Affiliation: | 1. Department of Forest and Conservation Sciences, University of British Columbia, Vancouver, British Columbia, Canada;2. Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, Ume?, SwedenLenka Kuglerová and Eliza Maher Hasselquist shared first authorship.;3. Department of Ecology and Environmental Science, Ume? University, Ume?, Sweden;4. Canadian Forest Service, Natural Resources Canada, Sault Ste. Marie, Ontario, Canada;5. Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, Ume?, Sweden |