance in testing palaeobiogeographic hypothesis of mammalian dispersal in to or out of India has necessitated a definite age of the sequence. Nummulites burdigalensis is the only age diagnostic foraminifer in the section. This species is widespread in the Indian sedimentary basins and used to mark Early Eocene strata. In the present study two subspecies of Nummulites burdigalensis, viz., Nummulites burdigalensis burdigalensis and Nummulites burdigalensis kuepperi, are distinguished which led to further biostratigraphic refinement. Their concurrence in the section implies basal Cuisian age and shallow benthic zone SBZ 10 (most likely the lower part). The examination of this species from Kutch indicates the occurrence of evolutionarily advanced subspecies Nummulites burdigalensis cantabricus of middle Cuisian age (SBZ 11). Contrary to some recent reports, the study rules out the occurrence of N. globulus or any other Ilerdian foraminifer from the mine section.     

<Emphasis Type="Italic">Guembelitria</Emphasis> (foraminifera) in the upper cretaceous-lower paleocene succession of the Langpar Formation,India and its paleoenvironmental implication

Guembelitria is an essential biotic component in the Late Cretaceous-Early Paleogene (K/Pg) marine successions to provide crucial information about the K/Pg boundary; however, it is not well studied in Indian subcontinent. Biostratigraphically well constrained K/Pg successions of Therriaghat and Mahadeo in the East Khasi Hills District, Meghalaya, India provided a scope to present a comprehensive account of the genus in the perspective of differences of opinion about its species. A total of six species including Guembelitria langparensis n. sp., are recognized and their stratigraphic distribution is recorded. A review of the taxonomic validity of the known species, an evaluation of the diverse concepts of Guembelitria cretacea Cushman, and semiquantitative analysis of the recovered species permitted their clustering into two morphogroups that had different ecology. Morphogroup I comprising Guembelitria cretacea, Guembelitria trifolia and Guembelitria langparensis n. sp., is characterized by flared and short spire tests that form a bioseries and thrived as surface floaters. Morphogroup II comprising Guembelitria irregularis, Guembelitria danica and Guembelitria sp a, possesses high spire, narrow tests that have morphological abnormalities and had preference for living in subsurface stressed environment. The occurrence of the species and accompanying other features like ratio of planktonic to benthic foraminifera (p/b), lithologic assemblage and incidence of phosphorite, are used to infer depositional environments and sea level changes during successive biozones of the sequence. The recognized Guembelitria events during the K/Pg transition are discussed with reference to world occurrences.     

What have natural and human changes wrought on the foraminifera of San Francisco Bay late Quaternary estuaries?

In this study we compare the foraminifera of modern South San Francisco Bay with fossils from sediments of a previous estuary at 125 ka to provide a basis for interpreting the impact of natural and human change on the benthic ecosystem. All the species found in the Pleistocene sediments of this study are estuarine and/or shallow-water species occurring commonly in San Francisco Bay today, except for the introduced foraminifer Trochammina hadai, a native of Japan that was not found in samples taken in San Francisco Bay before 1983. The biodiversity and species composition of the fossil and modern assemblages before the introduction of T. hadai are nearly identical, suggesting that the environmental and physical changes in the 125,000-year-old and modern estuaries have not had a significant effect on the meiofauna of the Bay. In contrast, modern anthropogenic change in the form of species introductions has impacted the modern foraminiferal assemblage: T. hadai began to dominate the modern assemblage a decade after its introduction. Similar to the recorded impacts of introductions of marine metazoan invertebrate species, the dominance of T. hadai changed species proportions in the post-1980s foraminiferal assemblage, however no known extinctions in the native foraminiferal fauna occurred.     

Conodont-based stratigraphy of the Mosolovian Regional Stage (Eifelian,Middle Devonian) of the Voronezh Anteclise

Conodonts from the deposits of the Mosolovian Regional Stage (Middle Devonian) of the Voronezh Anteclise are studied on the basis of the data from six boreholes. Three assemblages are recognized. The index species of the lowermost assemblage is Icriodus formosus Naz. The index species of the middle assemblage are I. formosus Naz. and Pseudobipennatus ziegleri Kon. et Kim, as well as I. arkonensis Stauff. and I. jejunus Naz. This assemblage is the most diverse and is recognized in all boreholes. The deposits containing this assemblage correspond to the maximum of the Nara transgression. The upper assemblage is also recognized in all the studied boreholes. Its index species is Ps. ziegleri Kon. et Kim. The Mosolovian Regional Stage is assigned to the kockelianus Zone. The distribution diagrams of species in the most diverse middle assemblage support the hypothesis that, in the southeastern regions of the Voronezh anteclise, the depth was greater than in its western part. The characteristic conodont species are shown in the plates.     

Oceanic events and biotic effects of the Cenomanian-Turonian anoxic event, Tarfaya Basin, Morocco

Profound biotic changes accompanied the late Cenomanian δ¹³C excursion and OAE2 in planktic foraminifera in the Tarfaya Basin of Morocco. Planktic foraminifera experienced a severe turnover, though no mass extinction, beginning with the rapid δ¹³C excursion and accelerating with the influx of oxic bottom waters during the first peak and trough of the excursion. Species extinctions equaled the number of evolving species, though only the disaster opportunists Guembelitria and Hedbergella thrived along with a low oxygen tolerant benthic assemblage. The succeeding δ¹³C plateau and organic-rich black shale deposition marks the anoxic event and maximum biotic stress accompanied by a prolonged drop in diversity to just two species, the dominant (80–90%) low oxygen tolerant Heterohelix moremani and surface dweller Hedbergella planispira. After the anoxic event other species returned, but remained rare and sporadically present well into the lower Turonian, whereas Heterohelix moremani remained the single dominant species. The OAE2 biotic turnover suggests that the stress to calcareous plankton was related to changes in the watermass stratification, intensity of upwelling, nutrient flux and oxic levels in the water column driven by changes in climate and oceanic circulation. Results presented here demonstrate a 4-stage pattern of biotic response to the onset, duration, and recovery of OAE2 that is observed widely across the Tethys and its bordering epicontinental seas.     

New data on diatoms from the marine Paleogene of Western Kamchatka

Diatom assemblages from the marine Paleogene of Western Kamchatka (the Kovachina, Viventek, and Kuluven formations) are studied. The shallow-water neritic assemblage from the Kovachina Formation of the Pyatibratskii Cape section is tentatively attributed to the middle Eocene, as it is lacking zonal index species. A high abundance and great taxonomic diversity are characteristic of diatoms from the upper part of the Viventek Formation in the Kovran River section. The zonal index species Cavitatus rectus identified here among the diatoms suggests that the respective assemblage is macerated from the synonymous diatom zone of the middle Oligocene (29.6?28.2 Ma). According to its taxonomic composition and structure, this assemblage is indicative of transgression environments and the presumable influence of the middle Oligocene climatic optimum. The Rocella gelida (upper part) and Thalassiosira praefraga diatom zones of the Oligocene-Miocene transition are established in the Kuluven Formation sediments of the Mt. Uvuch section in Kovachina Bay.