In many arid ecosystems, vegetation frequently occurs in high-cover patches interspersed in a matrix of low plant cover. However, theoretical explanations for shrub patch pattern dynamics along climate gradients remain unclear on a large scale. This context aimed to assess the variance of the Reaumuria soongorica patch structure along the precipitation gradient and the factors that affect patch structure formation in the middle and lower Heihe River Basin (HRB). Field investigations on vegetation patterns and heterogeneity in soil properties were conducted during 2014 and 2015. The results showed that patch height, size and plant-to-patch distance were smaller in high precipitation habitats than in low precipitation sites. Climate, soil and vegetation explained 82.5% of the variance in patch structure. Spatially, R. soongorica shifted from a clumped to a random pattern on the landscape towards the MAP gradient, and heterogeneity in the surface soil properties (the ratio of biological soil crust (BSC) to bare gravels (BG)) determined the R. soongorica population distribution pattern in the middle and lower HRB. A conceptual model, which integrated water availability and plant facilitation and competition effects, was revealed that R. soongorica changed from a flexible water use strategy in high precipitation regions to a consistent water use strategy in low precipitation areas. Our study provides a comprehensive quantification of the variance in shrub patch structure along a precipitation gradient and may improve our understanding of vegetation pattern dynamics in the Gobi Desert under future climate change.
In order to model non‐Fickian transport behaviour in groundwater aquifers, various forms of the time–space fractional advection–dispersion equation have been developed and used by several researchers in the last decade. The solute transport in groundwater aquifers in fractional time–space takes place by means of an underlying groundwater flow field. However, the governing equations for such groundwater flow in fractional time–space are yet to be developed in a comprehensive framework. In this study, a finite difference numerical scheme based on Caputo fractional derivative is proposed to investigate the properties of a newly developed time–space fractional governing equations of transient groundwater flow in confined aquifers in terms of the time–space fractional mass conservation equation and the time–space fractional water flux equation. Here, we apply these time–space fractional governing equations numerically to transient groundwater flow in a confined aquifer for different boundary conditions to explore their behaviour in modelling groundwater flow in fractional time–space. The numerical results demonstrate that the proposed time–space fractional governing equation for groundwater flow in confined aquifers may provide a new perspective on modelling groundwater flow and on interpreting the dynamics of groundwater level fluctuations. Additionally, the numerical results may imply that the newly derived fractional groundwater governing equation may help explain the observed heavy‐tailed solute transport behaviour in groundwater flow by incorporating nonlocal or long‐range dependence of the underlying groundwater flow field. 相似文献
Knowledge of stock structure is key for the effective management of any fish species. Amphidromous fish, which live and spawn in freshwater but spend a pelagic larval period at sea, have typically been assumed to disperse widely during their larval phase, resulting in populations being sourced from a single unstructured larval pool. We used otolith microchemical analysis to examine the stock structure of bluegill bully (Gobiomorphus hubbsi), a declining amphidromous eleotrid endemic to New Zealand, along the west coast of South Island, New Zealand. Some drainages – even those in close proximity (c. 20?km) – were readily distinguishable based on otolith trace element concentrations, while little structure was evident between other geographically disparate locations. These results indicate that, at least in some cases, locally retained larvae, rather than a single unstructured larval pool, dominates recruitment. Management of bluegill bully and other amphidromous species must therefore consider the possibility of regionally distinct populations. 相似文献
Reliable quantification of savanna vegetation structure is critical for accurate carbon accounting and biodiversity assessment under changing climate and land-use conditions. Inventories of fine-scale vegetation structural attributes are typically conducted from field-based plots or transects, while large-area monitoring relies on a combination of airborne and satellite remote sensing. Both of these approaches have their strengths and limitations, but terrestrial laser scanning (TLS) has emerged as the benchmark for vegetation structural parameterization – recording and quantifying 3D structural detail that is not possible from manual field-based or airborne/spaceborne methods. However, traditional TLS approaches suffer from similar spatial constraints as field-based inventories. Given their small areal coverage, standard TLS plots may fail to capture the heterogeneity of landscapes in which they are embedded. Here we test the potential of long-range (>2000 m) terrestrial laser scanning (LR-TLS) to provide rapid and robust assessment of savanna vegetation 3D structure at hillslope scales. We used LR-TLS to sample entire savanna hillslopes from topographic vantage points and collected coincident plot-scale (1 ha) TLS scans at increasing distances from the LR-TLS station. We merged multiple TLS scans at the plot scale to provide the reference structure, and evaluated how 3D metrics derived from LR-TLS deviated from this baseline with increasing distance. Our results show that despite diluted point density and increased beam divergence with distance, LR-TLS can reliably characterize tree height (RMSE = 0.25–1.45 m) and canopy cover (RMSE = 5.67–15.91%) at distances of up to 500 m in open savanna woodlands. When aggregated to the same sampling grain as leading spaceborne vegetation products (10–30 m), our findings show potential for LR-TLS to play a key role in constraining satellite-based structural estimates in savannas over larger areas than traditional TLS sampling can provide. 相似文献