Pseudo-cryptic species Arenicola defodiens and Arenicola marina (Polychaeta: Arenicolidae) in Wadden Sea,North Sea and Skagerrak: Morphological and molecular variation

The polychaete Arenicola defodiens, or black lug, was recently described as a morphologically highly similar species alongside the blow lug Arenicola marina. So far it was only known from the British Isles. A double spawning peak was observed earlier in lugworms of the western Wadden Sea. Here, we test the hypothesis that the two spawning peaks represent the two species in sympatry. This hypothesis is refuted on the basis of both morphological and mitochondrial DNA data; both spawning peaks are attributed to A. marina. In spite of this, we confirm, on the basis of new collections as well as re-examination of museum collections, the presence of A. defodiens in the western Wadden Sea, North Sea and also the Skagerrak; its distribution is restricted to habitats which are either submerged or have short emersion times, have relatively coarse sediments and high and stable salinities. Sympatry is common. The species differ strongly in the mitochondrial DNA fragment examined; the observed 14% uncorrected minimum difference amounts to an estimated 33–63 million years since sequence divergence. The amount of intraspecific molecular variation is larger for A. defodiens than for A. marina. This is evidence to suggest that A. marina may have undergone more recent and/or more severe population size bottlenecks.     

The angular momentum content of dwarf galaxies: new challenges for the theory of galaxy formation

We compute the specific angular momentum distributions for a sample of low-mass disc galaxies observed by Swaters. We compare these distributions to those of dark matter haloes obtained by Bullock et al. from high-resolution N -body simulations of structure formation in a ΛCDM universe. We find that although the disc mass fractions are significantly smaller than the universal baryon fraction, the total specific angular momenta of the discs are in good agreement with those of dark matter haloes. This suggests that discs form out of only a small fraction of the available baryons, but yet manage to draw most of the available angular momentum. In addition we find that the angular momentum distributions of discs are clearly distinct from those of the dark matter; discs lack predominantly both low and high specific angular momenta. Understanding these findings in terms of a coherent picture for disc formation is challenging. Cooling, feedback and stripping, which are the main mechanisms to explain the small disc mass fractions found, seem unable to simultaneously explain the angular momentum distributions of the discs. In fact, it seems that the baryons that make up the discs must have been born out of angular momentum distributions that are clearly distinct from those of ΛCDM haloes. However, the dark and baryonic mass components experience the same tidal forces, and it is therefore expected that they should have similar angular momentum distributions. Therefore, understanding the angular momentum content of disc galaxies remains an important challenge for our picture of galaxy formation.     

Amplification of vertically propagating SH waves by multiple layers of Gibson soils

Using the Haskell–Thomson transfer matrix approach, an analytical solution is obtained for SH wave amplification by multiple layers of Gibson soils (i.e. viscoelastic layers with linearly varying shear moduli). Amplification spectra for typical soil and basement rock properties are calculated. A comparison of the Gibson soil response with that obtained for homogeneous soil models shows generally stronger amplifications associated with the Gibson soil.     

A proposal for a new scenario framework to support research and assessment in different climate research communities

In this paper, we propose a scenario framework that could provide a scenario “thread” through the different climate research communities (climate change – vulnerability, impact, and adaptation - and mitigation) in order to support assessment of mitigation and adaptation strategies and climate impacts. The scenario framework is organized around a matrix with two main axes: radiative forcing levels and socio-economic conditions. The radiative forcing levels (and the associated climate signal) are described by the new Representative Concentration Pathways. The second axis, socio-economic developments comprises elements that affect the capacity for mitigation and adaptation, as well as the exposure to climate impacts. The proposed scenarios derived from this framework are limited in number, allow for comparison across various mitigation and adaptation levels, address a range of vulnerability characteristics, provide information across climate forcing and vulnerability states and span a full century time scale. Assessments based on the proposed scenario framework would strengthen cooperation between integrated-assessment modelers, climate modelers and vulnerability, impact and adaptation researchers, and most importantly, facilitate the development of more consistent and comparable research within and across these research communities.     

Quaternary uplift of northern England

Upland flats, attributable to erosion, have long been recognised in the landscape of the Lake District region of NW England, at altitudes of up to ~ 800 m O.D. Extrapolation using uplift rates derived from dated Pleistocene sites (karstic caves and other features) in the adjacent Pennine uplands suggests that if this succession of flats formed close to sea-level they date from the Middle Pliocene onwards, indicating a subsequent time-averaged uplift rate of almost 0.3 mm a^− 1. Numerical modelling indicates that erosion of surrounding areas at a typical rate of 0.2 mm a^− 1 since 3.1 Ma could have caused this uplift, as well as constraining the local effective viscosity of the lower crust as ~ 4 × 10¹⁸ Pa s and the typical local Moho temperature as ~ 650 °C. It is thus feasible that most of the topography of northern England has developed since the Middle Pliocene, as a consequence of coupling between erosion and the resulting induced flow in the lower continental crust. The much faster vertical crustal motions indicated in this part of northern England, compared with SE England, are thus mainly a consequence of much greater mobility of the lower crust in the north, due to its younger thermal age and the heating effect of radioactive Palaeozoic granites. Uplift of this magnitude, which has previously gone unrecognised, may have affected post-Pliocene global climate.