The Late Cretaceous Pterophyllum (Bennettitales) in the North-East of Russia

The Late Cretaceous Pterophyllum species survived as relict elements in areas influenced by volcanism in the North-East of Russia. Two new species are described: P. philippoviae Gnilovskaya from the Turonian-Coniacian deposits of the Vetvinskaya Unit (Penzhina and Oklan Rivers interfluve, Magadan Region) and P. terechoviae from the Maastrichtian deposits of the Kakanaut Formation (Kakanaut River Basin, Koryak Upland). The upper boundary of Pterophyllum stratigraphic range is extended from the Turonian-Coniacian up to the K/Pg boundary. Pterophyllum terechoviae is the youngest member of the genus and probably the latest occurrence of Bennettitales in the Northern Hemisphere.     

Integrated biostratigraphy and palaeoenvironmental interpretation of the Upper Cretaceous to Paleocene succession in the northern Moldavidian Domain (Eastern Carpathians,Romania)

This study of the upper Maastrichtian to Danian sedimentary succession from the northern part of the Romanian Eastern Carpathians (Varniţa section) aims to establish an integrated biostratigraphy based on calcareous nannofossils, organic-walled dinoflagellate cysts (dinocysts) and foraminiferal assemblages, and to reconstruct the depositional environments of the interval. The stratigraphic record across the studied section is incomplete, considering that an approximately 16 m thick strata interval from the top of the Maastrichtian to lowermost Danian cannot be analyzed due to a landslide covering the outcrop. The upper Maastrichtian is marked by a succession of biostratigraphic events, such as the First Appearance Datum (FAD) of the nannoplankton taxon Nephrolithus frequens and FAD of the dinocyst species Deflandrea galeata and Disphaerogena carposphaeropsis, and the Last Appearance Datum (LAD) of Isabelidinium cooksoniae in the lower part of the section. These bioevents are followed by the LAD of the Dinogymnium spp. and Palynodinium grallator dinocyst markers in the top of the Maastrichtian deposits analyzed. In terms of foraminiferal biostratigraphy, the upper Maastrichtian Abathomphalus mayaroensis Zone is documented in the lower part of the studied section. Some bioevents, such as the bloom of the calcareous dinoflagellate genus Thoracosphaera and the FAD of the organic-walled dinocysts Damassadinium californicum, Senoniasphaera inornata, Xenicodinium lubricum and X. reticulatum suggest an early Danian age for the middle part of the section. From the Danian deposits in the Varniţa section, we describe a new organic-walled dinocyst species, Pentadinium darmirae sp. nov., which is until now the only species of the Pentadinium genus discovered in the Paleocene. The occurrence of the global Danian dinocyst marker Senoniasphaera inornata in the top of the section, suggests an age not younger than middle Danian (62.6 Ma) for the analyzed deposits.The palynofacies constituents, as well as the agglutinated foraminiferal morphogroups, used to reconstruct the depositional environments, show that the late Maastrichtian sediments were deposited in an outer shelf to distal (bathyal) environment, followed by a marine transgression during the Danian.     

Latest Campanian and Maastrichtian Siderolitidae (larger benthic foraminifera) from the Pyrenees (S France and NE Spain)

The siderolitids from the uppermost Campanian and Maastrichtian deposits of the Pyrenees have been re-studied. This has revealed a high diversity and rapid replacement of taxa, confirming the group as a good tool for high resolution biostratigraphy. Two genera have been found in the uppermost Campanian–Maastrichtian interval in the Pyrenean deposits: Siderolites Lamarck, and Wannierina Robles-Salcedo. Siderolites, with canaliferous spines or denticulate periphery, is represented by four species replacing each other from the latest Campanian to Maastrichtian: Siderolites praecalcitrapoides (latest Campanian), S. pyrenaicus sp. nov. (early Maastrichtian), Siderolites calcitrapoides (late Maastrichtian) and Siderolites denticulatus (late Maastrichtian). Wannierina is characterised by well-developed keels and ramified marginal canals. Two species of Wannierina have been identified and they succeeded one another from latest Campanian to early Maastrichtian: Wannierina vilavellensis sp. nov. (latest Campanian) and Wannierina cataluniensis (early Maastrichtian). The species of the genus Siderolites inhabited shallow waters of tropical to subtropical platforms with moderate-to-high water-energy conditions and those of the genus Wannierina are typical of deep–water low-energy environments but still in the eutrophic zone.     

Dinoflagellate biostratigraphy at the Campanian-Maastrichtian boundary in Zumaia,northern Spain

Dinoflagellate cyst assemblages from a well-exposed uppermost Cretaceous section at Zumaia (northern Spain) provide a basis for comparison with previous biostratigraphic and magnetostratigraphic studies on the problematic location of the Campanian-Maastrichtian boundary in the section. The position of the last occurrence of Corradinisphaeridium horridum and first common occurrence of Alterbidinium acutulum, correspond well with the bioevents defining the Campanian-Maastrichtian boundary in the Global boundary Stratotype Section and Point of Tercis les Bains (130 km to the North). Together with other age-diagnostic dinoflagellate cyst bioevents, we suggest that the boundary should be placed between 239.75 and 224.75 m below the Cretaceous-Palaeogene boundary, about 46 m lower than an interpretation based on the first occurrence of the planktonic foraminifer Pseudoguembelina palpebra and the last occurrence of the nannofossil Broinsonia parca subsp. constricta. A conspicuous acme of the dinoflagellate cyst Thalassiphora cf. delicata is encountered around the lower-upper Maastrichtian boundary (calibrated by foraminiferal, calcareous nannoplankton and magnetic polarity data), which may prove to be a useful correlatable event.     

An Upper Cretaceous (Campanian-Maastrichtian) actinopterygian fish assemblage from the marginal marine Adaffa Formation of Saudi Arabia

Actinopterygian remains have been recovered from Upper Cretaceous (lower Campanian to lower Maastrichtian) marginal marine deposits of the Adaffa Formation in northwestern Saudi Arabia. The fossils comprise gars (Lepisosteidae), pachycormids (cf. Protosphyraena sp.), indeterminate pycnodontiforms, enchodontid teleosts (cf. Enchodus sp.) and other indeterminate Teleostei. This assemblage is significant because it includes a novel occurrence for the Middle East (Pachycormidae) together with taxa (Lepisosteidae, Pycnodontiformes, Enchodontidae) that have been previously recorded from Late Cretaceous faunas elsewhere in the Mediterranean Tethyan region.     

A mosasaur,cf. Plotosaurus,from the upper Maastrichtian Quiriquina Formation in Central Chile

Here we describe a mosasaurid from the upper Maastrichtian Quiriquina Formation near the town of Cocholgüe about 30 km north of Concepción, central Chile. The specimen comprises a concretion preserving the rostral part of a pair of mandibular rami. The bulbous base of the teeth allows referral to cf. Plotosaurus sp. This is southernmost record of this piscivorous and pelagic taxon. The southern dispersal of mosasaurs during the late Maastrichtian may not only have depended from warm south-bound Equatorial currents but mainly from the nutrient-rich oceanic regime along an upwelling zone of cold abyssal water that mixed with the warm surface water, resulting in an enormous primary production. Apparently mosasaur dispersal depended more on food than on suitable temperatures or currents.     

Upper Maastrichtian ammonite biostratigraphy of the Gulf Coastal Plain (Mississippi Embayment,southern USA)

The Cretaceous outcrop belt of the Mississippi Embayment in the Gulf Coastal Plain (GCP) spans the Cretaceous/Paleogene (K/Pg) boundary. A detailed reconstruction of this time interval is critical for understanding the nature of biotic and environmental changes preceding the end-Cretaceous Mass Extinction event and for deciphering the likely extinction mechanism (i.e., bolide impact versus volcanism). Eight sections encompassing the K/Pg succession across the Mississippi Embayment were analyzed using biostratigraphic sampling of ammonites, dinoflagellates, and nannofossils. An upper Maastrichtian ammonite zonation is proposed as follows, from oldest to youngest: Discoscaphites conradi Zone, D. minardi Zone, and D. iris Zone. Our study documents that the ammonite zonation established in the Atlantic Coastal Plain (ACP) extends to the GCP. This zonation is integrated with nannofossil and dinoflagellate biostratigraphy to provide a framework to more accurately determine the age relationships in this region. We demonstrate that ammonites and dinoflagellates are more reliable stratigraphic indicators in this area than nannofossils because age-diagnostic nannofossils are not consistently present within the upper Maastrichtian in the GCP. This biostratigraphic framework has the potential to become a useful tool for correlation of strata both within the GCP and between the GCP, Western Interior, and ACP. The presence of the uppermost Maastrichtian ammonite D. iris, calcareous nannofossil Micula prinsii, and dinoflagellates Palynodinium grallator and Disphaerogena carposphaeropsis suggests that the K/Pg succession in the GCP is nearly complete. Consequently, the GCP is an excellent setting for investigating fine scale temporal changes across the K/Pg boundary and ultimately elucidating the mechanisms causing extinction.     

Dinosaur eggs in the Upper Cretaceous of the Coll de Nargó area,Lleida Province,south-central Pyrenees,Spain: Oodiversity,biostratigraphy and their implications

The Coll de Nargó area (Lleida Province, south-central Pyrenees) has yielded thousands of dinosaur eggs distributed in more than 30 levels across 370 m of Upper Cretaceous Tremp Formation deposits. Two stratigraphic units yielding dinosaur eggshells, eggs and clutches, can be distinguished. Multiple eggshells were collected from up to 75 different in situ clutches, located in consecutive stratigraphical levels throughout the Mas de Pinyes section. Four different oospecies have been recognized: Megaloolithus aureliensis, Megaloolithus siruguei, Megaloolithus cf. baghensis and Cairanoolithus cf. roussetensis. In absence of typical biochronological markers, fossil eggshells may have potential as a tool for dating. Based on these ootaxa, three oozones can be established along the stratigraphic section. The lower assemblage is composed by M. aureliensis, M. siruguei and Cairanoolithus cf. roussetensis; the middle one consists only of M. siruguei, while the upper oozone is characterized by M. cf. baghensis. Results suggest that the age of Coll de Nargó rocks ranges from the latest Campanian to the beginning of the Late Maastrichtian. Also reported is the first evidence of the oogenus Cairanoolithus outside of France, where it was regarded as an endemic ootaxon. The Coll de Nargó area is considered to be one of the most important dinosaur nesting areas in Europe recurrently used by several dinosaurs during a long time span. Finally, the presence of fossilized eggshell membrane in several specimens from Coll de Nargó contributes to understanding the taphonomic process of megaloolithid eggs in this area.     

Caprinula and Sauvagesia rudist faunas (Bivalvia) from the Cenomanian of NW Jordan. Stratigraphy and taxonomy

Three species of a canaliculated rudist Caprinula d'Orbigny, 1847, C. sharpei (Choffat, 1885), C. cedrorum (Blanckenhorn, 1890) and C. cf. boissyi d'Orbigny, 1840 and a radiolitid Sauvagesia sharpei (Bayle, 1857) are described from the Hummar Formation (upper Cenomanian) in NW Jordan, in the vicinity of Ajlun. Caprinula sharpei, C. cedrorum and S. sharpei are described for the first time from Jordan. Many specimens of S. sharpei are characterized by the presence of cavities flanking the lamellar myophores in the left valve and the apparence of the dorsal cavity and teeth/socket system moulds in the inner part of the outer shell layer of the right valve. A hiatus (or erosional unconformity) between Hummar Formation and upper Turonian Wadi As Sir Limestone Formation is suggested by the presence of karstic structures, reworked limestone clasts, and rudist fragments and a sharp boundary. Early diagenetic processes such as dissolution and silicification present in the loose rudist material is described.     

Late Maastrichtian cephalopods,dinoflagellate cysts and foraminifera from the Cretaceous–Paleogene succession at Lechówka,southeast Poland: Stratigraphic and environmental implications

The Lechówka section comprises the most complete Cretaceous–Paleogene (K-Pg) boundary succession in Poland and is among 29 sites worldwide with the youngest ammonite record. Here, cephalopods (ammonites and nautilids), organic-walled dinoflagellates (dinocysts) and foraminifera from the uppermost Maastrichtian interval are studied. In terms of ammonite biostratigraphy, the upper Maastrichtian Hoploscaphites constrictus crassus Zone is documented up to a level 120 cm below the K-Pg boundary. There is no direct, ammonite-based evidence of the highest Maastrichtian H. constrictus johnjagti Zone. However, the predominance of the dinocyst marker taxon Palynodinium grallator suggests the presence of the equivalent of the uppermost Maastrichtian Thalassiphora pelagica Subzone, which is correlatable with the H. c. johnjagti ammonite Zone. The planktonic foraminiferal assemblage is coeval with that from the H. c. johnjagti Zone as well. These data indicate that the top of the Maastrichtian at Lechówka is complete within the limits of biostratigraphic resolution, albeit slightly condensed. The dinocyst and foraminiferal assemblages are dominated by taxa that are characteristic of high-energy, marginal marine environments. A reduction in test size among the calcareous epifaunal benthic foraminifera is observed at a level 50 cm below the K-Pg boundary, which is possibly related to environmental stress associated with Deccan volcanism.     

Turonian cephalopods (ammonoids and a nautiloid) from the Wadi Daya Formation of the Talerzha Basin (South Riffian Ridges Domain,northern Morocco)

The Wadi Daya Formation, or the Calcaires crayeux of the older literature, attains a thickness of 10–40 m in the Talerhza Basin of the South Riffian Ridges. Previously, this unit was first dated as “Vraconian” (i.e., late upper Albian), but then reinterpreted as Cenomanian-Turonian and Cenomanian-Coniacian on the basis of foraminiferal and ostracod assemblages, respectively. Here, we record for the first time in the South Riffian Ridges, some typically Turonian ammonoids and a nautiloid species, namely Romaniceras (Yubariceras) cf. ornatissimum (Stoliczka), Spathites (Jeanrogericeras) cf. reveliereanus (Courtiller), Neoptychites cephalotus (Courtiller), Pachydesmoceras linderi (de Grossouvre), Lewesiceras peramplum (Mantell) and Angulithes galea (Fritsch, in Fritsch & Schlönbach). These species are herein described and illustrated. In view of these data, the underlying Marnes et marno-calcaires jaunes Formation, formerly dated as “Vraconian”, could in fact be of a middle to late Cenomanian date, in accordance with the age assignment based on planktonic foraminifera. Deposition of the overlying Marnes jaunes Formation, previously dated as Cenomanian-“Senonian”, probably started during the latest Turonian or earliest Coniacian.     

Small sized charophyte gyrogonites in the Maastrichtian of Coll de Nargó, Eastern Pyrenees: An adaptation to temporary floodplain ponds

Charophytes bearing small sized fructifications dominated in fluviatile floodplain facies (red beds) from the Maastrichtian of Coll de Nargó and neighbouring basins in the southern Pyrenees (Catalonia, Spain). These charophytes mainly belong to the genus Microchara and are often disregarded in biostratigraphic studies, which focus their attention instead on facies from permanent lakes that are richer in species which usually bear fructifications with a larger size range. However, small sized gyrogonites are also significant for biostratigraphic purposes and even include some of the index species for Maastrichtian biozones in Europe. Indeed, the charophyte assemblages from the Maastrichtian of Coll de Nargó belong to the Microchara punctata biozone, recently calibrated to the middle-upper Maastrichtian.Floodplain ponds from the Maastrichtian red beds of Coll de Nargó (Lower Red Unit) were extremely shallow, received considerable terrigenous influx and were frequently exposed, probably resulting in turbid, warm waters with high light availability. These conditions could explain the almost exclusive occurrence of charophytes with small fructifications in the Lower Red Unit. The available data, mainly based on oospores from extant species, indicate that the small size observed in gyrogonites from temporary ponds may represent an adaptation to environmental stress. Fossil species with small gyrogonites of Microchara cristata, M. punctata and Microchara nana would thus develop massively in stressed shallow ponds on fluvial floodplains. To contrast these hypotheses, we compared our results to those of four well-known case studies with similar sedimentological contexts, ranging from the Lower Cretaceous to the upper Eocene–lower Oligocene. Gyrogonite size patterns were similar in all cases, possibly suggesting that characeans display a long history of adaptation to shallow, temporary and turbid floodplain ponds by means of producing a high number of small gyrogonites, probably representing short life cycles and opportunistic strategies.     

Sympatry of two ankylosaurs (Hungarosaurus and cf. Struthiosaurus) in the Santonian of Hungary

A complete and well-preserved right ankylosaurian humerus from the Upper Cretaceous (Santonian) Csehbánya Formation of Iharkút, western Hungary is described here. Based on its osteological features and 21.5 cm adult length, the new specimen is markedly different from the slender humerus of Hungarosaurus, the previously known ankylosaur from the locality, and more similar to that of Struthiosaurus. Thus, the new Hungarian specimen is tentatively assigned here to cf. Struthiosaurus thereby dating back the first occurrence of this genus to the Santonian. The new fossil demonstrates the sympatric co-existence of two different nodosaurid ankylosaurs (a smaller, robust form with 2–2.5 m total body length and a larger, cursorial form with 4–4.5 m body length) in the Iharkút fauna. This also suggests that the pattern of the European ankylosaur diversity was more complex than previously thought.     

Biometry of the Late Cretaceous Arkhangelskiella group: ecophenotypes controlled by nutrient flux

This study focuses on the morphometry and taxonomy of the Late Cretaceous coccolith genus Arkhangelskiella. Sixty samples from the Campanian–Maastrichtian interval of DSDP Hole 390A (Blake Nose) were investigated for their contents of Arkhangelskiella spp. In each sample one hundred specimens of Arkhangelskiella spp. were examined by measuring the coccolith length and width, as well as the length and width of the central area. In the samples investigated the Arkhangelskiella group exhibits a large size variation, specimens length varies from 4.95 μm to 14.52 μm. Former taxonomic concepts, based on morphometry, subdivided the Arkhangelskiella group into three species: Arkhangelskiella maastrichtiana, Arkhangelskiella confusa and Arkhangelskiella cymbiformis. Our data show a large variability of the morphometric data (coccolith length, width of the outer rim). There is no indication for three independant species; two of the quoted taxa (1. thick outer rim = Arkhangelskiella maastrichtiana; 2. very thin outer rim = Arkkhangelskiella cymbiformis) seem to be extreme forms of a continuous morphometric lineage. The lower part of the investigated succession (139.92–133.42 mbsf) is dominated by small specimens with an average length of 6.8 μm whereas the upper part (132.86–126.15 mbsf) is characterized by larger specimens (mean 8.7 μm). In DSDP Hole 390A the size increase appears to be very abrupt, within two samples (samples 133.42 mbsf, 132.86 mbsf) the mean size increases by 1.51 μm. Previous morphometric studies of Arkhangelskiella indicate a more continuous size increase throughout the late Campanian–Maastrichtian. The abrupt size increase observed here hints toward a minor hiatus in DSDP Hole 390A separating upper Campanian from lower Maastrichtian sediments. It seems likely that the size increase of Arkhangelskiella reflects changes of various environmental factors like nutrient supply and sea water chemistry (Mg/Ca ratio; Ca concentration). A comparison of morphometric results with previous palaeoecological studies documents a nutrient control for the growth of Arkhangelskiella. Small specimens can be related to more mesotrophic conditions whereas large specimens are linked to oligotrophic surface waters.     

Discovery of the fossiliferous Cu Brei Formation (Lower Devonian) in the Kon Tum Block (South Viet Nam)

Lower Devonian corals and stromatoporoids have recently been discovered in limestones among low grade metamorphic rocks on the western margin of the Kon Tum Block (South Viet Nam). This unit has been identified as the Cu Brei Formation. Coral and stromatoporoid species have been described including Squameofavosites aff. spongiosus, Parallelostroma cf. multicolumnum, Amphipora cf. rasilis, A. cf. raritalis, Simplexodictyon cf. artyschtense, Stromatopora cf. boriarchinovi and Stromatopora sp. indet. The Cu Brei Formation is exposed in a small area 6 km in length and 3 km wide at the foot of Cu Brei Mountain (Sa Thay District, Kon Tum Province). As this formation is in marine shelf facies it is probable that further exposures of Lower Devonian sediments may be discovered in the Kon Tum Block. This discovery raises the question of the tectonic history of the metamorphic Kon Tum Block. It is possible that the block was not an area of positive uplift from the beginning of Paleozoic as has been supposed, but was submerged in a marine environment, at least on its outer margins, in the Devonian, and possibly even earlier, in Early Paleozoic.     

Biostratigraphy of the Cretaceous-Paleogene boundary deposits in the Arctic region of West Siberia (Implications of Foraminifers)

Benthic foraminifers from borehole sections recovered by drilling in the Yamal Peninsula, West Siberia, characterize the Ceratobulimina cretacea Beds (the upper Campanian-lower Maastrichtian) and the Spiroplectammina variabilis-Gaudryina rugosa spinulosa and Spiroplectammina kasanzevi-Bulimina rosenkrantzi regional zones of the lower and upper Maastrichtian, respectively. The Danian Stage is missing from the sections, which include marine deposits of the Selandian Stage attributed to the Ceratolamarckina tuberculata Beds. Foraminiferal assemblages of the beds include the Siberian endemic species associated with Paleocene foraminifers of the Midway-type fauna of subglobal distribution range. Occurrence of the latter suggests that warm-water surface currents from the North Atlantic reached southern areas of the Kara Sea.     

A new mammal track from the Laramie Formation (Maastrichtian) at the Fossil Trace locality,Golden, Colorado

A previously unknown and morphologically distinct pentadactyl mammal track was recovered from the Upper Cretaceous (Maastrichtian) Laramie Formation at the Fossil Trace site, a National Natural Landmark which is the type locality for Schadipes crypticus, the only named mammal track known from North America. The track is different, and larger than S. crypticus, and thus is evidence of a diversity of mammal trackmakers at this site. Although Cretaceous mammal tracks are very rare, preliminary indications are that those currently known are all morphologically distinct and therefore indicative of a global diversity of different trackmakers, as the body fossil record suggests. Lack of well-preserved mammal trackways with morphologically distinct manus and pes footprints hampers efforts to name diagnostic ichnotaxa.     

Lepidorbitoides (Foraminifera) from the lower Maastrichtian Kallankuruchchi Formation,Cauvery Basin,India: Morphometry and paleobiogeography

The southernmost occurrence of the early Maastrichtian larger benthic foraminifera (LBF) in Tethys is known from the Kallankuruchchi Formation in the Cauvery Basin, SE India, represented by Lepidorbitoides-Siderolites assemblages. The systematics, age and paleobiogeography of Lepidorbitoides here have as yet remained unresolved due to lack of information particularly on the nepionic arrangement, whereas their links with the Western Tethyan and Caribbean biogeographic domains were speculated. Lepidorbitoides, studied from the same level in seven samples in two separate areas, invariably possess quadriserial nepionts and adauxiliary chamberlets, whose mean number ranges from 3.79 to 4.67. The ratio between the sample means of the internal diameter of deuteroconch and protoconch varies between 1.72 and 1.86. The equatorial layer in the early stage consists of arcuate chambers with basal stolons, and ogival-to spatulate chamberlets with annular and oblique stolons in the later stages. These features are consistent with the phylogenetically advanced members of the Western Tethyan Lepidorbitoides lineage, such as L. minor (Schlumberger) and L. socialis (Leymerie), and all samples were assigned to the transitional development stages of these species based on the morphometry. The taxonomic status of some Lepidorbitoides species, originally described from the Kallankuruchchi Formation and widely adopted in previous works, such as L. blanfordi (Rao) and L. inornata (Rao), are not justified. We extend the geographic range of Western Tethyan Lepidorbitoides to southern India.