A quantitative investigation into the temporal and spatial variations in the emergence of adult blackflies (Diptera: Simuliidae) from the Breitenbach, a small upland stream in Germany

As a contribution to the long-term emergence studies carried out as part of the “Breitenbach ecosystem project”, this paper presents the results obtained on emergence patterns and population dynamics of blackflies from 1984 to 1988. The Breitenbach is a small first order stream near Schlitz/Hesse, running into the Fulda river. Adult blackflies were caught in 4 greenhouse emergence traps, each spanning the whole width of the stream for a length of 6 m. A manual and then a partly automated method for collecting trapped insects was applied for three years and one year, respectively. Sixteen blackfly-species of different origin were found in the traps: a) autochthonous species: Prosimulium tomosvaryi, Simulium vernum s.l., S. cryophilum s.l., S. ornatum s.l. (common); S. costatum, S. trifasciatum, S. monticola, S. argyreatum (rare); b) species of doubtful origin: S. lundstromi, S. angustitarse, S. angustipes, S. aureum (very rare); c) allochthonous species: S. lineatum, S. equinum, S. erythrocephalum, S. noelleri. It was shown that the last 4 species had not emerged from the Breitenbach but had flown into the traps as adults. In addition, females of autochthonous species with blood or with mature eggs were trapped, which were also considered to have flown in. Besides revealing a limitation of the trap construction, they supported the detailed interpretation of some intricated patterns of appearance. Variations in emergence patterns and specimen numbers between years and traps were pronounced, but only in a few cases could they be attributed to changes in abiotic factors such as water temperature or discharge. P. tomosvaryi had the most simple and regular life cycle, with one well synchronised emergence peak annually from April to May (or even to June). No gradient of specimen numbers along the stream was evident. The two closely related species S. vernum and S. cryophilum had quite similar emergence patterns: There were two broad peaks per year, extending mainly from March to June and from July to October or November. In some cases the number and separation of consecutive generations was not clear. The abundance of both species clearly decreased downstream, more so for S. cryophilum than for S. vernum. In a trap closest to a tributary spring, S. cryophilum was the dominant species during three of the four years examined. The adults of S. ornatum displayed an intricate pattern of appearance, with very low specimen numbers in spring and medium to very high numbers in July/August and September/October. S. ornatum is the only blackfly species that inhabits both the Breitenbach and the adjacent section of the Fulda river. It was shown that females emerging from the Fulda river regularly invade the Breitenbach valley in greatly varying numbers and oviposit there. This leads to overlapping larval cohorts with corresponding emergence peaks. Although S. ornatum was the most abundant species in one year in the lower traps (60 to 80% of all individuals), it remains uncertain whether it is a long-term, permanent member of the autochthonous blackfly fauna of the Breitenbach. Estimates of total numbers of flown-in adults, actual emergence, dry weight biomass, the ecological separation of the species and their life cycle strategies are discussed.     

Latest Pleistocene and Holocene glacier variations in the European Alps

In the Alps, climatic conditions reflected in glacier and rock glacier activity in the earliest Holocene show a strong affinity to conditions in the latest Pleistocene (Younger Dryas). Glacier advances in the Alps related to Younger Dryas cooling led to the deposition of Egesen stadial moraines. Egesen stadial moraines can be divided into three or in some cases even more phases (sub-stadials). Moraines of the earliest and most extended advance, the Egesen maximum, stabilized at 12.2 ± 1.0 ka based on ¹⁰Be exposure dating at the Schönferwall (Tyrol, Austria) and the Julier Pass-outer moraine (Switzerland). Final stabilization of moraines at the end of the Egesen stadial was at 11.3 ± 0.9 ka as shown by ¹⁰Be data from four sites across the Alps. From west to east the sites are Piano del Praiet (northwestern Italy), Grosser Aletschgletscher (central Switzerland), Julier Pass-inner moraine (eastern Switzerland), and Val Viola (northeastern Italy). There is excellent agreement of the ¹⁰Be ages from the four sites. In the earliest Holocene, glaciers in the northernmost mountain ranges advanced at around 10.8 ± 1.1 ka as shown by ¹⁰Be data from the Kartell site (northern Tyrol, Austria). In more sheltered, drier regions rock glacier activity dominated as shown, for example, at Julier Pass and Larstig valley (Tyrol, Austria). New ¹⁰Be dates presented here for two rock glaciers in Larstig valley indicate final stabilization no later than 10.5 ± 0.8 ka. Based on this data, we conclude the earliest Holocene (between 11.6 and about 10.5 ka) was still strongly affected by the cold climatic conditions of the Younger Dryas and the Preboreal oscillation, with the intervening warming phase having had the effect of rapid downwasting of Egesen glaciers. At or slightly before 10.5 ka rapid shrinkage of glaciers to a size smaller than their late 20th century size reflects markedly warmer and possibly also drier climate. Between about 10.5 ka and 3.3 ka conditions in the Alps were not conducive to significant glacier expansion except possibly during rare brief intervals. Past tree-line data from Kaunertal (Tyrol, Austria) in concert with radiocarbon and dendrochronologically dated wood fragments found recently in the glacier forefields in both the Swiss and Austrian Alps points to long periods during the Holocene when glaciers were smaller than they were during the late 20th century. Equilibrium line altitudes (ELA) were about 200 m higher than they are today and about 300 m higher in comparison to Little Ice Age (LIA) ELAs. The Larstig rock glacier site we dated with ¹⁰Be is the type area for a postulated mid-Holocene cold period called the Larstig oscillation (presumed age about 7.0 ka). Our data point to final stabilization of those rock glaciers in the earliest Holocene and not in the middle Holocene. The combined data indicate there was no time window in the middle Holocene long enough for rock glaciers of the size and at the elevation of the Larstig site to have formed. During the short infrequent cold oscillations between 10.5 and 3.3 ka small glaciers (less than several km²) may have advanced to close to their LIA dimensions. Overall, the cold periods were just too short for large glaciers to advance. After 3.3 ka, climate conditions became generally colder and warm periods were brief and less frequent. Large glaciers (for example Grosser Aletschgletscher) advanced markedly at 3.0–2.6 ka, around 600 AD and during the LIA. Glaciers in the Alps attained their LIA maximum extents in the 14th, 17th, and 19th centuries, with most reaching their greatest LIA extent in the final 1850/1860 AD advance.     

Isochron‐burial dating of glaciofluvial deposits: First results from the Swiss Alps

In this study, we use isochron‐burial dating to date the Swiss Deckenschotter, the oldest Quaternary deposits of the northern Alpine Foreland. Concentrations of cosmogenic ¹⁰Be and ²⁶Al in individual clasts from a single stratigraphic horizon can be used to calculate an isochron‐burial age based on an assumed initial ratio and the measured ²⁶Al/¹⁰Be ratio. We suggest that, owing to deep and repeated glacial erosion, the initial isochron ratio of glacial landscapes at the time of burial varies between 6.75 and 8.4. Analysis of 22 clasts of different lithology, shape, and size from one 0.5 m thick gravel bed at Siglistorf (Canton Aargau) indicates low nuclide concentrations: <20 000 ¹⁰Be atoms/g and <150 000 ²⁶Al atoms/g. Using an ²⁶Al/¹⁰Be ratio of 7.6 (arithmetical mean of 6.75 and 8.4), we calculate a mean isochron‐burial age of 1.5 ± 0.2 Ma. This age points to an average bedrock incision rate between 0.13 and 0.17 mm/a. Age data from the Irchel, Stadlerberg, and Siglistorf sites show that the Higher Swiss Deckenschotter was deposited between 2.5 and 1.3 Ma. Our results indicate that isochron‐burial dating can be successfully applied to glaciofluvial sediments despite very low cosmogenic nuclide concentrations. Copyright © 2017 John Wiley & Sons, Ltd.     

Identifying slope processes over time and their imprint in soils of medium‐high mountains of Central Europe (the Karkonosze Mountains,Poland)

Soils in mountainous areas are often polygenetic, developed in slope covers that relate to glacial and periglacial activities of the Pleistocene and Holocene and reflect climatic variations. Landscape development during the Holocene may have been influenced by erosion/solifluction that often started after the Holocene climatic optimum. To trace back soil evolution and its timing, we applied a multi‐methodological approach. This approach helped us to outline scenario of soil transformation. According to our results, some aeolian input must have occurred in the late Pleistocene. During that time and the early Holocene, the soils most likely had features of Cryosols or Leptosols. Physico‐chemical and mineralogical analyses have indicated that the material was denudated (between late Boreal to the Atlantic) from the ridge and upper‐slope positions forming a colluvium at mid‐slope positions. Later, during the Sub‐Boreal, mass wasting of the remains of silt material deposited at the end of the Pleistocene age on the ridge top seems to have occurred. In addition, the cool and moist conditions caused the deposition of a colluvium at the lower‐slope positions. The next phase was characterized by the transformation of Leptosols/Cambisols into Podzols at upper‐slope or shoulder positions and to Albic Cambisols at mid‐slope positions. During the Sub‐Boreal period, Stagnosols started to form at the lower part of the slope catena. Overall, the highest erosion rates were calculated at the upper‐slope position and the lowest rates at mid‐slope sites. Berylium‐10 (¹⁰Be) data showed that the Bs, BC/C were covered during the Holocene by a colluvium with a different geological composition which complicated the calculation of erosion or accumulation rates. The interpretation of erosion and accumulation rates in such multi‐layered materials may, therefore, be hampered. However, the multi‐methodological reconstruction we applied shed light on the soil and landscape evolution of the eastern Karkonosze Mountains. Copyright © 2017 John Wiley & Sons, Ltd.     

The impact of storm-triggered landslides on sediment dynamics and catchment-wide denudation rates in the southern Central Range of Taiwan following the extreme rainfall event of Typhoon Morakot

Extreme erosion events can produce large short-term sediment fluxes. Such events complicate erosion rates estimated from cosmogenic nuclide concentrations in river sediment by providing sediment with a concentration different from the long-term basin average. We present a detrital ¹⁰Be study in southern Taiwan, with multiple samples obtained in a time sequence bracketing the 2009 Typhoon Morakot, to assess the impact of landslide sediment on ¹⁰Be concentrations (N¹⁰Be) in river sediment. Sediment samples were collected from 13 major basins, two or three times over the last decade, to observe the temporal variation of N¹⁰Be. Landslide inventories with time intervals of 5–6 years were used to quantify sediment flux changes. A negative correlation between N¹⁰Be and landslide areal density indicates dilution of N¹⁰Be by landslide sediment. Denudation rates estimated from the diluted N¹⁰Be can be up to three times higher than the lowest rate derived from the same basins. Observed increases imply that, 3 years after the passage of Typhoon Morakot, fluvial channels still contain a considerable amount of sediment provided by hillslope landslides during the event. However, higher N¹⁰Be in 2016 samples indicate that the contribution from landslide sediment at the sampled grain size has decreased with time. The correlation between changes in N¹⁰Be and landslide area and volume is not strong, likely resulting from the stochastic and complex nature of sediment transport. To simultaneously evaluate the volume of landslide-derived sediment and estimate the background denudation rate, associated with less impulsive sediment supply, we constructed a sediment-mixing model with the time series of N¹⁰Be and landslide inventories. The spatial pattern of background erosion rate in southern Taiwan is consistent with the regional tectonic framework, indicating that the landscape is evolving mainly in response to the tectonic forcing, and this signal is modified, but not obscured by impulsive sediment supply. © 2019 John Wiley & Sons, Ltd.     

Fluvial dynamics and 14C-10Be disequilibrium on the Bolivian Altiplano

Determining sediment transfer times is key to understanding source-to-sink dynamics and the transmission of environmental signals through the fluvial system. Previous work on the Bolivian Altiplano applied the in situ cosmogenic ¹⁴C-¹⁰Be-chronometer to river sands and proposed sediment storage times of ~10–20 kyr in four catchments southeast of Lake Titicaca. However, the fidelity of those results hinges upon isotopic steady-state within sediment supplied from the source area. With the aim of independently quantifying sediment storage times and testing the ¹⁴C-¹⁰Be steady-state assumption, we dated sediment storage units within one of the previously investigated catchments using radiocarbon dating, cosmogenic ¹⁰Be-²⁶Al isochron burial dating, and ¹⁰Be-²⁶Al depth-profile dating. Palaeosurfaces appear to preserve remnants of a former fluvial system, which has undergone drainage reversal, reduction in catchment area, and local isostatic uplift since ~2.8 Ma. From alluvium mantling the palaeosurfaces we gained a deposition age of ~580 ka, while lower down fluvial terraces yielded ≤34 ka, and floodplains ~3–1 ka. Owing to restricted channel connectivity with the terraces and palaeosurfaces, the main source of channel sediment is via reworking of the late Holocene floodplain. Yet modelling a set of feasible scenarios reveals that floodplain storage and burial depth are incompatible with the ¹⁴C-¹⁰Be disequilibrium measured in the channel. Instead we propose that the ¹⁴C-¹⁰Be offset results from: (i) non-uniform erosion whereby deep gullies supply hillslope-derived debris; and/or (ii) holocene landscape transience associated with climate or human impact. The reliability of the ¹⁴C-¹⁰Be chronometer vitally depends upon careful evaluation of sources of isotopic disequilibrium in a wide range of depositional and erosional landforms in the landscape. © 2018 John Wiley & Sons, Ltd.     

Evidence of central Alpine glacier advances during the Younger Dryas–early Holocene transition period

The transition phase from Lateglacial to Holocene climate conditions was accompanied by a pronounced reorganization of climate patterns in the Northern Hemisphere. Evidence of Alpine palaeoglaciers provides a basis for understanding climate downturns during a time of generally warming conditions. In this context a series of well‐preserved and previously undated moraines were investigated in the small Falgin cirque located in the central Alpine Langtaufers Valley (South Tyrol, Italy) and in the neighbouring Hinteres Bergle cirque of the Radurschl Valley (North Tyrol, Austria). Both localities are situated in the driest area of the eastern Alps. They lie well above prominent moraines associated with the Younger Dryas (YD) cold phase and represent the first moraines below Little Ice Age (LIA) positions. The corresponding equilibrium line altitude of the palaeoglaciers in both cirques was 100–120 m lower than during the LIA. Surface exposure dating (¹⁰Be) of the inner Falgin moraines shows a mean stabilization age of 11.2±0.9 ka, which is similar to the deglaciation age of 10.9±0.8 ka for the Hinteres Bergle cirque. The ages indicate glacier activity most likely during the earliest Holocene or the YD/Holocene transition. These findings point to a climate with mean summer temperatures about 1.5 °C lower than during the 20th century in the Alps.     

The 10Be deglaciation chronology of the Göschenertal,central Swiss Alps,and new insights into the Göschenen Cold Phases

The Göschenertal (Göschenen valley) is the type locality of the so‐called Göschenen Cold Phases I (~3–2.3 ka) and II (~1.8–1.1 ka). According to earlier studies, these Late Holocene climatic cooling periods were characterized by changes in vegetation and pronounced glacier advances. As a peculiarity, the Göschenen Cold Phase I was thought to be connected to a local surge‐type advance of the Chelengletscher (Chelen glacier) – an exceptional event of unparalleled dimension in the European Alps. Based on cosmogenic ¹⁰Be exposure ages from moraine boulders, we investigated the local glacier chronology. In contrast to former research, moraines at different positions within the Göschenen valley (central Swiss Alps) have been dated to the Younger Dryas and the Early Holocene. This questions the applicability of palaeo‐Equilibrium Line Altitude (ELA) calculations for stadial attributions without additional numerical age constraints. Furthermore, we have found compelling evidence that the proposed non‐climatic glacier advance attributed to the Göschenen Cold Phase I did not occur. The present results, along with a reappraisal of the original study, question the scientific reliability and the glaciological definition of the Göschenen Cold Phases as glacier advances that clearly exceeded the Little Ice Age positions. While our data do not exclude potential changes in climate and vegetation, we nonetheless show that the Göschenen Cold Phases are not suitable as reference stadials in the system of Alpine Holocene glacier fluctuations.