Adriatic ‘opisthobranchs’ (Gastropoda,Heterobranchia): shedding light on biodiversity issues

We herein review the Adriatic opisthobranch fauna, provide an updated checklist of 223 species and assess their distribution at regional and country levels. New Adriatic records are provided for 67 opisthobranch taxa, adding three new records for the Italian coastline, five new records for Albania, eight for Croatia and 15 for Montenegro. The presence of Hermaea bifida (Montagu, 1815), Hermaea variopicta (Costa A., 1869) and Facelina annulicornis (Chamisso & Eysenhardt, 1821) is reported for the first time from anywhere in the Adriatic Sea. Including the new findings, our inventory includes 28 species from Albania, 163 from Croatia, 178 from the Italian coastline of the Adriatic, 41 from Montenegro and 74 from Slovenia. No records were available from Bosnia and Hercegovina. Ninety species (40.4%) are widespread, and were recorded from all three main divisions (Northern, Western and Eastern Adriatic), whilst 79 species (35.4%) were reported from only one of them. At sub‐division levels, the Albanian inventory is the most dissimilar to the other country/regional lists, presumably because of the lack of targeted field surveys. The highest similarity is observed amongst the Western Adriatic, Croatia and the Italian Ionian coastline. Seven alien, one possible alien and three cryptogenic opisthobranchs were reliably recorded from the Adriatic Sea. The first Mediterranean records of five species were in the Adriatic Sea, of which four species have not been found elsewhere in the Mediterranean. Shipping and aquaculture are the probable pathways for most of the alien species. A steady rate of one to two alien introductions per decade since the 1970s was evident. The Northern Adriatic coasts are the most affected by alien invasions, probably due to repeated local introductions along with shellfish transfers (oysters, mussels, Manila clams), and heavy maritime transport.     

Multistage metamorphism and deformation in high‐pressure metabasites of the northern Adula Nappe Complex (Central Alps,Switzerland)

Metabasites from the northern Adula Nappe Complex (ANC) display a complex microstructural evolution recording episodes of deformation and metamorphic re‐equilibration that were obliterated in the surrounding phengite‐bearing schists. Pre‐D1 and D1 deformation episodes are preserved as mineral inclusions within garnet cores of some amphibole‐bearing eclogites and record high‐temperature greenschist‐/amphibolite‐facies conditions. D2 produced an eclogite‐facies foliation which developed at 580 ± 70°C and 19 ± 3 kbar. D3 was a composite deformation episode which can be divided into three sub‐episodes D3m, D3a and D3b which occurred as the metamorphism evolved from post‐eclogitic high‐pressure and low‐temperature conditions through to amphibolite‐facies conditions at 590 ± 30°C and 11.7 ± 1.3 kbar. The D3 deformation episode was responsible for the development of the S3 regional‐scale foliation in the surrounding schists, whilst D4 caused the development of an S4 greenschist foliation. The composite nature of the D3 episode indicates that rocks of the northern ANC experienced a protracted post‐eclogitic structural reworking and that the current structure of this part of the Alps is a late‐Alpine feature. Copyright © 2010 John Wiley & Sons, Ltd.     

Sar Interferometry And Its Applications

Satellite mounted Synthetic Aperture Radars (SAR) provide informationof the amplitude of the backscatterer its phase. This corresponds to the totaltravel time (source receiver and back), plus the phase of the scatterer itself.SAR interferometry yields the image of the differences of the phases takenin two successive passes. Or in the same pass if two receivers are availableon board; these images give a good digital elevation model of the terrain,with a vertical resolution that could be in the meter range. Millimetric motionof large areas of the terrain or of corner reflectors can also be measured withgood reliability; the system then measures subsidence, co-seismic motions andsmall motions of buildings and constructions. The coherence of t e interferometric pair, i.e., the permanence of the scattering characteristics of the terrain from one pass to the next combined with the more usual backscatterer amplitude, leads to high quality image segmentation. The possibilities and the limits of this technique are summarized.     

A hybrid optimization scheme for self-potential measurements due to multiple sheet-like bodies in arbitrary 2D resistivity distributions

Self-potential is a passive geophysical method that can be applied in a straightforward manner with minimum requirements in the field. Nonetheless, interpretation of self-potential data is particularly challenging due to the inherited non-uniqueness present in all potential methods. Incorporating information regarding the target of interest can facilitate interpretation and increase the reliability of the final output. In the current paper, a novel method for detecting multiple sheet-like targets is presented. A numerical framework is initially described that simulates sheet-like bodies in an arbitrary 2D resistivity distribution. A scattered field formulation based on finite differences is employed that allows the edges of the sheet to be independent of the grid geometry. A novel analytical solution for two-layered models is derived and subsequently used to validate the accuracy of the proposed numerical scheme. Lastly, a hybrid optimization is proposed that couples linear least-squares with particle-swarm optimization in order to effectively locate the edges of multiple sheet-like bodies. Through numerical and real data, it is proven that the hybrid optimization overcomes local minimal that occurs in complex resistivity distributions and converges substantially faster compared to traditional particle-swarm optimization.     

A discriminatory diagram of massive versus stratified deposits based on the sedimentation and bedload transportation rates. Experimental investigation and application to pyroclastic density currents

Dense gas-particle jets similar to collapsing eruption columns were generated by large-scale experiments. The column collapse resulted in a ground-hugging current forming stratified layers with bedding similar to natural pyroclastic density current deposits. At the impact of the collapsing column on the ground, a thick, massive bed was formed due to a high sedimentation rate that dumped turbulence due to high clast concentration. Down-current, flow expansion favoured turbulence and dilute gas-particle current that formed thin rippled layers deposited under traction. Experiments fed with fine ash (median size 0·066 mm) formed deposits without tractional structures, because fine particles, as other sedimentary fine material, is cohesive and exposes a limited surface to the shear stress. Experimental outcomes show that massive beds are formed where the sedimentation rate per unit width S_rw exceeds the bedload transportation rate Q_b by two orders of magnitude. A lower ratio generates traction at the base of the flow and formation of shear structures that increase in wavelength and height with a decreasing flux. This study presents a diagram that provides a useful addition for facies analysis of pyroclastic density currents, provided that deposits representing sustained sedimentation can be identified in the field. In the diagram a decrease in the S_rw/Q_b ratio corresponds to an increase in bedform size. Application of the diagram for hazard assessment purposes allows the reconstruction of the mass eruption rate of the Agnano–Monte Spina eruption at Campi Flegrei, which is the main variable defining the intensity of past eruptions, and of the Bingham rheology of the massive underflow of the Mercato pyroclastic density current at Vesuvius.     

Compound and hybrid clinothems of the last lowstand Mid-Adriatic Deep: Processes,depositional environments,controls and implications for stratigraphic analysis of prograding systems

Clinoforms with a range of scales are essential elements of prograding continental margins. Different types of clinoforms develop during margin growth, depending on combined changes in relative sea level, sediment supply and oceanographic processes. In studies of continental margin stratigraphy, trajectories of clinoform ‘rollover’ points are often used as proxies for relative sea-level variation and as predictors of the character of deposits beyond the shelf-break. The analysis of clinoform dynamics and rollover trajectory often suffers from the low resolution of geophysical data, the small scale of outcrops with respect to the dimensions of clinoform packages and low chronostratigraphic resolution. Here, through high-resolution seismic reflection data and sediment cores, we show how compound clinoforms were the most common architectural style of margin progradation of the late Pleistocene lowstand in the Adriatic Sea. During compound clinoform development, the shoreline was located landward of the shelf-break. It comprised a wave-dominated delta to the west and a barrier and back-barrier depositional system in the central and eastern area. Storm-enhanced hyperpycnal flows were responsible for the deposition of a sandy lobe in the river mouth, whereas a heterolithic succession formed elsewhere on the shelf. The storm-enhanced hyperpycnal flows built an apron of sand on the slope that interrupted an otherwise homogeneous progradational mudbelt. Locally, the late lowstand compound clinoforms have a flat to falling shelf-break trajectory. However, the main phase of shelf-break bypass and basin deposition coincides with a younger steeply rising shelf-break trajectory. We interpret divergence from standard models, linking shelf-break trajectory to deep-sea sand deposition, as resulting from a great efficiency of oceanographic processes in reworking sediment in the shelf, and from a high sediment supply. The slope foresets had a large progradational attitude during the late lowstand sea-level rise, showing that oceanographic processes can inhibit coastal systems to reach the shelf-edge. In general, our study suggests that where the shoreline does not coincide with the shelf-break, trajectory analysis can lead to inaccurate reconstruction of the depositional history of a margin.     

A systematic analysis of the XMM-Newton background: III. Impact of the magnetospheric environment

A detailed characterization of the particle induced background is fundamental for many of the scientific objectives of the Athena X-ray telescope, thus an adequate knowledge of the background that will be encountered by Athena is desirable. Current X-ray telescopes have shown that the intensity of the particle induced background can be highly variable. Different regions of the magnetosphere can have very different environmental conditions, which can, in principle, differently affect the particle induced background detected by the instruments. We present results concerning the influence of the magnetospheric environment on the background detected by EPIC instrument onboard XMM-Newton through the estimate of the variation of the in-Field-of-View background excess along the XMM-Newton orbit. An important contribution to the XMM background, which may affect the Athena background as well, comes from soft proton flares. Along with the flaring component a low-intensity component is also present. We find that both show modest variations in the different magnetozones and that the soft proton component shows a strong trend with the distance from Earth.     

Ecosystem exploitation and trophodynamic indicators: A comparison between the Northern Adriatic Sea and Southern New England

In an ecosystem-based resource management context, it is crucial to assess the relationships between community structure and ecosystem function and how those relationships change with resource extraction. To elucidate how changes in resource use can affect community structure and ecosystem function, we executed a comparative analysis of two different ecosystems subjected to notable fishing pressure. We contrasted the Northern Adriatic Sea (NAS) and Southern New England (SNE) ecosystems by examining outputs from comparable steady-state models. Both ecosystems have relatively high fishing pressure and a high biomass of benthic invertebrates. The basic structure of the food webs shows differences both in the number and definition of the functional groups, as described in the models. Fisheries, on the contrary, show similarities both in terms of catches and discards. Almost all statistics summarizing the structure and flows showed values three times higher in the SNE than in the NAS ecosystem, but despite this difference the two ecosystems exhibited similar, overall properties. Biomass ratios and the Mixed Trophic Impact (MTI) analysis showed that both ecosystems are dominated by the benthic compartment. Removing the biomass effect, however, shows a clear top-down effect, with a high rank achieved by fishing activities. In general terms, the low mean trophic level of catches and the high primary production required (PPR) values result in a high overexploitation level of the ecosystem, as highlighted by the L index. We conclude by exploring how comparative studies will continue to be valuable as ecosystem-based management is further implemented.     

The operational sex ratio of the sea urchin Paracentrotus lividus populations: the case of the Mediterranean marine protected area of Ustica Island (Tyrrhenian Sea, Italy)

We investigated, with a series of field and laboratory observations, the possible effect of the starfish Marthasteria glacialis predation on the operational sex ratio (OSR), i.e. the number of sexually mature males divided by the total number of sexually mature adults of both sexes at any one time, of the edible sea urchin Paracentrotus lividus . The OSR was estimated three times during the sea urchin summer spawning period (July 2004, June 2005 and July 2006) on barren substrates of Ustica Island Marine Protected Area (Southern Tyrrhenian Sea, Italy). Four sites were selected: two characterized by high M. glacialis density (take zone C) and two controls with low starfish density (no-take zone A). Mature sea urchins were independently collected by SCUBA diving and sexed. The adult sex ratio was skewed towards males at high M. glacialis density sites, whereas it was balanced (1:1) at predator low-density sites. Results of sex-selective feeding experiments in the laboratory showed that females of P. lividus were more vulnerable to M. glacialis predation. These outcomes underline the possible importance of M. glacialis in regulating the OSR of P. lividus populations.